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<document id="648999D1CCC73D959585E790DA837035" ID-DOI="10.5281/zenodo.5735795" ID-ISSN="2064-2474" ID-Zenodo-Dep="5735795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638060571117" checkinUser="felipe" docAuthor="Glöer, P. &amp; Georgiev, D." docDate="2012" docId="9E352638FFA3FFEFFDBEFC012155FC50" docLanguage="en" docName="ActaZoolAcadSciH.58.2.193-197.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 58 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Bithynia danubialis Glöer &amp; Georgiev 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="196" masterDocId="620C5E40FFA2FFECFFC2FFDB2564FFEE" masterDocTitle="Bithynia Danubialis, A New Species From The Bulgarian Danube (Gastropoda: Rissooidea: Bithyniidae)" masterLastPageNumber="197" masterPageNumber="193" pageNumber="194" updateTime="1699430515034" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="007702B8264866B5B14FDEE54B533CEA">Bithynia Danubialis, A New Species From The Bulgarian Danube (Gastropoda: Rissooidea: Bithyniidae)</mods:title>
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<mods:name id="4DE45C8563CAAF53E5C51A38E264E049" type="personal">
<mods:role id="8B9AC27F413B8DED058557587D8E1CE0">
<mods:roleTerm id="C5286FDF6471E6584927A51558CA5B66">Author</mods:roleTerm>
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<mods:namePart id="29D3423BB4CAB8DFDE42C7AA657F7651">Glöer, P.</mods:namePart>
<mods:affiliation id="DD724E71C697B21DE1CE93E24315F485">and G, D. G. &amp; Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany e-mail: gloeer @ malaco. de</mods:affiliation>
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<mods:name id="6B28B062861A4642EE11D93988D2C49C" type="personal">
<mods:role id="9DCB3B24009847F339D6003356C7019C">
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<mods:namePart id="83C527CAC8FD3FF8123BDED56988E207">Georgiev, D.</mods:namePart>
<mods:affiliation id="B99CD592B848A74801FC3EF638ED12F6">Department of Ecology and Environmental Conservation, University of Plovdiv Tzar Assen Str. 24, BG- 4000 Plovdiv, Bulgaria, e-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:typeOfResource id="C943B68B9AFF4138C136BCF3B3200D2A">text</mods:typeOfResource>
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<mods:title id="9886686C4B85A68804F7ECF9EA43E887">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:part id="8A254CDABD02419AF75555848404E293">
<mods:date id="7A113F68A8A0D57A5BCC69D78907530B">2012</mods:date>
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<mods:number id="0ED86CD11B9F424D79CD1F01F528385A">2012-06-18</mods:number>
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<mods:number id="F4B2EDF80E3A0ACE9C3B961FA0339948">58</mods:number>
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<mods:number id="73A674C2CA0684D4B746D8591299D3BD">2</mods:number>
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<mods:identifier id="E77E8F8F0E0A790DF5640994F0CE24F0" type="DOI">10.5281/zenodo.5735795</mods:identifier>
<mods:identifier id="3652F7B2FDBA42CD46E4E71C2187491A" type="ISSN">2064-2474</mods:identifier>
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<treatment id="9E352638FFA3FFEFFDBEFC012155FC50" LSID="urn:lsid:plazi:treatment:9E352638FFA3FFEFFDBEFC012155FC50" httpUri="http://treatment.plazi.org/id/9E352638FFA3FFEFFDBEFC012155FC50" lastPageId="3" lastPageNumber="196" pageId="1" pageNumber="194">
<subSubSection id="5E86C4A5FFA3FFEDFDBEFC0126D3FC1B" box="[636,951,986,1013]" pageId="1" pageNumber="194" type="nomenclature">
<paragraph id="1623972EFFA3FFEDFDBEFC0126D3FC1B" blockId="1.[636,951,986,1013]" box="[636,951,986,1013]" pageId="1" pageNumber="194">
<heading id="4D6B2042FFA3FFEDFDBEFC0126D3FC1B" box="[636,951,986,1013]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="194" reason="2">
<taxonomicName id="D19CECADFFA3FFEDFDBEFC012614FC1A" ID-CoL="5WKDF" authority="Glöer &amp; Georgiev, 2012" authorityName="Glöer &amp; Georgiev" authorityYear="2012" box="[636,880,986,1012]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="194" phylum="Mollusca" rank="species" species="danubialis" status="sp. nov.">
<emphasis id="24E84B3CFFA3FFEDFDBEFC012614FC1A" bold="true" box="[636,880,986,1012]" pageId="1" pageNumber="194">Bithynia danubialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FDBF647FFA3FFEDFCBAFC0026D3FC1B" box="[888,951,987,1013]" pageId="1" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5E86C4A5FFA3FFEFFD28FC252155FC50" lastPageId="3" lastPageNumber="196" pageId="1" pageNumber="194" type="description">
<paragraph id="1623972EFFA3FFEDFD28FC252626FBF6" blockId="1.[746,834,1022,1048]" box="[746,834,1022,1048]" pageId="1" pageNumber="194">
<heading id="4D6B2042FFA3FFEDFD28FC252626FBF6" box="[746,834,1022,1048]" centered="true" fontSize="11" level="3" pageId="1" pageNumber="194" reason="3">
(
<figureCitation id="8EA78BABFFA3FFEDFD30FC25265DFBF6" box="[754,825,1022,1048]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="1" pageNumber="194">Fig. 2</figureCitation>
)
</heading>
</paragraph>
<paragraph id="1623972EFFA3FFEDFEBCFB9A24C0FB9D" blockId="1.[317,1270,1089,1139]" pageId="1" pageNumber="194">
Material examined:
<specimenCount id="009A5CA7FFA3FFEDFDFDFB9A271FFBB8" box="[575,635,1089,1110]" pageId="1" pageNumber="194" type="generic">30 ex.</specimenCount>
, DILIAN GEORGIEV leg.
<date id="6222B1EEFFA3FFEDFCA2FB9A26A9FBB8" box="[864,973,1089,1110]" pageId="1" pageNumber="194" value="2009-05-14">14.05.2009</date>
and
<date id="6222B1EEFFA3FFEDFC38FB9A2103FBB8" box="[1018,1127,1089,1110]" pageId="1" pageNumber="194" value="2011-07-21">21.07.2011</date>
(
<specimenCount id="009A5CA7FFA3FFEDFBB1FB9A2192FBB8" box="[1139,1270,1089,1110]" pageId="1" pageNumber="194" type="generic">10 specimens</specimenCount>
dissected).
</paragraph>
<caption id="42E3C7A6FFA3FFEDFE3EF8D72153F8CC" ID-DOI="http://doi.org/10.5281/zenodo.5735797" ID-Zenodo-Dep="5735797" box="[508,1079,1804,1826]" httpUri="https://zenodo.org/record/5735797/files/figure.png" pageId="1" pageNumber="194" startId="1.[508,543,1804,1825]" targetBox="[356,1233,1166,1775]" targetPageId="1">
<paragraph id="1623972EFFA3FFEDFE3EF8D72153F8CC" blockId="1.[508,1079,1804,1826]" box="[508,1079,1804,1826]" pageId="1" pageNumber="194">
<emphasis id="24E84B3CFFA3FFEDFE3EF8D72758F8CF" bold="true" box="[508,572,1804,1825]" pageId="1" pageNumber="194">Fig. 1.</emphasis>
The sampling site of
<taxonomicName id="D19CECADFFA3FFEDFCCEF8D626ACF8CC" authority="Glöer &amp; Georgiev, 2012" authorityName="Glöer &amp; Georgiev" authorityYear="2012" box="[780,968,1805,1826]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="194" phylum="Mollusca" rank="species" species="danubialis" status="sp. nov.">
<emphasis id="24E84B3CFFA3FFEDFCCEF8D626ACF8CC" box="[780,968,1805,1826]" italics="true" pageId="1" pageNumber="194">Bithynia danubialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FDBF647FFA3FFEDFC0DF8D62167F8CC" box="[975,1027,1805,1826]" pageId="1" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
(dot)
</paragraph>
</caption>
<paragraph id="1623972EFFA0FFEEFEBCFE6F26A9FE27" blockId="2.[382,1061,436,486]" box="[382,973,436,457]" pageId="2" pageNumber="195">
<materialsCitation id="A6F49D73FFA0FFEEFEBCFE6F26A9FE27" accessionNumber="ZMH 79308" box="[382,973,436,457]" collectionCode="ZMH" pageId="2" pageNumber="195" specimenCount="1" typeStatus="holotype">
<typeStatus id="C927298CFFA0FFEEFEBCFE6F24BFFE27" box="[382,475,436,457]" pageId="2" pageNumber="195" type="holotype">Holotype</typeStatus>
: shell height
<quantity id="D1643ACBFFA0FFEEFD9CFE6F27CFFE27" box="[606,683,436,457]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="195" unit="mm" value="6.5">6.5 mm</quantity>
, width
<quantity id="D1643ACBFFA0FFEEFD36FE6F2626FE27" box="[756,834,436,457]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="195" unit="mm" value="4.4">4.4 mm</quantity>
,
<collectionCode id="708D0FEBFFA0FFEEFC8FFE6F26E6FE27" box="[845,898,436,457]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="195" type="Museum">ZMH</collectionCode>
79308.
</materialsCitation>
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFE0A2141FE0B" blockId="2.[382,1061,436,486]" box="[382,1061,465,486]" pageId="2" pageNumber="195">
<materialsCitation id="A6F49D73FFA0FFEEFEBCFE0A27C6FE08" accessionNumber="ZMH 79309" box="[382,674,465,486]" collectionCode="ZMH" pageId="2" pageNumber="195" specimenCount="5" typeStatus="paratype">
<typeStatus id="C927298CFFA0FFEEFEBCFE0A24BBFE08" box="[382,479,465,486]" pageId="2" pageNumber="195" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="009A5CA7FFA0FFEEFE28FE0A2778FE08" box="[490,540,465,486]" pageId="2" pageNumber="195" type="generic">5 ex.</specimenCount>
,
<collectionCode id="708D0FEBFFA0FFEEFDE5FE0A2738FE08" box="[551,604,465,486]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="195" type="Museum">ZMH</collectionCode>
79309
</materialsCitation>
,
<materialsCitation id="A6F49D73FFA0FFEEFD6CFE0A263DFE0B" box="[686,857,465,486]" collectionCode="GLOER" pageId="2" pageNumber="195" specimenCount="5" typeStatus="paratype">
<specimenCount id="009A5CA7FFA0FFEEFD6CFE0A27BBFE08" box="[686,735,465,486]" pageId="2" pageNumber="195" type="generic">5 ex.</specimenCount>
coll.
<collectionCode id="708D0FEBFFA0FFEEFCD5FE0A263DFE0B" box="[791,857,465,486]" pageId="2" pageNumber="195">GLÖER</collectionCode>
</materialsCitation>
, rest coll. GEORGIEV
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFDD027C9FDD3" blockId="2.[317,1270,523,573]" pageId="2" pageNumber="195">
Locus typicus: Bulgarian sector of the Danube River at the village of Marten (
<collectingRegion id="D45859CCFFA0FFEEFB41FDD021DAFDCE" box="[1155,1214,523,544]" country="Bulgaria" name="Ruse" pageId="2" pageNumber="195">Russe</collectingRegion>
town area)
<geoCoordinate id="73A8F1E9FFA0FFEEFEB7FDF3276AFDD3" box="[373,526,552,573]" degrees="43" direction="north" minutes="56" orientation="latitude" pageId="2" pageNumber="195" precision="1" seconds="19.4" value="43.938725">N43° 56 19.4”</geoCoordinate>
;
<geoCoordinate id="73A8F1E9FFA0FFEEFDDBFDF327C9FDD3" box="[537,685,552,573]" degrees="26" direction="east" minutes="05" orientation="longitude" pageId="2" pageNumber="195" precision="1" seconds="20.7" value="26.089085">E26° 05 20.7”</geoCoordinate>
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFDBF261CFD90" blockId="2.[317,1270,612,850]" box="[382,888,612,638]" pageId="2" pageNumber="195">Habitat. Littoral zone of the Danube River.</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFD5C27C8FD06" blockId="2.[317,1270,612,850]" pageId="2" pageNumber="195">
Distribution. Only known from the
<typeStatus id="C927298CFFA0FFEEFCD4FD5C262DFD4F" box="[790,841,647,673]" pageId="2" pageNumber="195">type</typeStatus>
locality as yet. But it possibly will be dispersed by the current in most of the Bulgarian and Romanian Danube River sectors down to the Danube Delta.
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFD2A26BBFCBF" blockId="2.[317,1270,612,850]" pageId="2" pageNumber="195">
Associated gastropods.
<taxonomicName id="D19CECADFFA0FFEEFD4FFD2A211FFCE5" authority="(C. PFEIFFER, 1828)" baseAuthorityName="C. PFEIFFER" baseAuthorityYear="1828" box="[653,1147,753,779]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="195" phylum="Mollusca" rank="species" species="danubialis">
<emphasis id="24E84B3CFFA0FFEEFD4FFD2A26EEFCE5" box="[653,906,753,779]" italics="true" pageId="2" pageNumber="195">Theodoxus danubialis</emphasis>
(C. PFEIFFER, 1828)
</taxonomicName>
,
<taxonomicName id="D19CECADFFA0FFEEFB44FD2A2404FCC0" class="Gastropoda" family="Viviparidae" genus="Viviparus" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="195" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="24E84B3CFFA0FFEEFB44FD2A2192FCE5" box="[1158,1270,753,779]" italics="true" pageId="2" pageNumber="195">Viviparus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="D19CECADFFA0FFEEFEB2FCCF262CFCC0" authority="(LINNAEUS, 1758)" baseAuthorityName="LINNAEUS" baseAuthorityYear="1758" box="[368,840,788,814]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="195" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="24E84B3CFFA0FFEEFEB2FCCF2705FCC0" box="[368,609,788,814]" italics="true" pageId="2" pageNumber="195">Bithynia tentaculata</emphasis>
(LINNAEUS, 1758)
</taxonomicName>
,
<taxonomicName id="D19CECADFFA0FFEEFC94FCCF2116FCC0" baseAuthorityName="C. Pfeiffer" baseAuthorityYear="1828" box="[854,1138,788,814]" class="Gastropoda" family="Lithoglyphidae" genus="Lithoglyphus" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="195" phylum="Mollusca" rank="species" species="naticoides">
<emphasis id="24E84B3CFFA0FFEEFC94FCCF2116FCC0" box="[854,1138,788,814]" italics="true" pageId="2" pageNumber="195">Lithoglyphus naticoides</emphasis>
</taxonomicName>
(C. PFEIF- FER, 1828), and
<emphasis id="24E84B3CFFA0FFEEFE3FFCE32661FCBC" box="[509,773,824,850]" italics="true" pageId="2" pageNumber="195">
<taxonomicName id="D19CECADFFA0FFEEFE3FFCE32712FCBC" authorityName="Bourguignat" authorityYear="1877" box="[509,630,824,850]" class="Gastropoda" family="Melanopsidae" genus="Esperiana" kingdom="Animalia" pageId="2" pageNumber="195" phylum="Mollusca" rank="genus">Esperiana</taxonomicName>
daudebartii
</emphasis>
(PREVOST, 1821).
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFCAF215DFC67" blockId="2.[382,1081,884,905]" box="[382,1081,884,905]" pageId="2" pageNumber="195">Etymology: named after the Danube River were the species was found.</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFC7526C3FBBD" blockId="2.[317,1270,942,1165]" pageId="2" pageNumber="195">
Description. Shell glossy and light yellowish to horn-coloured, surface finely striated, 4.55 whorls which are slightly convex with a clear visible but not deep suture, umbilicus closed, the aperture height takes 0.5 of the shell height, edge of aperture sharp, thickened at the columella (
<figureCitation id="8EA78BABFFA0FFEEFB5FFC33218EFC13" box="[1181,1258,1000,1021]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195">Fig. 2.1</figureCitation>
), outer margin of aperture slightly sinuated (
<figureCitation id="8EA78BABFFA0FFEEFD12FBDF2679FBF7" box="[720,797,1028,1049]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195">Fig. 2.2</figureCitation>
). The females are as large as the males, thus a sexual dimorphism is not visible. The operculum is oval and slightly angled at the top (
<figureCitation id="8EA78BABFFA0FFEEFBA7FBFA21D7FBD8" box="[1125,1203,1057,1078]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195">Fig. 2.3</figureCitation>
). Shell height 6.46±
<quantity id="D1643ACBFFA0FFEEFE7AFBE52767FBBD" box="[440,515,1086,1107]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="195" unit="mm" value="0.4">0.4 mm</quantity>
(s = 0.35), width 4.51±
<quantity id="D1643ACBFFA0FFEEFD2FFBE5265CFBBD" box="[749,824,1086,1107]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="195" unit="mm" value="0.4">0.4 mm</quantity>
(s = 0.17).
</paragraph>
<paragraph id="1623972EFFA0FFEEFEBCFB80217DFB63" blockId="2.[317,1270,942,1165]" pageId="2" pageNumber="195">
Morphology of the penis. The distal part of the penis is as long as the penial apendix (
<figureCitation id="8EA78BABFFA0FFEEFB5CFB80218FFB9E" box="[1182,1259,1115,1136]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195">Fig. 2.5</figureCitation>
). The flagellum is approximately four times longer than the penis (
<figureCitation id="8EA78BABFFA0FFEEFC7CFBA3216AFB63" box="[958,1038,1144,1165]" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195">Fig. 2.6</figureCitation>
).
</paragraph>
<caption id="42E3C7A6FFA0FFEEFEFFF90F2192F8E8" ID-DOI="http://doi.org/10.5281/zenodo.5735799" ID-Zenodo-Dep="5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="2" pageNumber="195" startId="2.[317,352,1748,1769]" targetBox="[317,1269,1273,1718]" targetPageId="2">
<paragraph id="1623972EFFA0FFEEFEFFF90F2192F8E8" blockId="2.[317,1270,1748,1827]" pageId="2" pageNumber="195">
<emphasis id="24E84B3CFFA0FFEEFEFFF90F2418F907" bold="true" box="[317,380,1748,1769]" pageId="2" pageNumber="195">Fig. 2.</emphasis>
<taxonomicName id="D19CECADFFA0FFEEFE43F90E2758F904" authority="Glöer &amp; Georgiev, 2012" authorityName="Glöer &amp; Georgiev" authorityYear="2012" box="[385,572,1749,1770]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="195" phylum="Mollusca" rank="species" species="danubialis" status="sp. nov.">
<emphasis id="24E84B3CFFA0FFEEFE43F90E2758F904" box="[385,572,1749,1770]" italics="true" pageId="2" pageNumber="195">Bithynia danubialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FDBF647FFA0FFEEFD80F90E2713F904" box="[578,631,1749,1770]" pageId="2" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-
</paragraph>
</caption>
<paragraph id="1623972EFFA0FFEEFDF7F8D5269AF8CD" blockId="2.[317,1270,1748,1827]" box="[565,1022,1806,1827]" pageId="2" pageNumber="195">pendix, pp = pseudopenis, s= snout, t = tentacle</paragraph>
<paragraph id="1623972EFFA1FFEFFEBCFE6C268BFD26" blockId="3.[317,1270,439,958]" pageId="3" pageNumber="196">
Differentiating features. From
<taxonomicName id="D19CECADFFA1FFEFFD31FE632682FE3C" baseAuthorityName="LINNAEUS" baseAuthorityYear="1758" box="[755,998,440,466]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="24E84B3CFFA1FFEFFD31FE632682FE3C" box="[755,998,440,466]" italics="true" pageId="3" pageNumber="196">Bithynia tentaculata</emphasis>
</taxonomicName>
, which also occurs in
<collectingCountry id="6E8BD7BEFFA1FFEFFEFFFE0024C1FE1B" box="[317,421,475,501]" name="Bulgaria" pageId="3" pageNumber="196">Bulgaria</collectingCountry>
, the new species can be distinguished by the deeper suture, the more convex whorls and the sinuated margin of the aperture, which is in
<taxonomicName id="D19CECADFFA1FFEFFB8CFE252191FDF6" baseAuthorityName="LINNAEUS" baseAuthorityYear="1758" box="[1102,1269,510,536]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="24E84B3CFFA1FFEFFB8CFE252191FDF6" box="[1102,1269,510,536]" italics="true" pageId="3" pageNumber="196">B. tentaculata</emphasis>
</taxonomicName>
straight. The penis morphology is similar to that of
<taxonomicName id="D19CECADFFA1FFEFFC52FDFA241CFDB0" authority="BIELZ, 1853" authorityName="BIELZ" authorityYear="1853" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="transsilvanica">
<emphasis id="24E84B3CFFA1FFEFFC52FDFA21C4FDD5" box="[912,1184,545,571]" italics="true" pageId="3" pageNumber="196">Bithynia transsilvanica</emphasis>
BIELZ, 1853
</taxonomicName>
(
<bibRefCitation id="720DEADFFFA1FFEFFE44FD9F27EAFDB0" author="GLOER, P. &amp; FEHER, Z." box="[390,654,580,606]" pageId="3" pageNumber="196" pagination="285 - 297" refId="ref1751" refString="GLOER, P. &amp; FEHER, Z. (2004) Bithynia leachii (Sheppard, 1823) and Bithynia troschelii (Paasch, 1842) in Hungary (Prosobranchia: Bithyniidae). Annales historico-naturales Musei nationalis hungarici 96: 285 - 297." type="journal article" year="2004">GLÖER &amp; FEHÉR 2004</bibRefCitation>
, =
<taxonomicName id="D19CECADFFA1FFEFFD6CFD9F265FFDB0" baseAuthorityName="Paasch" baseAuthorityYear="1842" box="[686,827,580,606]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="troschelii">
<emphasis id="24E84B3CFFA1FFEFFD6CFD9F265FFDB0" box="[686,827,580,606]" italics="true" pageId="3" pageNumber="196">B. troschelii</emphasis>
</taxonomicName>
), which occurs in
<collectingCountry id="6E8BD7BEFFA1FFEFFBCEFD9F2112FDB0" box="[1036,1142,580,606]" name="Romania" pageId="3" pageNumber="196">Romania</collectingCountry>
(
<bibRefCitation id="720DEADFFFA1FFEFFB46FD9F24ABFD6F" author="GLOER, P. &amp; SIRBU, I." pageId="3" pageNumber="196" pagination="229 - 238" refId="ref1834" refString="GLOER, P. &amp; SIRBU, I. (2005) New freshwater molluscs species found in the Romanian fauna. Heldia 6 (5 - 6): 229 - 238." type="journal article" year="2005">GLÖER &amp; SÎRBU 2005</bibRefCitation>
), but the whorls of
<taxonomicName id="D19CECADFFA1FFEFFD79FDB326E7FD6C" authorityName="BIELZ" authorityYear="1853" box="[699,899,616,642]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="transsilvanica">
<emphasis id="24E84B3CFFA1FFEFFD79FDB326E7FD6C" box="[699,899,616,642]" italics="true" pageId="3" pageNumber="196">B. transsilvanica</emphasis>
</taxonomicName>
are more convex, the suture is deeper and the operculum is rounded. In addition
<taxonomicName id="D19CECADFFA1FFEFFC42FD502178FD4B" authority="Glöer &amp; Georgiev, 2012" authorityName="Glöer &amp; Georgiev" authorityYear="2012" box="[896,1052,651,677]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="danubialis" status="sp. nov.">
<emphasis id="24E84B3CFFA1FFEFFC42FD502178FD4B" box="[896,1052,651,677]" italics="true" pageId="3" pageNumber="196">B. danubialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FDBF647FFA1FFEFFBE1FD502105FD4B" box="[1059,1121,651,677]" pageId="3" pageNumber="196" rank="species">sp. n.</taxonomicNameLabel>
is, however, smaller in height than
<taxonomicName id="D19CECADFFA1FFEFFD81FD75278CFD26" baseAuthorityName="LINNAEUS" baseAuthorityYear="1758" box="[579,744,686,712]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="tentaculata">
<emphasis id="24E84B3CFFA1FFEFFD81FD75278CFD26" box="[579,744,686,712]" italics="true" pageId="3" pageNumber="196">B. tentaculata</emphasis>
</taxonomicName>
and
<taxonomicName id="D19CECADFFA1FFEFFCE3FD75268CFD26" authorityName="BIELZ" authorityYear="1853" box="[801,1000,686,712]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="transsilvanica">
<emphasis id="24E84B3CFFA1FFEFFCE3FD75268CFD26" box="[801,1000,686,712]" italics="true" pageId="3" pageNumber="196">B. transsilvanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1623972EFFA1FFEFFEBCFD0A2697FCBB" blockId="3.[317,1270,439,958]" pageId="3" pageNumber="196">
Notes on the ecology. The species was found on sandy bottom at the littoral zone of the Danube, the river banks were dominated by
<taxonomicName id="D19CECADFFA1FFEFFC08FD2F214BFCE0" box="[970,1071,756,782]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="24E84B3CFFA1FFEFFC08FD2F2161FCE0" box="[970,1029,756,782]" italics="true" pageId="3" pageNumber="196">Salix</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="D19CECADFFA1FFEFFBAFFD2F2191FCE0" box="[1133,1269,756,782]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="24E84B3CFFA1FFEFFBAFFD2F21A9FCE0" box="[1133,1229,756,782]" italics="true" pageId="3" pageNumber="196">Populus</emphasis>
sp.
</taxonomicName>
Maybe this new species survives short periods of desiccation on the banks under wet water vegetation, and mud, like other
<taxonomicName id="D19CECADFFA1FFEFFCEEFCE026A3FCBB" box="[812,967,827,853]" class="Gastropoda" family="Bithyniidae" genus="Bithynia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="24E84B3CFFA1FFEFFCEEFCE026EBFCBB" box="[812,911,827,853]" italics="true" pageId="3" pageNumber="196">Bithynia</emphasis>
spp.
</taxonomicName>
do.
</paragraph>
<paragraph id="1623972EFFA1FFEFFEBCFC852155FC50" blockId="3.[317,1270,439,958]" pageId="3" pageNumber="196">
Remarks. In some females we found specimens with a pseudopenis (
<figureCitation id="8EA78BABFFA1FFEFFB04FC852401FC75" captionStart="Fig" captionStartId="2.[317,352,1748,1769]" captionTargetBox="[317,1269,1273,1718]" captionTargetId="figure-278@2.[317,1269,1273,1718]" captionTargetPageId="2" captionText="Fig. 2. Bithynia danubialis sp. n.: 12 = the shell (holotype), 3 = operculum, 4 = pseudopenis in situ, 5 = penis in situ, 6 = penis with flagellum. Legend: e = eye, fl = flagellum, p = penis, pa = penial ap-" figureDoi="http://doi.org/10.5281/zenodo.5735799" httpUri="https://zenodo.org/record/5735799/files/figure.png" pageId="3" pageNumber="196">Fig. 2.4</figureCitation>
), a very small, not completely developed penis. This phenomenon can be found in species of the genus
<taxonomicName id="D19CECADFFA1FFEFFD8CFC7F2667FC50" authorityName="Paulucci" authorityYear="1878" box="[590,771,932,958]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="196" phylum="Mollusca" rank="genus">
<emphasis id="24E84B3CFFA1FFEFFD8CFC7F2667FC50" box="[590,771,932,958]" italics="true" pageId="3" pageNumber="196">Pseudamnicola</emphasis>
</taxonomicName>
, too (GLÖER
<emphasis id="24E84B3CFFA1FFEFFC61FC7F26BAFC50" box="[931,990,932,958]" italics="true" pageId="3" pageNumber="196">et al.</emphasis>
2010).
</paragraph>
</subSubSection>
</treatment>
</document>