262 lines
22 KiB
XML
262 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.585.7503" ID-GBIF-Dataset="284e15e6-931f-4863-b037-68a83c8b53a8" ID-PMC="PMC4857038" ID-Pensoft-Pub="1313-2970-585-51" ID-PubMed="27199604" ID-ZBK="B914D8CF92A14C948EDC7CE8B0202076" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-585-51" ModsDocOrigin="ZooKeys 585" ModsDocTitle="Megastigmus seed chalcids (Hymenoptera, Torymidae) radiated much more on Angiosperms than previously considered. I- Description of 8 new species from Kenya, with a key to the females of Eastern and Southern Africa" checkinTime="1461901611862" checkinUser="pensoft" docAuthor="Roques, Alain, Copeland, Robert S., Soldati, Laurent, Denux, Olivier & Auger-Rozenberg, Marie-Anne" docDate="2016" docId="9B4D902763104BB35610798C95FCD40E" docLanguage="en" docName="ZooKeys 585: 51-124" docOrigin="ZooKeys 585" docSource="http://dx.doi.org/10.3897/zookeys.585.7503" docTitle="Megastigmus grewianae Roques & Copeland, sp. n." docType="treatment" docUuid="5AA95119-9BC7-4DF3-AE4E-3CB3409147EB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="90" masterDocId="FF812555FFDD75799D3DFF99922B9D29" masterDocTitle="Megastigmus seed chalcids (Hymenoptera, Torymidae) radiated much more on Angiosperms than previously considered. I- Description of 8 new species from Kenya, with a key to the females of Eastern and Southern Africa" masterLastPageNumber="124" masterPageNumber="51" pageNumber="86" updateTime="1668163066542" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Megastigmus seed chalcids (Hymenoptera, Torymidae) radiated much more on Angiosperms than previously considered. I- Description of 8 new species from Kenya, with a key to the females of Eastern and Southern Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Roques, Alain</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Copeland, Robert S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soldati, Laurent</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Denux, Olivier</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Auger-Rozenberg, Marie-Anne</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>585</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>124</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.585.7503</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.585.7503</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-585-51</mods:identifier>
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<mods:identifier type="ZBK">B914D8CF92A14C948EDC7CE8B0202076</mods:identifier>
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<mods:identifier type="ZooBank">B914D8CF92A14C948EDC7CE8B0202076</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127898981" LSID="urn:lsid:zoobank.org:act:5AA95119-9BC7-4DF3-AE4E-3CB3409147EB" httpUri="http://treatment.plazi.org/id/9B4D902763104BB35610798C95FCD40E" lastPageId="39" lastPageNumber="90" pageId="35" pageNumber="86">
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<subSubSection pageId="35" pageNumber="86" type="multiple">
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<paragraph pageId="35" pageNumber="86">Taxon classification Animalia Hymenoptera Torymidae</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="86" type="nomenclature">
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<paragraph pageId="35" pageNumber="86">
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<taxonomicName LSID="http://zoobank.org/5AA95119-9BC7-4DF3-AE4E-3CB3409147EB" authority="Roques & Copeland" class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus grewianae" order="Hymenoptera" pageId="35" pageNumber="86" phylum="Arthropoda" rank="species" species="grewianae">Megastigmus grewianae Roques & Copeland</taxonomicName>
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<taxonomicNameLabel pageId="35" pageNumber="86">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="86" type="type material">
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<paragraph pageId="35" pageNumber="86">Type material.</paragraph>
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<paragraph pageId="35" pageNumber="86">
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Holotype ♀, Kenya, Coast Province, Arabuko-Sokoke forest,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-3.2997">3.2997°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="39.9869">39.9869°E</geoCoordinate>
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, 55 m, 17 Feb 2000, ex.
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia stuhlmanii" order="Malvales" pageId="35" pageNumber="86" phylum="Tracheophyta" rank="species" species="stuhlmanii">Grewia stuhlmanii</taxonomicName>
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fruits, Coll. N° Kip-356, R Copeland leg. (NMKE)
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</paragraph>
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<paragraph pageId="35" pageNumber="86">
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Paratypes: Kenya, 1♀, 1♂, same collection data as holotype (RSC); 2♀♀, 1♂, same collection data as holotype (ARC); 1♀, 7♂♂, Eastern Province, Tsavo,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-2.676">2.6760°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="38.3325">38.3325°E</geoCoordinate>
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, 638 m, 20 Feb 2000, ex. fruits of
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia tephrodermis" order="Malvales" pageId="35" pageNumber="86" phylum="Tracheophyta" rank="species" species="tephrodermis">Grewia tephrodermis</taxonomicName>
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, A&M Coll. N°514, R. Copeland leg. (1♀, 3♂♂ NMKE, 4♂♂ ICIPE); 1♀, Coast Province, Kasigau Mountain,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-3.827">3.82700°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="38.64875">38.64875°E</geoCoordinate>
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, 1065 m, Malaise trap in campsite clearance, 19 May to 2 Jun 2011, R. Copeland leg. (SAMC); 1♀, Eastern Province, Kasaala area,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-2.07486">2.07486°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="38.2253">38.22530°E</geoCoordinate>
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, 741 m, Malaise trap, 28 Nov. to 4 Dec 2013, leg. J. Bukhebi & R. Copeland (ICIPE).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="86" type="materials_examined">
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<paragraph pageId="35" pageNumber="86">Additional material in alcohol.</paragraph>
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<paragraph pageId="35" pageNumber="86">
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Kenya, 7♀♀, 1♂, same as holotype (ARC); 1♀, Eastern Province, Tsavo,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-2.676">2.6760°S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="38.3325">38.3325°E</geoCoordinate>
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, 638 m, 20 Feb 2000, ex. fruits of
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia tephrodermis" order="Malvales" pageId="35" pageNumber="86" phylum="Tracheophyta" rank="species" species="tephrodermis">Grewia tephrodermis</taxonomicName>
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, Coll. #514, R. Copeland leg. (ARC)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="37" lastPageNumber="88" pageId="35" pageNumber="86" type="description">
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<paragraph pageId="35" pageNumber="86">Description.</paragraph>
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<paragraph pageId="35" pageNumber="86">Holotype ♀. Body length (without ovipositor) 4.1mm; length of ovipositor sheaths 3.0 mm. Body colour orange with some darker patterns (Figures 81-83). Head colour orange with a very narrow occipital black line and outer part of ocelli black. Pilosity on face pale, dark on dorsum of head but hairs not strong (Figure 85). Antenna brownish except scape and pedicel yellowish (Figure 85). Pronotum orange with a conspicuous longitudinal black stripe running for 7/8 of the lateral part of the pronotum, interrupted just before the suture with mid-lobe of mesoscutum (Figures 82-83). Remainder of thorax orange except black spots on the posterior part of axilla at wing insertion. Pilosity on thorax black but hairs not strong; a few on pronotum; 5 pairs on latero-posterior part of mid-lobe of mesoscutum; 4 pairs on lateral lobe of mesoscutum along suture with mid-lobe of mesoscutum; 4 pairs on axilla. Scutellum orange with 5 pairs of lateral hairs with conspicuous insertion dots (Figures 83-84).</paragraph>
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<paragraph lastPageId="36" lastPageNumber="87" pageId="35" pageNumber="86">
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Legs entirely pale yellow except claws brown, coxae with conspicuous hair dots. Forewing stigma brown without infuscation; basal cell closed, with 13 setae on disc; basal setal line with 6 long setae; costal setal line with 16 small setae, costal cell with 11 setae in 2 rows (Figure 87). Propodeum dark orange, with a small oblique brownish spot around the spiracle; not wrinkled transversally; propodeum hairs dark. Gaster orange with two lateral rows composed of 5 brown elongated spots, one on each of the first five segments, placed latero-medially, the third spot shaped like a comma, larger
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<pageBreakToken pageId="36" pageNumber="87" start="start">than</pageBreakToken>
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the others, followed in size by the 4th one. Ovipositor sheaths black, 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as gaster, 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as body (Figure 82).
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</paragraph>
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<paragraph lastPageId="37" lastPageNumber="88" pageId="36" pageNumber="87">
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Head rounded, width: height ratio: 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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(Figure 85); POL: OOL: 1.3; torulus ca. 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; inter-antennal area much smaller (0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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) than torulus width; scrobe elongate, ca. 3.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide. Scape elongate, 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as combined length of pedicel, anellus and F1 (Figure 86); scape 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as combined length
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<pageBreakToken pageId="37" pageNumber="88" start="start">of</pageBreakToken>
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pedicel, anellus, F1 and F2; Pedicel elongate, 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than F1; anellus subquadrate; F1 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, shorter than the other segments, 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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shorter than F2; F2 1.9
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; following funicular segments same as F2, except F7 and F8 tending to subquadrate, with F7 1.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide. Pronotum with 8 very strong transverse carinae (Figure 84), mid- lobe of mesoscutum with 7 coarse transverse carinae extending onto lateral lobes; axillae with 3 weaker, longitudinal, carinae. Mid-lobe of mesoscutum 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as scutellum length. Scutellum 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with transverse, irregular striae anteriorly; frenum smooth, 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as scutellum length. Stigma oval, about 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide (Figure 87); upper part of stigmal vein very short, 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as stigma length; uncus comparatively elongate, 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than upper part of stigmal vein; marginal vein 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as postmarginal vein. Propodeum reticulate with 2 oblique carinae diverging from the center of the anterior suture (Figure 84).
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</paragraph>
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<paragraph pageId="37" pageNumber="88">Male. Body length 4.2 mm; Body colour, pilosity and black patterns similar to female (Figures 88-91) except gaster with 6 rows of brown spots laterally on dorsum; those on the first two segments elongate with the second longer, the 3 following more rounded, comma-like, and the last a simple spot (Figure 88). Forewing stigma brown without infuscation; basal cell closed, with 21 setae on disc; basal setal line with 7 setae; costal setal line with 21 setae; costal cell with 20 setae arranged in 3 rows (Figure 94). Thorax pilosity as in female.</paragraph>
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<paragraph pageId="37" pageNumber="88">
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Head subquadrate, width: height ratio: 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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(Figure 91); POL: OOL 1.3; torulus ca. 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; inter-antennal area short, 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as broad as torulus width; scrobe elongate, ca. 3.9
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; eyes protruding. Scape as long as combined length of pedicel, anellus and F1 (Figure 92); scape 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as combined length of pedicel, anellus, F1 and F2; pedicel 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than F1; anellus subquadrate; F1 2.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; F2 1.9
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; following funicular segments similar to F2, only F7 tending to suquadrate (1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide). Pronotum with coarse transverse carinae as in female but irregular; some carinae interrupted in their middle on mid-lobe of mesoscutum (Figure 93); axilla with curved longitudinal striae; mid-lobe of mesoscutum 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as scutellum length. Scutellum 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with transverse striae in the anterior part only, then smooth; frenum entirely smooth, 0.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as scutellum length (Figure 93). Forewing stigma subquadrate, 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide (Figure 94); upper part of stigmal vein short, 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as stigma length; uncus as long as upper part of stigmal vein; marginal vein 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as postmarginal vein. Propodeum with several oblique, irregular carinae (Figure 93). Genitalia with aedeagus relatively elongate, its part above digitus about 1.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as digitus length; digitus enlarged at its extremity, only twice as long than its maximum width, with 3 teeth (Figure 95).
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</paragraph>
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<paragraph pageId="37" pageNumber="88">Variation. Females range in length from 3.5 to 4.0mm. The number of lateral spots on gaster varies from 6 (with two on T3) to 2 (only the large ones remaining), the 3 last lateral spots sometimes fused into a line. Pilosity on thorax paler in one female paratype. Males range in length from 4.1 to 4.5 mm. The spots on gaster sometimes larger, the pairs of lateral hairs on scutellum ranging up to 6 in both sexes.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="39" lastPageNumber="90" pageId="38" pageNumber="89" type="host plants">
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<paragraph pageId="38" pageNumber="89">
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<pageBreakToken pageId="38" pageNumber="89" start="start">Host</pageBreakToken>
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plants.
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</paragraph>
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<paragraph lastPageId="39" lastPageNumber="90" pageId="38" pageNumber="89">
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia stuhlmannii" order="Malvales" pageId="38" pageNumber="89" phylum="Tracheophyta" rank="species" species="stuhlmannii">Grewia stuhlmannii</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia tephrodermis" order="Malvales" pageId="38" pageNumber="89" phylum="Tracheophyta" rank="species" species="tephrodermis">Grewia tephrodermis</taxonomicName>
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(
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<taxonomicName family="Malvaceae" lsidName="Anacardiaceae" pageId="38" pageNumber="89" rank="family">Malvaceae</taxonomicName>
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). Probably a seed feeder, based on its place in the molecular phylogeny of
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<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus" order="Hymenoptera" pageId="38" pageNumber="89" phylum="Arthropoda" rank="genus">Megastigmus</taxonomicName>
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spp. (Figure 14). Sequencing of four specimens from Arabuko-Sokoke forest showed they differed by less than 0.3% in COI, confirming the validity of the single specimen that we used
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<pageBreakToken pageId="39" pageNumber="90" start="start">in</pageBreakToken>
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the analyses, even if the position remains unresolved. A large number of species of
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia" order="Malvales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="genus">Grewia</taxonomicName>
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coexist in East Africa (Brink and Achigan-Dako 2012) and it would be interesting to find if other
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<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus" order="Hymenoptera" pageId="39" pageNumber="90" phylum="Arthropoda" rank="genus">Megastigmus</taxonomicName>
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species or sub-species are able to attack their seeds.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="90" type="distribution">
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<paragraph pageId="39" pageNumber="90">Distribution.</paragraph>
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<paragraph pageId="39" pageNumber="90">
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Known from dry
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
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/
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<taxonomicName class="Magnoliopsida" family="Burseraceae" genus="Commiphora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commiphora" order="Sapindales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="genus">Commiphora</taxonomicName>
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savanna, moderately wet mid-altitude mountain forest, and bush associated with costal forest. Adults emerged from 2.7% of
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia stuhlmannii" order="Malvales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="species" species="stuhlmannii">Grewia stuhlmannii</taxonomicName>
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fruits and 13% of
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia tephrodermis" order="Malvales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="species" species="tephrodermis">Grewia tephrodermis</taxonomicName>
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fruits (Table 3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="90" type="etymology">
|
||
<paragraph pageId="39" pageNumber="90">Etymology.</paragraph>
|
||
<paragraph pageId="39" pageNumber="90">Named after the genus of its host plant.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="39" pageNumber="90" type="diagnosis">
|
||
<paragraph pageId="39" pageNumber="90">Diagnosis.</paragraph>
|
||
<paragraph pageId="39" pageNumber="90">
|
||
Females are easily separated from those of
|
||
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus copelandi" order="Hymenoptera" pageId="39" pageNumber="90" phylum="Arthropoda" rank="species" species="copelandi">Megastigmus copelandi</taxonomicName>
|
||
, the other species reared from
|
||
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Grewia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Grewia" order="Malvales" pageId="39" pageNumber="90" phylum="Tracheophyta" rank="genus">Grewia</taxonomicName>
|
||
fruits, by their significantly larger size (4.1 mm vs. 2.6-2.7) and the relatively longer ovipositor (ca. 1.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than gaster length vs. 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
in
|
||
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus copelandi" order="Hymenoptera" pageId="39" pageNumber="90" phylum="Arthropoda" rank="species" species="copelandi">Megastigmus copelandi</taxonomicName>
|
||
). Females and males are also easily distinguished from the other species of Afrotropical
|
||
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus" order="Hymenoptera" pageId="39" pageNumber="90" phylum="Arthropoda" rank="genus">Megastigmus</taxonomicName>
|
||
by the unique pattern of a longitudinal black stripe extending 7/8 of the length of the side of the pronotum (Figures 82-83, 89-90).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |