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<document id="39552C8AB32F0EFF5CCD2E5EE245BA6C" ID-DOI="10.11646/zootaxa.4813.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3944139" ID-ZooBank="0ED34D69-0BC1-4E7D-A50D-6C0A31AB0374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594719214043" checkinUser="plazi" docAuthor="Jordal, Bjarte H. &amp; Smith, Sarah M." docDate="2020" docId="9E3D1C0AFB2DFFA8FF62FA1B1DAAFCB1" docLanguage="en" docName="zootaxa.4813.1.1.pdf" docOrigin="Zootaxa 4813 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scolytodes virgatus Jordal and Smith 2020, sp. nov." docType="treatment" docUuid="6B55F132-942E-484C-9F5C-34A3B9ABD836" docUuidSource="ZooBank" docVersion="5" lastPageNumber="37" masterDocId="62046472FB0EFF8CFFF5FFEA1918FF8A" masterDocTitle="Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences" masterLastPageNumber="67" masterPageNumber="1" pageNumber="36" updateTime="1698781377542" updateUser="plazi">
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<mods:title id="D3E36A2C168B6EE2011FE933CD15E983">Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences</mods:title>
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<mods:namePart id="1892FDBC2A35A86D3AB56159E5C68F61">Jordal, Bjarte H.</mods:namePart>
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<mods:namePart id="1BCDF6B6FB3AA933E2CE2EEF2DA424C0">Smith, Sarah M.</mods:namePart>
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<mods:title id="543F0870ABED561F74B0CDEE23A5DFC6">Zootaxa</mods:title>
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<mods:date id="869F935C796DCBC162D49319F4EE8F95">2020</mods:date>
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<mods:number id="A0BD713A0DD7C3FFB80FAE444ADB1073">2020-07-14</mods:number>
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<mods:number id="40E5E7D915EEF73A2CDC26BBB3E89E6C">4813</mods:number>
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<treatment id="9E3D1C0AFB2DFFA8FF62FA1B1DAAFCB1" ID-DOI="http://doi.org/10.5281/zenodo.4338702" ID-Zenodo-Dep="4338702" LSID="urn:lsid:zoobank.org:act:6B55F132-942E-484C-9F5C-34A3B9ABD836" httpUri="http://treatment.plazi.org/id/9E3D1C0AFB2DFFA8FF62FA1B1DAAFCB1" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="5E8EFE97FB2DFFAFFF62FA1B1ADDF9BA" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="162BAD1CFB2DFFAFFF62FA1B1BD5F986" blockId="35.[151,965,1521,1619]" box="[151,717,1521,1548]" pageId="35" pageNumber="36">
<heading id="4D631A70FB2DFFAFFF62FA1B1BD5F986" bold="true" box="[151,717,1521,1548]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="24E0710EFB2DFFAFFF62FA1B1BD5F986" bold="true" box="[151,717,1521,1548]" pageId="35" pageNumber="36">
<taxonomicName id="D194D69FFB2DFFAFFF62FA1B1B7BF981" authority="Jordal and Smith" authorityName="Jordal and Smith" authorityYear="2020" box="[151,611,1521,1548]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="virgatus" status="sp. nov.">
<emphasis id="24E0710EFB2DFFAFFF62FA1B1898F981" bold="true" box="[151,384,1521,1548]" italics="true" pageId="35" pageNumber="36">Scolytodes virgatus</emphasis>
Jordal and Smith
</taxonomicName>
,
<taxonomicNameLabel id="3FD3CC75FB2DFFAFFD85FA181BD5F986" box="[624,717,1522,1548]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="162BAD1CFB2DFFAFFF62F9FF1ADDF9BA" blockId="35.[151,965,1521,1619]" box="[151,965,1557,1584]" pageId="35" pageNumber="36">
<uri id="6205A11EFB2DFFAFFF62F9FF1ADDF9BA" box="[151,965,1557,1584]" pageId="35" pageNumber="36">
urn:lsid:zoobank.org:act:
<uuid id="623297C9FB2DFFAFFE5EF9FF1ADDF9BA" box="[427,965,1557,1584]" pageId="35" pageNumber="36">6B55F132-942E-484C-9F5C-34A3B9ABD836</uuid>
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</paragraph>
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<subSubSection id="5E8EFE97FB2DFFAFFF62F9D31849F9D9" box="[151,337,1593,1619]" pageId="35" pageNumber="36" type="description">
<paragraph id="162BAD1CFB2DFFAFFF62F9D31849F9D9" blockId="35.[151,965,1521,1619]" box="[151,337,1593,1619]" pageId="35" pageNumber="36">
(
<figureCitation id="8EAFB199FB2DFFAFFF6AF9D31851F9D9" box="[159,329,1593,1619]" captionStart="FIGURES 6472" captionStartId="37.[151,264,1890,1915]" captionTargetBox="[156,1432,195,1865]" captionTargetId="figure@37.[155,1433,181,1865]" captionTargetPageId="37" captionText="FIGURES 6472. Dorsal, lateral and frontal views of the female holotype of Scolytodes virgatus (64, 67, 70), Scolytodes criniger (65, 68, 71), and Scolytodes pseudocrassus (66, 69, 72)." figureDoi="http://doi.org/10.5281/zenodo.3944155" httpUri="https://zenodo.org/record/3944155/files/figure.png" pageId="35" pageNumber="36">Figs 64, 67, 70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2DFFAFFF62F9681A8EF969" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="162BAD1CFB2DFFAFFF62F9681A8EF969" blockId="35.[151,1437,1665,2016]" pageId="35" pageNumber="36">
<emphasis id="24E0710EFB2DFFAFFF62F968185DF916" bold="true" box="[151,325,1665,1692]" pageId="35" pageNumber="36">Type material.</emphasis>
<materialsCitation id="A6FCA741FB2DFFAFFEB8F9681875F935" collectingDate="2007-08-10" collectorName="A. C. Proano &amp; A. Barragan" country="Ecuador" elevation="1975" latitude="-79.0" location="Otonga" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Cotopaxi" typeStatus="holotype">
<emphasis id="24E0710EFB2DFFAFFEB8F96818DBF916" bold="true" box="[333,451,1666,1692]" pageId="35" pageNumber="36">
<typeStatus id="C92F13BEFB2DFFAFFEB8F96818A6F916" box="[333,446,1666,1692]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="6E83ED8CFB2DFFAFFE3FF96B1B34F911" box="[458,556,1665,1691]" name="Ecuador" pageId="35" pageNumber="36">Ecuador</collectingCountry>
:
<collectingRegion id="D45063FEFB2DFFAFFDC2F96B1BB9F911" box="[567,673,1665,1691]" country="Ecuador" name="Cotopaxi" pageId="35" pageNumber="36">Cotopaxi</collectingRegion>
,
<location id="134BFBC7FB2DFFAFFD59F96B1A19F911" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB2DFFA8FF62FA1B1DAAFCB1:134BFBC7FB2DFFAFFD59F96B1A19F911" box="[684,769,1665,1691]" country="Ecuador" latitude="-79.0" name="Otonga" pageId="35" pageNumber="36" stateProvince="Cotopaxi">Otonga</location>
,
<quantity id="D16C00F9FB2DFFAFFCF9F96B1A45F916" box="[780,861,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="1.975" pageId="35" pageNumber="36" unit="m" value="1975.0">
<elevation id="9DB94A2FFB2DFFAFFCF9F96B1A45F916" box="[780,861,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="1.975" pageId="35" pageNumber="36" unit="m" value="1975.0">1975m</elevation>
</quantity>
, W
<geoCoordinate id="73A0CBDBFB2DFFAFFC73F96B1D08F916" box="[902,1040,1665,1692]" degrees="79" direction="south" minutes="00" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="00" value="-79.0">79°0000 S</geoCoordinate>
00°2500,
<date id="622A8BDCFB2DFFAFFB6DF96B1C3EF911" box="[1176,1318,1665,1691]" pageId="35" pageNumber="36" value="2007-08-10">
<collectingDate id="726E7234FB2DFFAFFB6DF96B1C3EF911" box="[1176,1318,1665,1691]" pageId="35" pageNumber="36" value="2007-08-10">10.Aug2007</collectingDate>
</date>
,
<collectorName id="BB61C8CAFB2DFFAFFAC5F96B19A7F94A" pageId="35" pageNumber="36">A.C. Proaño</collectorName>
&amp;
<collectorName id="BB61C8CAFB2DFFAFFF14F94C1875F935" box="[225,365,1701,1728]" pageId="35" pageNumber="36">A. Barragán</collectorName>
</materialsCitation>
.
<materialsCitation id="A6FCA741FB2DFFAFFE82F94C1A06F935" box="[375,798,1701,1728]" collectingDate="2007-08-10" collectorName="A. C. Proano &amp; A. Barragan" country="Ecuador" elevation="1975" latitude="-79.0" location="Otonga" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Cotopaxi" typeStatus="paratype">
<emphasis id="24E0710EFB2DFFAFFE82F94C18E0F94A" bold="true" box="[375,504,1702,1728]" pageId="35" pageNumber="36">
<typeStatus id="C92F13BEFB2DFFAFFE82F94C18EBF94A" box="[375,499,1702,1728]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as holotype (1)
</materialsCitation>
;
<materialsCitation id="A6FCA741FB2DFFAFFCDFF94C1D3BF935" box="[810,1059,1701,1728]" collectingDate="2007-08-10" collectorName="A. C. Proano &amp; A. Barragan" country="Ecuador" elevation="1975" latitude="-79.0" location="Otonga" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Cotopaxi" typeStatus="paratype">same data as holotype</materialsCitation>
, except
<date id="622A8BDCFB2DFFAFFB88F94F1C10F935" box="[1149,1288,1701,1727]" pageId="35" pageNumber="36" value="2007-07-11">11. Jul 2007</date>
, A.C. Proaño (1).
<materialsCitation id="A6FCA741FB2DFFAFFF33F9231A8BF969" box="[198,915,1737,1764]" collectionCode="QCAZ, USNM" country="Ecuador" pageId="35" pageNumber="36" specimenCount="2" stateProvince="Cotopaxi" typeStatus="holotype">
<typeStatus id="C92F13BEFB2DFFAFFF33F9231836F969" box="[198,302,1737,1763]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
and one
<typeStatus id="C92F13BEFB2DFFAFFE61F92018ECF96E" box="[404,500,1738,1764]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
deposited in
<collectionCode id="708535D9FB2DFFAFFD7FF9231BC1F969" box="[650,729,1737,1763]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="35" pageNumber="36" type="Museum">QCAZ</collectionCode>
, 1 PT in
<collectionCode id="708535D9FB2DFFAFFCCBF9231A8BF969" box="[830,915,1737,1763]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">USNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2DFFAFFF32F9071A40F8F9" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="162BAD1CFB2DFFAFFF32F9071A40F8F9" blockId="35.[151,1437,1665,2016]" pageId="35" pageNumber="36">
<emphasis id="24E0710EFB2DFFAFFF32F907185AF882" bold="true" box="[199,322,1773,1800]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
Interstriae 10 elevated to level of ventrite 1. Protibiae without an additional mesal tooth. Pronotum smooth. Elytra with erect interstrial bristles and fine strial and interstrial setae. Distinguished from
<taxonomicName id="D194D69FFB2DFFAFFB2EF8F819C1F8C5" authority="(Eggers, 1928)" baseAuthorityName="Eggers" baseAuthorityYear="1928" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="24E0710EFB2DFFAFFB2EF8F81C20F8A1" box="[1243,1336,1810,1836]" italics="true" pageId="35" pageNumber="36">S. minor</emphasis>
(Eggers, 1928)
</taxonomicName>
by longer interstrial and strial setae, and from
<taxonomicName id="D194D69FFB2DFFAFFD12F8DC1AC6F8C5" authority="(Eggers, 1943)" baseAuthorityName="Eggers" baseAuthorityYear="1943" box="[743,990,1845,1872]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="24E0710EFB2DFFAFFD12F8DC1A28F8C5" box="[743,816,1846,1872]" italics="true" pageId="35" pageNumber="36">S. ater</emphasis>
(Eggers, 1943)
</taxonomicName>
and similar species by the regular rows of strial and interstrial setae and the oval and stout body shape.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2DFFA8FF32F8971DD2FDB5" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="162BAD1CFB2DFFA8FF32F8971C2BFD91" blockId="35.[151,1437,1665,2016]" lastBlockId="36.[151,1437,152,827]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="24E0710EFB2DFFAFFF32F89718B8F812" bold="true" box="[199,416,1917,1944]" pageId="35" pageNumber="36">Description, male.</emphasis>
Length 1.72.0 mm, 2.02.1 × as long as wide; colour black.
<emphasis id="24E0710EFB2DFFAFFBACF8941D81F812" bold="true" box="[1113,1177,1918,1944]" pageId="35" pageNumber="36">Head</emphasis>
. Eyes entire, separated above by 2.02.2 × their width. Frons convex, flattened near epistoma, surface reticulated, with moderately coarse punctures scattered from epistoma to vertex; vestiture consisting of scant, short setae near and on epistoma. An- tennal club with segment 1 and 2 corneous, two transverse sutures marked by light setae. Funiculus 5-segmented.
<emphasis id="24E0710EFB2AFFA8FF62FF571808FF5D" bold="true" box="[151,272,189,215]" pageId="36" pageNumber="37">Pronotum</emphasis>
reticulated, dull, with dense, deep punctures reaching anterior margin. Vestiture consisting of very fine, short, setae, and approximately 8 longer, more erect setae (4-2-2).
<emphasis id="24E0710EFB2AFFA8FC6EFF0B1AFFFF71" bold="true" box="[923,999,225,251]" pageId="36" pageNumber="37">Elytra</emphasis>
smooth, shiny; striae regular, not impressed, punctures deep, spaced by their diameter; interstrial punctures nearly as large, in regular rows except slightly confused along suture; interstriae 10 elevated to level of ventrite 1. Vestiture consisting of erect or slightly curved interstrial setae, and rows of finer and shorter, semirecumbent, strial and interstrial setae.
<emphasis id="24E0710EFB2AFFA8FB19FEA71C33FEED" bold="true" box="[1260,1323,333,359]" pageId="36" pageNumber="37">Legs.</emphasis>
Procoxae separated by 0.7 × the width of one procoxa. Mesocoxae separated by 1.01.1 × the width of a procoxa. Protibiae broader towards apex, distal tooth 1 and 2 of equal size, cuticle extended between teeth 2 and 3, with 3 additional small spines or granules along the edge towards tibial base; protibial mucro obtuse. Meso- and metatibiae with 5 and 56 lateral socketed teeth on distal half and third, respectively.
<emphasis id="24E0710EFB2AFFA8FC70FE371D55FE7D" bold="true" box="[901,1101,476,503]" pageId="36" pageNumber="37">Ventral vestiture</emphasis>
. Setae on mes- and metanepisternum trifid or plumose, on anterior part of metasternum trifid or bifid, posterior part with simple setae.
</paragraph>
<paragraph id="162BAD1CFB2AFFA8FF32FDCF1DD2FDB5" blockId="36.[151,1437,152,827]" box="[199,1226,549,575]" pageId="36" pageNumber="37">
<emphasis id="24E0710EFB2AFFA8FF32FDCF183DFDB5" bold="true" box="[199,293,549,575]" pageId="36" pageNumber="37">Female.</emphasis>
Sex undetermined, except one male
<typeStatus id="C92F13BEFB2AFFA8FD37FDCF1A3EFDB5" box="[706,806,549,575]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
. Female likely near identical to male.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2AFFA8FF32FDA31A6DFD45" pageId="36" pageNumber="37" type="key">
<paragraph id="162BAD1CFB2AFFA8FF32FDA31A6DFD45" blockId="36.[151,1437,152,827]" pageId="36" pageNumber="37">
<emphasis id="24E0710EFB2AFFA8FF32FDA31885FDE9" bold="true" box="[199,413,584,611]" pageId="36" pageNumber="37">
Key (
<bibRefCitation id="7205D0EDFB2AFFA8FEF3FDA21888FDE9" author="Wood, S. L." box="[262,400,584,611]" pageId="36" pageNumber="37" refId="ref46297" refString="Wood, S. L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Brigham Young University Press, Provo, Utah, 900 pp." type="book" year="2007">Wood 2007</bibRefCitation>
).
</emphasis>
Goes wrong at couplet 1 due to the combination of a short interstriae 10 and the lack of an additional mesal tooth on protibiae. Related species have an additional tooth on protibiae and if allowed to couplet 59, the new species fits approximately to couplet 78,
<taxonomicName id="D194D69FFB2AFFA8FD09FD7B1D22FD21" ID-CoL="4VT7W" authority="(Eggers, 1943)" baseAuthorityName="Eggers" baseAuthorityYear="1943" box="[764,1082,657,683]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="24E0710EFB2AFFA8FD09FD7B1A9FFD21" box="[764,903,657,683]" italics="true" pageId="36" pageNumber="37">S. punctatus</emphasis>
(Eggers, 1943)
</taxonomicName>
, but differs by a much broader pronotum, more widely spaced setae, and thicker interstrial setae.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2AFFA8FF32FD331AD8FC9D" pageId="36" pageNumber="37" type="etymology">
<paragraph id="162BAD1CFB2AFFA8FF32FD331AD8FC9D" blockId="36.[151,1437,152,827]" pageId="36" pageNumber="37">
<emphasis id="24E0710EFB2AFFA8FF32FD331856FD79" bold="true" box="[199,334,729,755]" pageId="36" pageNumber="37">Etymology.</emphasis>
The Latin name
<taxonomicName id="D194D69FFB2AFFA8FDEDFD301B6BFD79" authorityName="Jordal and Smith" authorityYear="2020" box="[536,627,730,755]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="virgatus">
<emphasis id="24E0710EFB2AFFA8FDEDFD301B6BFD79" box="[536,627,730,755]" italics="true" pageId="36" pageNumber="37">virgatus</emphasis>
</taxonomicName>
is a masculine nominative adjective, meaning striped, referring to the alternate strial and interstrial bands of short and long setae on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB2AFFA8FF32FCCA1DAAFCB1" box="[199,1202,800,827]" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="162BAD1CFB2AFFA8FF32FCCA1DAAFCB1" blockId="36.[151,1437,152,827]" box="[199,1202,800,827]" pageId="36" pageNumber="37">
<emphasis id="24E0710EFB2AFFA8FF32FCCA18E9FCB1" bold="true" box="[199,497,800,827]" pageId="36" pageNumber="37">Biology and distribution.</emphasis>
<materialsCitation id="A6FCA741FB2AFFA8FE0DFCCB1DB6FCB1" box="[504,1198,801,827]" country="Ecuador" location="Ecuador" pageId="36" pageNumber="37" specimenCount="1">
Only known from the
<typeStatus id="C92F13BEFB2AFFA8FD04FCCB1A39FCB1" box="[753,801,801,827]" pageId="36" pageNumber="37">type</typeStatus>
locality at high altitude in
<collectingCountry id="6E83ED8CFB2AFFA8FBBAFCCB1DB6FCB1" box="[1103,1198,801,827]" name="Ecuador" pageId="36" pageNumber="37">Ecuador</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>