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<document id="EA226595ABEF4F49C8BF74733AD81374" ID-DOI="10.11646/zootaxa.3701.4.5" ID-GBIF-Dataset="bf7d9bbf-b798-4d03-8fb3-c72c138ed14e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217182" ID-ZooBank="A86F25FC-0ECB-4D78-AF44-9B3007C593EF" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460668210448" checkinUser="plazi" docAuthor="Liu, Luxi, Yoshizawa, Kazunori, Li, Fasheng &amp; Liu, Zhiqi" docDate="2013" docId="9C1C87839B44FFB2FF042CF7FE21FE6C" docLanguage="en" docName="zt03701p466.pdf" docOrigin="Zootaxa 3701 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrichadenotecnum multispinosus Liu, Yoshizawa, Li &amp; Liu, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="464" masterDocId="6025FFFB9B46FFB6FF932815FFC1FFE4" masterDocTitle="Atrichadenotecnum multispinosus sp. n. (Psocoptera: Psocidae) from southwestern China, with new synonyms and new combinations from Psocomesites and Clematostigma" masterLastPageNumber="466" masterPageNumber="460" pageNumber="462" updateTime="1698347992017" updateUser="plazi">
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<mods:title id="9FD075652B334468E4BBC29DA9831930">Atrichadenotecnum multispinosus sp. n. (Psocoptera: Psocidae) from southwestern China, with new synonyms and new combinations from Psocomesites and Clematostigma</mods:title>
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<mods:namePart id="B6F832D5C0AF37FD5B4A990B53AEF231">Liu, Luxi</mods:namePart>
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<mods:namePart id="CFF5E0B1DCACE7960D69FD0C01E2FD6C">Yoshizawa, Kazunori</mods:namePart>
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<mods:namePart id="91AE7AA2CB4A637039E7E2F11804C4E1">Li, Fasheng</mods:namePart>
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<mods:namePart id="B93B2F3D745756194F3620416732DA3D">Liu, Zhiqi</mods:namePart>
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<mods:title id="28DC290DF090DD89B77F7B679524E792">Zootaxa</mods:title>
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<mods:date id="AF0FB0619A3E9BB1652736DDB748B282">2013</mods:date>
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<mods:number id="57E6A49BE7E7F93915B0D855DC90A513">3701</mods:number>
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<mods:identifier id="F36BF5EA06770D91DCD99A2C4265C4F8" type="DOI">10.11646/zootaxa.3701.4.5</mods:identifier>
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<treatment id="9C1C87839B44FFB2FF042CF7FE21FE6C" ID-DOI="http://doi.org/10.5281/zenodo.6145735" ID-GBIF-Taxon="119578941" ID-Zenodo-Dep="6145735" LSID="urn:lsid:plazi:treatment:9C1C87839B44FFB2FF042CF7FE21FE6C" httpUri="http://treatment.plazi.org/id/9C1C87839B44FFB2FF042CF7FE21FE6C" lastPageId="4" lastPageNumber="464" pageId="2" pageNumber="462">
<subSubSection id="5CAF651E9B44FFB4FF042CF7FAB1F9A4" pageId="2" pageNumber="462" type="nomenclature">
<paragraph id="140A36959B44FFB4FF042CF7FDBEFB18" blockId="2.[151,639,1250,1276]" box="[151,639,1250,1276]" pageId="2" pageNumber="462">
<heading id="4F4281F99B44FFB4FF042CF7FDBEFB18" bold="true" box="[151,639,1250,1276]" fontSize="11" level="1" pageId="2" pageNumber="462" reason="1">
<emphasis id="26C1EA879B44FFB4FF042CF7FDBEFB18" bold="true" box="[151,639,1250,1276]" pageId="2" pageNumber="462">
<taxonomicName id="D3B54D169B44FFB4FF042CF7FDF5FB18" ID-CoL="JKFF" box="[151,564,1250,1276]" class="Insecta" family="Psocidae" genus="Atrichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="462" phylum="Arthropoda" rank="species" species="multispinosus" status="sp. nov.">
<emphasis id="26C1EA879B44FFB4FF042CF7FDF5FB18" bold="true" box="[151,564,1250,1276]" italics="true" pageId="2" pageNumber="462">Atrichadenotecnum multispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DF257FC9B44FFB4FDAF2CF7FDBEFB18" box="[572,639,1250,1276]" pageId="2" pageNumber="462" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="140A36959B44FFB4FF042D3CFAB1F9A4" blockId="2.[151,1436,1321,2038]" pageId="2" pageNumber="462">
<emphasis id="26C1EA879B44FFB4FF042D3CFEA6FAA6" bold="true" box="[151,359,1321,1346]" pageId="2" pageNumber="462">Male. Coloration</emphasis>
(in alcohol). Body generally brown. Compound eyes grayish black; ocelli brown with dark brown ocellar field. Rest of head brown with dark brown markings. Antennae dark brown except scape, pedicel and first flagellar segment yellowish. Maxillary palpi dark brown. Thorax dark brown, with membranous regions paler. Legs brown, with margin of femora, distal part of tibiae, and tarsi dark brown. Fore wing (
<figureCitation id="8C8E2A109B44FFB4FB252D80FB38FA49" box="[1206,1273,1429,1454]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Fig. 1</figureCitation>
) hyaline with brown markings; basal region dark brown; nodal band continuous, without distinct spots; pterostigma entirely pigmented, without hyaline spots basally; submarginal markings continuous, ranging from cell R3 to middle of areola postica.
<collectingCountry id="6CA276059B44FFB4FED12E11FEBBF9F9" box="[322,378,1540,1565]" name="India" pageId="2" pageNumber="462">Hind</collectingCountry>
wing (
<figureCitation id="8C8E2A109B44FFB4FE582E11FDCEF9F8" box="[459,527,1540,1565]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Fig. 2</figureCitation>
) hyaline with pale brownish along anterior margin; veins brown except in basal regions white. Abdominal segments creamy brown, with brown irregular markings; apical regions dark brown.
</paragraph>
</subSubSection>
<subSubSection id="5CAF651E9B44FFB2FF542E59FE51FF54" lastPageId="4" lastPageNumber="464" pageId="2" pageNumber="462" type="description">
<paragraph id="140A36959B44FFB4FF542E59FAB4F84A" blockId="2.[151,1436,1321,2038]" pageId="2" pageNumber="462">
<emphasis id="26C1EA879B44FFB4FF542E59FEA1F981" bold="true" box="[199,352,1612,1637]" pageId="2" pageNumber="462">Morphology.</emphasis>
Fore wing membranous, glabrous; Sc ending free in membrane, Rs and M fused for a distance, slightly shorter in length than 1st section of Rs; pterostigma triangular with apex moderately rounded; distal closure of discoidal cell concave; areola postica triangular, with short roof almost of equal length as roof of cell M3; radial fork branches diverge at an angle less than 90°.
<collectingCountry id="6CA276059B44FFB4FC9D2EAFFC87F937" box="[782,838,1722,1747]" name="India" pageId="2" pageNumber="462">Hind</collectingCountry>
wing without marginal setae between R2+3 and R4+5. Phallosome (
<figureCitation id="8C8E2A109B44FFB4FEBA2EF4FE55F91E" box="[297,404,1761,1786]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Figs. 3, 4</figureCitation>
) much longer than wide, gradually tapered apically, its left margin almost straight and right margin strongly curved to the left, apex with tiny projections. Hypandrium (
<figureCitation id="8C8E2A109B44FFB4FC6D2F10FBADF8FA" box="[1022,1132,1797,1822]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Figs. 5, 6</figureCitation>
) almost symmetrical, with five developed spines dorso-medially (two in left and three in right); pair of serrated ventral projections strongly projected dorso-laterally; distal margin with pair of processes bearing with fine spines. Clunium (
<figureCitation id="8C8E2A109B44FFB4FA9C2F58FA96F882" box="[1295,1367,1869,1894]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Fig. 7</figureCitation>
) with dorsal margin straight and slightly extended posteriorly. Epiproct (
<figureCitation id="8C8E2A109B44FFB4FC072F64FC1BF86E" box="[916,986,1905,1930]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1900,1923]" captionTargetBox="[190,1403,238,1861]" captionTargetId="figure@3.[151,1436,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 7. Atrichadenotecnum multispinosus sp. n., male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view." httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="2" pageNumber="462">Fig. 7</figureCitation>
) with almost straight posterior margin, unsclerotized medially; paraprocts broad, with a strong apical spine, sensory fields with about 28 trichobothria.
</paragraph>
<paragraph id="140A36959B44FFB4FF542FACFE25F812" blockId="2.[151,1436,1321,2038]" pageId="2" pageNumber="462">
<emphasis id="26C1EA879B44FFB4FF542FACFEBCF836" bold="true" box="[199,381,1977,2002]" pageId="2" pageNumber="462">Measurements.</emphasis>
Bw: 3935, B: 2267, FW: 3233, HW: 2367; F: 573, T: 1253, t1: 393; t2: 146, f1: 492, f2: 372, f3: 338, Mx4: 121, IO/d: 705.
</paragraph>
<caption id="40CA661D9B45FFB5FF042F79FA5DF845" httpUri="https://zenodo.org/record/217183/files/figure.png" pageId="3" pageNumber="463" targetBox="[190,1403,238,1861]" targetPageId="3">
<paragraph id="140A36959B45FFB5FF042F79FA5DF845" blockId="3.[151,1436,1900,1953]" pageId="3" pageNumber="463">
<emphasis id="26C1EA879B45FFB5FF042F79FEFBF865" bold="true" box="[151,314,1900,1923]" pageId="3" pageNumber="463">FIGURES 17.</emphasis>
<taxonomicName id="D3B54D169B45FFB5FED32F79FD59F866" box="[320,664,1900,1922]" class="Insecta" family="Psocidae" genus="Atrichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="463" phylum="Arthropoda" rank="species" species="multispinosus" status="sp. nov.">
<emphasis id="26C1EA879B45FFB5FED32F79FD59F866" box="[320,664,1900,1922]" italics="true" pageId="3" pageNumber="463">Atrichadenotecnum multispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DF257FC9B45FFB5FD0D2F78FD16F866" box="[670,727,1901,1922]" pageId="3" pageNumber="463" rank="species">
<emphasis id="26C1EA879B45FFB5FD0D2F78FD16F866" bold="true" box="[670,727,1901,1922]" pageId="3" pageNumber="463">sp. n.</emphasis>
</taxonomicNameLabel>
, male. (1) Fore wing. (2) Hind wing. (3) Phallosome, lateral view. (4) Phallosome, ventral view. (5) Hypandrium, lateral view. (6) Hypandrium, posterior view. (7) Epiproct &amp; paraproct, dorsal view.
</paragraph>
</caption>
<paragraph id="140A36959B42FFB2FF542882FE51FF54" blockId="4.[151,1436,151,392]" box="[199,400,151,176]" pageId="4" pageNumber="464">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="5CAF651E9B42FFB2FF5428A9FE96FF1C" pageId="4" pageNumber="464" type="materials_examined">
<paragraph id="140A36959B42FFB2FF5428A9FE96FF1C" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="464">
<emphasis id="26C1EA879B42FFB2FF5428A9FE70FF31" bold="true" box="[199,433,188,213]" pageId="4" pageNumber="464">Material examined.</emphasis>
<typeStatus id="CB0E88379B42FFB2FE2B28A9FDE1FF31" box="[440,544,188,213]" pageId="4" pageNumber="464" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="6CA276059B42FFB2FDFE28A9FD0BFF31" box="[621,714,188,213]" name="China" pageId="4" pageNumber="464">
<emphasis id="26C1EA879B42FFB2FDFE28A9FD0BFF31" bold="true" box="[621,714,188,213]" pageId="4" pageNumber="464">CHINA</emphasis>
</collectingCountry>
: Guizhou Province, Kuankuoshui Nature Reserve,
<date id="600B10559B42FFB2FA9E28A9FA56FF31" box="[1293,1431,188,213]" pageId="4" pageNumber="464" value="2010-08-13">13.viii.2010</date>
, Luxi Liu (CAU).
</paragraph>
</subSubSection>
<subSubSection id="5CAF651E9B42FFB2FF542911FB0AFEF9" box="[199,1227,260,285]" pageId="4" pageNumber="464" type="etymology">
<paragraph id="140A36959B42FFB2FF542911FB0AFEF9" blockId="4.[151,1436,151,392]" box="[199,1227,260,285]" pageId="4" pageNumber="464">
<emphasis id="26C1EA879B42FFB2FF542911FE8FFEF9" bold="true" box="[199,334,260,285]" pageId="4" pageNumber="464">Etymology.</emphasis>
The species name refers to the prominent developed spines on the hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="5CAF651E9B42FFB2FF542932FE21FE6C" pageId="4" pageNumber="464" type="discussion">
<paragraph id="140A36959B42FFB2FF542932FE21FE6C" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="464">
<emphasis id="26C1EA879B42FFB2FF542932FEFAFEA4" bold="true" box="[199,315,295,320]" pageId="4" pageNumber="464">Remarks.</emphasis>
The new species appears to be closely related to
<taxonomicName id="D3B54D169B42FFB2FCE22932FAAAFEA4" authority="Thornton, 1961" authorityName="Thornton" authorityYear="1961" box="[881,1387,295,320]" class="Insecta" family="Psocidae" genus="Atrichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="464" phylum="Arthropoda" rank="species" species="nudum">
<emphasis id="26C1EA879B42FFB2FCE22932FB61FEA4" box="[881,1184,295,320]" italics="true" pageId="4" pageNumber="464">Atrichadenotecnum nudum</emphasis>
(Thornton, 1961)
</taxonomicName>
and
<taxonomicName id="D3B54D169B42FFB2FF042959FE3CFE80" authority="Yoshizawa, 1998" authorityName="Yoshizawa" authorityYear="1998" box="[151,509,332,357]" class="Insecta" family="Psocidae" genus="Atrichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="464" phylum="Arthropoda" rank="species" species="ryukyuense">
<emphasis id="26C1EA879B42FFB2FF042959FEF5FE81" box="[151,308,332,357]" italics="true" pageId="4" pageNumber="464">A. ryukyuense</emphasis>
Yoshizawa, 1998
</taxonomicName>
, but can be distinguished from the two species by hypandrium characters with five strong spines dorso-medially.
</paragraph>
</subSubSection>
</treatment>
</document>