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<document id="7EFE7338EA58FBE716750519BDA78B20" ID-DOI="10.11646/zootaxa.4813.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3944139" ID-ZooBank="0ED34D69-0BC1-4E7D-A50D-6C0A31AB0374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594719214043" checkinUser="plazi" docAuthor="Jordal, Bjarte H. &amp; Smith, Sarah M." docDate="2020" docId="9E3D1C0AFB37FFB5FF62FDEB1804F831" docLanguage="en" docName="zootaxa.4813.1.1.pdf" docOrigin="Zootaxa 4813 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scolytodes lapillus Jordal and Smith 2020, sp. nov." docType="treatment" docUuid="AA1272A7-3E32-426C-94B6-505D3EA1E84B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="58" masterDocId="62046472FB0EFF8CFFF5FFEA1918FF8A" masterDocTitle="Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences" masterLastPageNumber="67" masterPageNumber="1" pageNumber="58" updateTime="1698781377542" updateUser="plazi">
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<mods:title id="275386FBB2972DB7EE13677602730FCB">Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences</mods:title>
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<mods:namePart id="509FC8F1B63170561D27C9E3FC0AC433">Jordal, Bjarte H.</mods:namePart>
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<mods:namePart id="BCE964089DABCB445B1856B7B81D9AD6">Smith, Sarah M.</mods:namePart>
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<mods:title id="3DA48FB94FA674D873E8CEE6CF809FF5">Zootaxa</mods:title>
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<mods:date id="CC8B93B7A66641B47B35D962EBC370C7">2020</mods:date>
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<mods:number id="1B33F00D4C6B2B64125B2D3F5FA29695">2020-07-14</mods:number>
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<treatment id="9E3D1C0AFB37FFB5FF62FDEB1804F831" ID-DOI="http://doi.org/10.5281/zenodo.4338648" ID-Zenodo-Dep="4338648" LSID="urn:lsid:zoobank.org:act:AA1272A7-3E32-426C-94B6-505D3EA1E84B" httpUri="http://treatment.plazi.org/id/9E3D1C0AFB37FFB5FF62FDEB1804F831" lastPageNumber="58" pageId="57" pageNumber="58">
<subSubSection id="5E8EFE97FB37FFB5FF62FDEB1AD3FDB5" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="162BAD1CFB37FFB5FF62FDEB1BDDFD91" blockId="57.[151,971,513,611]" box="[151,709,513,540]" pageId="57" pageNumber="58">
<heading id="4D631A70FB37FFB5FF62FDEB1BDDFD91" bold="true" box="[151,709,513,540]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="24E0710EFB37FFB5FF62FDEB1BDDFD91" bold="true" box="[151,709,513,540]" pageId="57" pageNumber="58">
<taxonomicName id="D194D69FFB37FFB5FF62FDEB1B43FD91" authority="Jordal and Smith" authorityName="Jordal and Smith" authorityYear="2020" box="[151,603,513,540]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="lapillus" status="sp. nov.">
<emphasis id="24E0710EFB37FFB5FF62FDEB1860FD96" bold="true" box="[151,376,513,540]" italics="true" pageId="57" pageNumber="58">Scolytodes lapillus</emphasis>
Jordal and Smith
</taxonomicName>
,
<taxonomicNameLabel id="3FD3CC75FB37FFB5FD9DFDEB1BDDFD91" box="[616,709,513,539]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
<uuid id="623297C9FB37FFB5FE58FDCE1AD3FDB5" box="[429,971,548,575]" pageId="57" pageNumber="58">AA1272A7-3E32-426C-94B6-505D3EA1E84B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF62FDA31863FDE9" box="[151,379,585,611]" pageId="57" pageNumber="58" type="description">
<paragraph id="162BAD1CFB37FFB5FF62FDA31863FDE9" blockId="57.[151,971,513,611]" box="[151,379,585,611]" pageId="57" pageNumber="58">
(
<figureCitation id="8EAFB199FB37FFB5FF6AFDA3186BFDE9" box="[159,371,585,611]" captionStart="FIGURES 127135" captionStartId="58.[151,264,1867,1892]" captionTargetBox="[155,1430,191,1841]" captionTargetId="figure@58.[151,1436,181,1842]" captionTargetPageId="58" captionText="FIGURES 127135. Dorsal, lateral and frontal views of the female holotype of Scolytodes lapillus (127, 130, 133), and Scolytodes coronatus (128, 131, 134), and a female specimen of Scolytodes grandis (129, 132, 135)." figureDoi="http://doi.org/10.5281/zenodo.3944170" httpUri="https://zenodo.org/record/3944170/files/figure.png" pageId="57" pageNumber="58">Figs 127, 130, 133</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF62FD7B1D78FD79" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="162BAD1CFB37FFB5FF62FD7B1D78FD79" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF62FD7B185FFD21" bold="true" box="[151,327,656,683]" pageId="57" pageNumber="58">Type material.</emphasis>
<materialsCitation id="A6FCA741FB37FFB5FEA5FD7B1BCAFD45" collectingDate="2012-07-27" collectorName="J. Hulcr" country="Ecuador" location="Tiputini" municipality="Prov" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="24E0710EFB37FFB5FEA5FD7B1B3DFD21" bold="true" box="[336,549,656,683]" pageId="57" pageNumber="58">
<typeStatus id="C92F13BEFB37FFB5FEA5FD7B18A7FD21" box="[336,447,657,683]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="6E83ED8CFB37FFB5FDDBFD7B1B88FD21" box="[558,656,657,683]" name="Ecuador" pageId="57" pageNumber="58">Ecuador</collectingCountry>
:
<collectingRegion id="D45063FEFB37FFB5FD69FD7B1BE4FD21" box="[668,764,657,683]" country="Ecuador" name="Orellana" pageId="57" pageNumber="58">Orellana</collectingRegion>
<collectingMunicipality id="F64F3766FB37FFB5FCF0FD7B1A23FD21" box="[773,827,657,683]" pageId="57" pageNumber="58">Prov</collectingMunicipality>
,
<location id="134BFBC7FB37FFB5FCB2FD7B1AB8FD21" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB37FFB5FF62FDEB1804F831:134BFBC7FB37FFB5FCB2FD7B1AB8FD21" box="[839,928,657,683]" country="Ecuador" municipality="Prov" name="Tiputini" pageId="57" pageNumber="58" stateProvince="Orellana">Tiputini</location>
biodiversity station, GIS: -0.637°, -76.150°, ex bark on tree branch,
<date id="622A8BDCFB37FFB5FE6AFD5F1B21FD45" box="[415,569,693,719]" pageId="57" pageNumber="58" value="2012-07-27">
<collectingDate id="726E7234FB37FFB5FE6AFD5F1B21FD45" box="[415,569,693,719]" pageId="57" pageNumber="58" value="2012-07-27">27. July 2012</collectingDate>
</date>
,
<collectorName id="BB61C8CAFB37FFB5FDB1FD5F1B86FD45" box="[580,670,693,719]" pageId="57" pageNumber="58">J. Hulcr</collectorName>
, leg.
</materialsCitation>
<emphasis id="24E0710EFB37FFB5FD2CFD5E1A9FFD45" bold="true" box="[729,903,692,719]" pageId="57" pageNumber="58">
<typeStatus id="C92F13BEFB37FFB5FD2CFD5E1A26FD45" box="[729,830,692,719]" pageId="57" pageNumber="58" type="allotype">Allotype</typeStatus>
male:
</emphasis>
same data as holotype.
<materialsCitation id="A6FCA741FB37FFB5FB67FD5F187AFD7E" collectingDate="2012-07-27" collectorName="J. Hulcr" country="Ecuador" location="Tiputini" municipality="Prov" pageId="57" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Orellana" typeStatus="paratype">
<emphasis id="24E0710EFB37FFB5FB67FD5F1C10FD45" bold="true" box="[1170,1288,693,719]" pageId="57" pageNumber="58">
<typeStatus id="C92F13BEFB37FFB5FB67FD5F1C1BFD45" box="[1170,1283,693,719]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
:
</emphasis>
same data as holotype (
<specimenCount id="00926695FB37FFB5FEFEFD331842FD7E" box="[267,346,729,756]" pageId="57" pageNumber="58" type="male">1 male</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="A6FCA741FB37FFB5FE99FD331A52FD79" box="[364,842,729,755]" collectionCode="QCAZ" country="Ecuador" pageId="57" pageNumber="58" specimenCount="2" stateProvince="Orellana" typeStatus="holotype">
<typeStatus id="C92F13BEFB37FFB5FE99FD3318CCFD79" box="[364,468,729,755]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
and
<typeStatus id="C92F13BEFB37FFB5FDFEFD331B7DFD79" box="[523,613,729,755]" pageId="57" pageNumber="58" type="allotype">allotype</typeStatus>
deposited in
<collectionCode id="708535D9FB37FFB5FD0EFD331A52FD79" box="[763,842,729,755]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="57" pageNumber="58" type="Museum">QCAZ</collectionCode>
</materialsCitation>
,
<materialsCitation id="A6FCA741FB37FFB5FCA0FD331D44FD79" box="[853,1116,729,755]" collectionCode="MSUC" country="Ecuador" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
one
<typeStatus id="C92F13BEFB37FFB5FC70FD331AFDFD79" box="[901,997,729,755]" pageId="57" pageNumber="58" type="paratype">paratype</typeStatus>
in
<collectionCode id="708535D9FB37FFB5FBFCFD331D44FD79" box="[1033,1116,729,755]" pageId="57" pageNumber="58">MSUC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF32FD161829FC2D" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="162BAD1CFB37FFB5FF32FD161829FC2D" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32FD16185AFC9D" bold="true" box="[199,322,764,791]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
Interstriae 10 sharply raised to near apex. Protibiae with a tiny additional mesal spine near tarsal insertion. Pronotum strongly asperate on anterior half. Most closely related to
<taxonomicName id="D194D69FFB37FFB5FBE0FCCB1C18FCB1" authority="Jordal, 2018" authorityName="Jordal" authorityYear="2018" box="[1045,1280,801,827]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="24E0710EFB37FFB5FBE0FCCB1D74FCB1" box="[1045,1132,801,827]" italics="true" pageId="57" pageNumber="58">S. rufus</emphasis>
<bibRefCitation id="7205D0EDFB37FFB5FB81FCCB1C18FCB1" author="Jordal, B. H." box="[1140,1280,801,827]" pageId="57" pageNumber="58" pagination="1" refId="ref45287" refString="Jordal, B. H. (2018) Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa, 4504 (1), 076 - 104. https: // doi. org / 10.11646 / zootaxa. 4504.1.4" type="journal article" year="2018">Jordal, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D194D69FFB37FFB5FACFFCCB183BFCD5" authority="Wood, 1983" authorityName="Wood" authorityYear="1983" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="retifer">
<emphasis id="24E0710EFB37FFB5FACFFCCB1C84FCB1" box="[1338,1436,801,827]" italics="true" pageId="57" pageNumber="58">S. retifer</emphasis>
Wood, 1983
</taxonomicName>
, but is distinguished from both species by the more impressed striae and the longer and thicker interstrial setae, and further from
<taxonomicName id="D194D69FFB37FFB5FE14FC831B5BFC09" authorityName="Wood" authorityYear="1983" box="[481,579,873,899]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="retifer">
<emphasis id="24E0710EFB37FFB5FE14FC831B5BFC09" box="[481,579,873,899]" italics="true" pageId="57" pageNumber="58">S. retifer</emphasis>
</taxonomicName>
by stronger asperities on pronotum, and further from
<taxonomicName id="D194D69FFB37FFB5FB5DFC831DE7FC09" authorityName="Jordal" authorityYear="2018" box="[1192,1279,873,899]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="24E0710EFB37FFB5FB5DFC831DE7FC09" box="[1192,1279,873,899]" italics="true" pageId="57" pageNumber="58">S. rufus</emphasis>
</taxonomicName>
by the longer interstriae 10.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF32FC5A1AC5F9FD" pageId="57" pageNumber="58" type="description">
<paragraph id="162BAD1CFB37FFB5FF32FC5A18CEF9A5" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32FC5A18AFFC41" bold="true" box="[199,439,944,971]" pageId="57" pageNumber="58">Description, female.</emphasis>
Length 2.0 mm, 2.4 × as long as wide; colour light brown, presumably much darker on older specimens.
<emphasis id="24E0710EFB37FFB5FEA8FC3F18BBFC65" bold="true" box="[349,419,981,1007]" pageId="57" pageNumber="58">Head.</emphasis>
Eyes lightly emarginated, separated above by 2.73.1 × their width. Frons flattened from vertex between eyes to epistoma, except lightly elevated and impunctate area on median two-thirds from epistoma to just below upper level of eyes, densely punctured elsewhere; vestiture consisting of a fringe of rather short setae from vertex along the margin of the impressed area to epistoma, pointing orad. Antennal club thick, asymmetrically attached to funiculus, pilose, setae short, a faint V-shaped septum barely visible. Funiculus possibly 6-segmented.
<emphasis id="24E0710EFB37FFB5FF62FB631808FB29" bold="true" box="[151,272,1161,1187]" pageId="57" pageNumber="58">Pronotum</emphasis>
reticulate, dull, punctures small, shallow, spaced by 13 × their diameter, punctures not reaching anterior margin, replaced on anterior half by rough, sharp asperities; anterior part of lateral carinae curved towards upper anterior margin, anterior row of asperities forming a low rim. Vestiture consisting mainly of 8 long, erect setae (4-2- 2).
<emphasis id="24E0710EFB37FFB5FF49FB1F1810FA85" bold="true" box="[188,264,1269,1295]" pageId="57" pageNumber="58">Elytra</emphasis>
shiny, smooth; striae regular, 1 and 2, sometimes 3 and 4 lightly impressed, more strongly so on declivity, punctures small, deep, separated by 12 × their diameter; interstriae 34 × broader than striae, punctures slightly smaller, more widely separated in irregular rows. Interstriae 10 weakly raised to apex. Vestiture consisting of 1620 long, erect, almost spatulate, setae on posterior part of interstriae 3, 5, 7 and 9.
<emphasis id="24E0710EFB37FFB5FBF7FA8B1D59FAF1" bold="true" box="[1026,1089,1377,1403]" pageId="57" pageNumber="58">Legs.</emphasis>
Procoxae separated by 0.30.4 × the width of one procoxa. Mesocoxae separated by 0.70.8 × the width of a procoxa. Protibiae narrow, distal teeth 1 similar to 2, with 3 additional transverse rugae towards tibial base; an additional mesal tooth or spine present on posterior face near base of tooth 2; protibial mucro short, thin, weakly curved posteriorly. Meso- and metatibiae with 7 or 67 socketed teeth on distal half.
<emphasis id="24E0710EFB37FFB5FDCAFA1B1A1FF981" bold="true" box="[575,775,1520,1547]" pageId="57" pageNumber="58">Ventral vestiture</emphasis>
. Setae on mesanepisternum trifid, on metanepisternum and metasternum mainly simple.
</paragraph>
<paragraph id="162BAD1CFB37FFB5FF32F9D31AC5F9FD" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32F9D31813F9D9" bold="true" box="[199,267,1593,1619]" pageId="57" pageNumber="58">Male.</emphasis>
Similar to female except frons convex, transversely impressed just above epistoma, surface strongly reticulate, dull, with deep punctures; vestiture consisting scant, short setae.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF32F96B1A9EF8A1" pageId="57" pageNumber="58" type="key">
<paragraph id="162BAD1CFB37FFB5FF32F96B1A9EF8A1" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32F96B1884F911" bold="true" box="[199,412,1664,1691]" pageId="57" pageNumber="58">
Key (
<bibRefCitation id="7205D0EDFB37FFB5FEF0F96A1897F911" author="Wood, S. L." box="[261,399,1664,1691]" pageId="57" pageNumber="58" refId="ref46297" refString="Wood, S. L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Brigham Young University Press, Provo, Utah, 900 pp." type="book" year="2007">Wood 2007</bibRefCitation>
).
</emphasis>
Depending on the judgement of the additional spine on protibia (possibly not a proper additional tooth), one may arrive at couplet 58b,
<taxonomicName id="D194D69FFB37FFB5FD73F94F1A44F935" ID-CoL="4VT2X" authority="(Eggers)" baseAuthorityName="Eggers" box="[646,860,1701,1727]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="24E0710EFB37FFB5FD73F94F1BEBF935" box="[646,755,1701,1727]" italics="true" pageId="57" pageNumber="58">S. bicolor</emphasis>
(Eggers)
</taxonomicName>
, but differs by having erect interstrial setae, smaller elytral punctures and larger asperities on the pronotum, and by the different female frons. If one recognizes the spine as an additional tooth, one may reach couplet 93,
<taxonomicName id="D194D69FFB37FFB5FD4EF9071A03F88D" ID-CoL="4VT87" authorityName="Wood" authorityYear="1983" box="[699,795,1773,1799]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="retifer">
<emphasis id="24E0710EFB37FFB5FD4EF9071A03F88D" box="[699,795,1773,1799]" italics="true" pageId="57" pageNumber="58">S. retifer</emphasis>
</taxonomicName>
, but differs by stronger asperities on the pronotum, by the less plumose setae in the female frons, and by the impressed striae.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF32F8DF1D03F8F9" pageId="57" pageNumber="58" type="etymology">
<paragraph id="162BAD1CFB37FFB5FF32F8DF1D03F8F9" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32F8DF1856F8C5" bold="true" box="[199,334,1845,1871]" pageId="57" pageNumber="58">Etymology.</emphasis>
The Latin name
<taxonomicName id="D194D69FFB37FFB5FDE0F8DF1B71F8C5" authorityName="Jordal and Smith" authorityYear="2020" box="[533,617,1845,1871]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="lapillus">
<emphasis id="24E0710EFB37FFB5FDE0F8DF1B71F8C5" box="[533,617,1845,1871]" italics="true" pageId="57" pageNumber="58">lapillus</emphasis>
</taxonomicName>
is a masculine noun, meaning precious stone (such as a gem or jewel), referring to the large, shiny and lightly elevated central field in the female frons.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB37FFB5FF32F8961804F831" pageId="57" pageNumber="58" type="biology_ecology">
<paragraph id="162BAD1CFB37FFB5FF32F8961804F831" blockId="57.[151,1437,656,1979]" pageId="57" pageNumber="58">
<emphasis id="24E0710EFB37FFB5FF32F89618EAF81D" bold="true" box="[199,498,1916,1943]" pageId="57" pageNumber="58">Biology and distribution.</emphasis>
<materialsCitation id="A6FCA741FB37FFB5FE0CF8971800F831" country="Ecuador" location="It" pageId="57" pageNumber="58" specimenCount="1">
Only known from the lowland
<typeStatus id="C92F13BEFB37FFB5FCA6F8971A9BF81D" box="[851,899,1917,1943]" pageId="57" pageNumber="58">type</typeStatus>
locality in
<collectingCountry id="6E83ED8CFB37FFB5FBF7F8971D7AF81D" box="[1026,1122,1917,1943]" name="Ecuador" pageId="57" pageNumber="58">Ecuador</collectingCountry>
.
<location id="134BFBC7FB37FFB5FB99F8971D65F81D" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB37FFB5FF62FDEB1804F831:134BFBC7FB37FFB5FB99F8971D65F81D" box="[1132,1149,1917,1943]" country="Ecuador" name="It" pageId="57" pageNumber="58">It</location>
was taken from bark of a tree branch
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>