179 lines
22 KiB
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179 lines
22 KiB
XML
<document id="AE8C75C8BF2204463CB2B2D07C496D72" ID-CLB-Dataset="35168" ID-DOI="http://dx.doi.org/10.3897/subtbiol.16.4467" ID-GBIF-Dataset="16642a21-060d-4a21-96e0-18cfb3940761" ID-Pensoft-Pub="1314-2615-16-1" ID-ZooBank="C93851B1764545D4B9EBF09586DD56F5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1314-2615-16-1" ModsDocOrigin="Subterranean Biology 16" ModsDocTitle="Four new representatives of the genus Allocyclops Kiefer, 1932 from semi-consolidated subsoil aquifers in Benin (Copepoda, Cyclopoida, Cyclopidae)" checkinTime="1435211255059" checkinUser="pensoft" docAuthor="Fiers, Frank & Lagnika, Moissou" docDate="2015" docId="9ACD6F92DBF3D5C3C50CAB13F8EB09C7" docLanguage="en" docName="SubterBiol 16: 1-36" docOrigin="Subterranean Biology 16" docSource="http://dx.doi.org/10.3897/subtbiol.16.4467" docTitle="Allocyclops sakitii Fiers & Lagnika, 2015, sp. n." docType="treatment" docUuid="316EB4BF-2BA4-406E-B289-81CE29B3C34D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="20" masterDocId="FF94FF809B3DFFA5CE7B96404B5C8160" masterDocTitle="Four new representatives of the genus Allocyclops Kiefer, 1932 from semi-consolidated subsoil aquifers in Benin (Copepoda, Cyclopoida, Cyclopidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="13" updateTime="1701381930107" updateUser="ExternalLinkService">
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<mods:title id="5DFF7C1FD4FFE5872DC7456E0A3830F6">Four new representatives of the genus Allocyclops Kiefer, 1932 from semi-consolidated subsoil aquifers in Benin (Copepoda, Cyclopoida, Cyclopidae)</mods:title>
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<mods:name id="4F4D098E388301871750B112E5BDACE4" type="personal">
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<mods:roleTerm id="502795838F2E5B1B5ADE36AD26140B0A">Author</mods:roleTerm>
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<mods:namePart id="6504611B3D749A77EBA3261C55CB5BB0">Fiers, Frank</mods:namePart>
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<mods:name id="0DA300F3B2D123D34DCDBB3AA41DDB40" type="personal">
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<mods:roleTerm id="1EDAF2F9CBC34B4F8F4AA5B902F768CA">Author</mods:roleTerm>
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<mods:namePart id="7E0FBEFB6038B1E26249CA4098D93988">Lagnika, Moissou</mods:namePart>
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<mods:title id="717EF7BE4712E12A6B0174B594A12E08">Subterranean Biology</mods:title>
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<mods:part id="64B009108B1B436E8982488A48760B72">
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<mods:date id="D8C469271DDC753135FE1CE2D7250F99">2015</mods:date>
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<mods:detail id="067083DD8A545D8CE7FCB8CC2607C4E0" type="volume">
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<mods:number id="E62DB08D82976E31B46E427895ACEACC">16</mods:number>
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<mods:start id="AD37689EA5C511C13411EA3B1E9558DD">1</mods:start>
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<mods:end id="4D41C348218AFE5FE4D527F34904DDC0">36</mods:end>
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<mods:url id="85204C914E4FFC552A4F4BD1A611F20F">http://dx.doi.org/10.3897/subtbiol.16.4467</mods:url>
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<mods:classification id="247549607498FD87F58EFCD7CF5FE0E7">journal article</mods:classification>
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<mods:identifier id="3E98ED04B03D000620C523FA054FDFF3" type="DOI">http://dx.doi.org/10.3897/subtbiol.16.4467</mods:identifier>
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<mods:identifier id="DEC1A2EAC986782FD524227F4C5DA430" type="Pensoft-Pub">1314-2615-16-1</mods:identifier>
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<treatment id="9ACD6F92DBF3D5C3C50CAB13F8EB09C7" ID-GBIF-Taxon="127873577" LSID="urn:lsid:zoobank.org:act:316EB4BF-2BA4-406E-B289-81CE29B3C34D" httpUri="http://treatment.plazi.org/id/9ACD6F92DBF3D5C3C50CAB13F8EB09C7" lastPageId="19" lastPageNumber="20" pageId="12" pageNumber="13">
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<subSubSection id="30AEA0BBAD6C75E5925A0613449E66BF" pageId="12" pageNumber="13" type="multiple">
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<paragraph id="B9A2E0A9F93DFA5CBF8094AF27E97F70" pageId="12" pageNumber="13">Taxon classification Animalia Cyclopoida Cyclopidae</paragraph>
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</subSubSection>
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<subSubSection id="42116D71D44F0417EC06B68E0B1C6F78" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="3748B0122FF5A914F3C202EE33F11FEC" pageId="12" pageNumber="13">
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<taxonomicName id="9C5DDE68064D7A9AED8792754E535911" ID-CoL="BVYZ" LSID="http://zoobank.org/316EB4BF-2BA4-406E-B289-81CE29B3C34D" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops sakitii" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sakitii">Allocyclops sakitii</taxonomicName>
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<taxonomicNameLabel id="1E8917AB10AEA164F18AD69C03D5D6B9" pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
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Figs 4G, H, L, 5I, J, 13, 14, 15
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</paragraph>
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</subSubSection>
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<subSubSection id="1B72F78C687CC5A1BB9D2AB407CEE463" pageId="13" pageNumber="14" type="type locality">
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<paragraph id="37C5717338416EF4957703F5C63DE128" pageId="13" pageNumber="14">
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<pageBreakToken id="5C7BA558C53D3EE514877130E753B9DD" pageId="13" pageNumber="14" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph id="A1C8E093FE5AEBABDD33A019B73D2A40" pageId="13" pageNumber="14">at Addrodji, Lokossa District, Mono Couffo drainage basin: Well P2 (details in Table 1).</paragraph>
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</subSubSection>
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<subSubSection id="C6A2D3797529740C98238EB36F84104E" pageId="13" pageNumber="14" type="type material">
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<paragraph id="D3169FE5DAFB476D18E34040F3158652" pageId="13" pageNumber="14">Type material.</paragraph>
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<paragraph id="4279E5C02F08D71AB2DD3A96C9D0D8A4" pageId="13" pageNumber="14">Female holotype, dissected and mounted on 3 slides, RBINSc COP 10.354; allotype male, dissected and mounted on 2 slides, RBINSc COP 10.356; female paratypes dissected: RBINSc COP 10.355A-C, COP 10.357 A, COP 10. 358 A-C; 14 females and 5 males preserved (RBINSc COP 10.359), all collected 1-12-2012. Additional paratypic material (topotypic), collected 19/06/2012: 185 females, 15 males (preserved material RBINSc COP 10.360).</paragraph>
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</subSubSection>
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<subSubSection id="5268326E0AB145C0F99F50F168B5AC1C" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="539A783FB8CD2A1B69800136139AEA3F" pageId="13" pageNumber="14">Etymology.</paragraph>
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<paragraph id="198B72C7051199D8B58DE8AAE57C4814" pageId="13" pageNumber="14">
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Named after Nestor G. Sakiti who initiated, in collaboration with Claude Boutin, subterranean biology research in
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<normalizedToken id="D6D25F90481073B41A874EBBAD0C1172" originalValue="Bénin">Benin</normalizedToken>
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, and supported the junior author initializing his investigations.
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</paragraph>
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</subSubSection>
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<subSubSection id="E5F91D32F08AF1ABB7D5151CCC697CBB" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="additional material">
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<paragraph id="6D349F88B82C0F9C40FF378885EF7CAF" pageId="13" pageNumber="14">Additional material.</paragraph>
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<paragraph id="D04257E9DA71F481443CFD27E92C4EBB" pageId="13" pageNumber="14">(1) Lokossa, Mono District, Mono-Couffo drainage basin:</paragraph>
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<paragraph id="A9DCE4B4091101B41C8813F60B704FAC" pageId="13" pageNumber="14">- Well P6: 17-06-2012: 1 ♀ (preserved material RBINSc COP 10.364); 06/09/2012: 18 ♀♀ 1 ♂ (preserved material RBINSc COP 10.378); Well PM (about 20 m from well P6, cemented, closed and abandoned, and considered here as identical with well P6): 07/01/2012: 65 ♀♀, 4 ♂♂ (dissected ♀ RBINSc COP 10.370A-C; preserved material RBINSc COP 10.365).</paragraph>
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<paragraph id="8B3554AF33C947F8E8429FB50C6DF9AE" pageId="14" pageNumber="15">
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<pageBreakToken id="859EE21C193F9C933818245297008460" pageId="14" pageNumber="15" start="start">-</pageBreakToken>
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Well P7: 13-06-2012: 74 ♀♀, 4 ♂♂, 2 juveniles (dissected ♀ RBINSc COP 10.369A-C; preserved material RBINSc 10.363);
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</paragraph>
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<paragraph id="9DFADDBB0AEB699C8E412ACF638DF29E" pageId="14" pageNumber="15">- Well P9: 11-06-2012: 116 ♀♀, 11 ♂♂, 9 juveniles (preserved material RBINSc COP 10.361); 30-11-2012: 9 ♀♀, 2 ♂♂ (preserved material RBINSc COP 10.362).</paragraph>
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<paragraph id="470AAED2D1256E7F6B17259A48F162FE" pageId="14" pageNumber="15">
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(2) Abomey-Calavi District,
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<normalizedToken id="4E228DFB211BE33148CAD1AD8047E54F" originalValue="Ouémé">Oueme</normalizedToken>
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drainage basin:
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</paragraph>
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<paragraph id="3F820D13DC39E5B1E3F65955A900B7FB" pageId="14" pageNumber="15">- Well near Campus: 27-02-2013: 24 ♀♀, 3 ♂♂, 1 juvenile (preserved material RBINSc COP 10.366).</paragraph>
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</subSubSection>
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<subSubSection id="7104D98C2F9AC49C017ECF1645A4ECE0" lastPageId="19" lastPageNumber="20" pageId="15" pageNumber="16" type="description">
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<paragraph id="ACDC3B78B41D0ABD621B9520CB84AC5E" pageId="15" pageNumber="16">
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<pageBreakToken id="639EE613DE5554F6DDF0D3019F44FDD5" pageId="15" pageNumber="16" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph id="B490AC54D5FC2A1429057635BA5C1E5A" pageId="15" pageNumber="16">
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Female: Body (Fig. 13A) typically cyclopoid with prominent prosome (1.8-1.9 times urosome length). Genital double-somite widest in anterior third, and as long as widest part. Posterodorsal fringe of prosomites and urosomite 1 narrow and straight. Posterior fringe of genital double-somite and urosomites 3-4 narrow and discretely serrate (Fig. 14A, B). Anal somite with shallow crescent operculum (Fig. 13C). Anal sinus apparently smooth. Posterior margin of anal somite with uninterrupted row of spinules: rather short dorsally, laterally and along outer half of ventral margin, twice as long along medial half of ventral margin (Fig. 14A). Body length: 833
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<normalizedToken id="937556820D6F9E7333FC46E42ADED435" originalValue="μm">μm</normalizedToken>
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(holotype, paratypes ranging from 807-876
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<normalizedToken id="006AD6CE745B7EC07E653AD33381D58C" originalValue="μm">μm</normalizedToken>
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, n=5).
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</paragraph>
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<caption id="71EDC7A88B6CB58BB0D77C31A47DDA66" pageId="15" pageNumber="16">
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<paragraph id="F6FA769C6837E56132371EE9D68CFD43" pageId="15" pageNumber="16">
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Figure 13.
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<taxonomicName id="ECEBD87487C82055C47A68C61436AB24" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops sakitii" order="Cyclopoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sakitii">Allocyclops sakitii</taxonomicName>
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sp. n. A habitus in dorsal view B principal caudal setae of left caudal ramus C anal somite and caudal rami, in dorsal view D habitus in dorsal view (Female RBINSc 10.357: A, B; female RBINSc COP 10.354: C; male RBINSc COP 10.356: D).
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</paragraph>
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</caption>
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<caption id="3BA0B59D10F67477AC2138227E17DAB1" pageId="15" pageNumber="16">
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<paragraph id="564E2A3E79FEE398B40D4FFBD9579E46" pageId="15" pageNumber="16">
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Figure 14.
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<taxonomicName id="C0E0004E34E18D22B247F467053FACBF" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops sakitii" order="Cyclopoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sakitii">Allocyclops sakitii</taxonomicName>
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sp. n. A urosome, ventral view B pediger 5 and genital double-somite, in lateral view C leg 1 intercoxal sclerite, frontal view D leg 2 intercoxal sclerite, frontal view E leg 3 intercoxal sclerite, frontal view F leg 4 intercoxal sclerite, frontal view G pediger 5 and urosomite 2 in ventral view (Female RBINSc COP 10.355: A, B; COP 10.358:
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<normalizedToken id="523A844447CB88FFBF2E8B34CFB04DCD" originalValue="C–F">C-F</normalizedToken>
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; male RBINSc COP 10.356: G).
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</paragraph>
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</caption>
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<paragraph id="E46C334476194ABAFB32D363B33D8C3C" pageId="15" pageNumber="16">Caudal rami (Fig. 13C) cylindrical, between 2.5 and 2.6 times longer than wide, with triangular medial depression reaching halfway the margin. Integument of caudal rami smooth. Anterolateral seta short and pinnate, without spinules at insertion. Distolateral element equally or slightly longer than ramus (ratio: 1.0-1.1/1), furnished with widely spaced setules on both sides of the stem. Spinules at insertion of distolateral element present. Terminal medial element longer than ramus (ratio: 1.3-1.5/1) and pinnate. Principal terminal setae with breaking plane and pinnate. Dorsal seta, articulating on small basal part, and as long as medial element.</paragraph>
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<paragraph id="267E51BDE6E79757E72A47BB7DE34C89" pageId="15" pageNumber="16">Antennule 11-segmented, not reaching to posterior edge of cephalothorax. Segment 1 with crescent row of spinules in proximal half (Fig. 4H). Aesthetasc on segment 8 linguiform, as long as accompanying seta, and reaching middle of segment 9. Aesthetasc on segment 10 filiform, as long as accompanying seta, and reaching nearly to distal end of terminal segment. Aesthetasc on segment 11 tubular, as long as segments 9-11 combined, and shorter than accompanying seta (Fig. 4G).</paragraph>
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<paragraph id="80A4B6AB4B1A3A61DB5707F89A350B70" pageId="15" pageNumber="16">
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Antennary general morphology and armature as in
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<taxonomicName id="AF540D80789DC65B8EBE99E6A7C6CCA4" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops chappuisi" order="Cyclopoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chappuisi">Allocyclops chappuisi</taxonomicName>
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. Frontal surface of coxobasis with a cluster of minute spinules (6-7 spinules) in middle of outer margin and a short transverse set of slender ones in the proximal half bear the abexopodal margin (Fig. 5J). Caudal surface nearly completely naked only furnished with a few slender spinules in the proximal half near the abexopodal margin. (Fig. 5I).
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</paragraph>
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<paragraph id="E1D221B346266A691E14FD3E63CF5D87" pageId="15" pageNumber="16">
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Mandible, maxillule, maxilliped and general aspect of maxilla as in
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<taxonomicName id="4C574F78EA000D71F773634C81D7A3A2" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops cavicola" order="Cyclopoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavicola">Allocyclops cavicola</taxonomicName>
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and the preceding species. Claw of maxillary basis blunt, armed with 8-9 large spinules, increasing in width and length medially. Claw shorter than accessory element (Fig. 4L). The latter furnished with groups of long spinules on both sides in proximal half and along one side in distal half, besides the finely serrate ornamentation of the distal part of the stem. Both elements on proximal endopodite segment and terminal element on distal segment furnished partially with long spinules along one side of the stem, finely serrate at the opposite side. Additional setae on distal endopodite segment slender, unequal in length.
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</paragraph>
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<paragraph id="744D707EF6A96C54AAC59F039DF06B64" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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Leg 1-4 protopodite without ornamentation on praecoxa (Fig. 15A, C, E, F). Frontal surface of coxa unadorned except for the usual minute spinules along distal margin. Caudal coxal surface smooth in leg 1, with some slender and short spinules near outer proximal corner in legs 2 and 3 (Fig. 15D). Caudal surface of leg 4 coxa with 2 small clusters of spinules near outer proximal distal corner and a median row of narrow spinules parallel with distal margin (Fig. 15F). Medial coxal seta present in
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<pageBreakToken id="F65041FDD3839409907AE85816C4DE99" pageId="16" pageNumber="17" start="start">all</pageBreakToken>
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legs, pinnate and reaching to middle of proximal endopodite segment in legs 1-3, but hardly beyond basis in leg 4. Intercoxal sclerites as in preceding species. Basis in leg 1 with long medial spine, reaching to middle of distal endopodite segment and with spinules at insertion. Medial margin of legs 1-3 crescent and hairy, naked in leg 4. Medial distal edge of basis in leg 1 rounded, slightly triangularly produced in legs 2-4. Outer seta on basis of legs 1 and 4 as long as exopodite, of legs 2 and 3 half as long.
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</paragraph>
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<caption id="892E336B1677D0593D8E5A2D80F8B42A" pageId="16" pageNumber="17">
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<paragraph id="57A07451CBE321AC7740562C7F08005A" pageId="16" pageNumber="17">
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Figure 15.
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<taxonomicName id="95D5399E54039B720FB90B57A8D86AAA" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops sakitii" order="Cyclopoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sakitii">Allocyclops sakitii</taxonomicName>
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sp. n. A leg 1, frontal view B distal endopodite segment of leg 2, frontal view C leg 3, frontal view D Outer region of coxa and basis of leg 3, caudal view E leg 4, frontal view F leg 4 protopodite and endopodite, caudal view G leg 4 endopodite, caudal view (Female RBINSc COP 10.355:
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<normalizedToken id="D87938F15FA08F309E8867AA0E2523E6" originalValue="A–C">A-C</normalizedToken>
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, F; COP 10.358: D, E; male RBINSc COP 10.356: G).
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</paragraph>
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</caption>
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<paragraph id="C8C1D6052DE6511C9FDDC88C75A57BE5" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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Leg 1-4 rami 2-segmented with general appearance and armament distribution as in preceding species (Fig. 15
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<normalizedToken id="678298BAE74C1FE71B97F07202755DBF" originalValue="A–C">A-C</normalizedToken>
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,
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<normalizedToken id="722C467F5C97D06940D29DE2EF45804F" originalValue="E–F">E-F</normalizedToken>
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). Proximal segment of leg 4 endopodite slightly
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<pageBreakToken id="400F22D13ECD61B3130E719B406C9C42" pageId="17" pageNumber="18" start="start">expanded</pageBreakToken>
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caudally. Distal endopodite segment of leg 4 2.2-2.4 times longer than wide. Inner terminal spine shorter than segment (ratio: 0.9/1), outer spine half as long as inner one. Medial setae on distal endopodite segment of leg 4 reaching distinctly beyond longest terminal spine.
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</paragraph>
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<paragraph id="2283B1A552EF9813B9EF7397A89BFE67" pageId="18" pageNumber="19">
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<pageBreakToken id="F82A9C66A2D57F091A37109E91B784D3" pageId="18" pageNumber="19" start="start">Leg</pageBreakToken>
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5 (Fig. 14A, B) confluent with pediger, represented as a semi-circular expansion bearing 3 elements. Ancestral basal segment represented by long setiform element, ancestral distal segment by two equally long elements: outer one setiform, inner one spiniform. Surface of leg 5 expansion with some short and hairy-like spinules.
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</paragraph>
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<paragraph id="EA9F3C2120A4F75449A133650FD46B02" pageId="19" pageNumber="20">
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<pageBreakToken id="33CC1E41202108E405F5B7C5C379059F" pageId="19" pageNumber="20" start="start">Leg</pageBreakToken>
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6 (Fig. 14B) located laterally on genital double-somite, typically with 3 elements: outer one setiform, middle and medial one dwarfed, hyaline and blunt. Valve unadorned. Genital complex with small ovate anterior receptacle. Caudally expanded receptacle with single medioventral lobe. Copulatory pore leading to U-shaped duct. Lateral arms straight.
|
||
</paragraph>
|
||
<paragraph id="FE7DF1BA39C0B748256935C03F1A3870" pageId="19" pageNumber="20">
|
||
Male: General appearance of body as in the female with prosome 1.5 times longer than urosome. The latter with pediger 6 half as long as wide (Fig. 14G). Posterodorsal fringes of prosomite and first urosomite, and posterior fringes of urosomites as in the female. Body length: 676
|
||
<normalizedToken id="3600444CF5A2811FCE1FDA124EF679F5" originalValue="µm">µm</normalizedToken>
|
||
(allotype and other male paratypes ranging between 650 and 695
|
||
<normalizedToken id="51F96CC0F1C259437D36F13B639A2074" originalValue="μm">μm</normalizedToken>
|
||
; n=4).
|
||
</paragraph>
|
||
<paragraph id="D5C8512555C90E4E1C51D3F4032E76EF" pageId="19" pageNumber="20">Caudal rami 2.5-2.7 times longer than wide, distolateral element 1-1.15 times longer than caudal rami, and terminal medial element 1.30-1.45 times longer than ramus.</paragraph>
|
||
<paragraph id="004D8ECF0C94DBDA9529E68CDDCE6499" pageId="19" pageNumber="20">
|
||
Antennule as described for
|
||
<taxonomicName id="6447E7C2C8D5CBF557EE5F9DBAED69F9" class="Copepoda" family="Cyclopidae" genus="Allocyclops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allocyclops cavicola" order="Cyclopoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cavicola">Allocyclops cavicola</taxonomicName>
|
||
in
|
||
<bibRefCitation id="41B59FD63B5EE64861D7FB9CD160DA4A" author="Fiers, F" journalOrPublisher="Journal of Natural History" pageId="24" pageNumber="25" pagination="175 - 247" title="The generic concept of Allocyclops Kiefer, 1932 (Copepoda: Cyclopoida: Cyclopidae) and alternative view." url="10.1080/00222933.2011.626530" volume="46" year="2012">Fiers (2012)</bibRefCitation>
|
||
. Mouthparts and legs as in the female but proximal endopodite segment of leg 4 without expansion.
|
||
</paragraph>
|
||
<paragraph id="A262C21490BC28326A0CAC00A11E838F" pageId="19" pageNumber="20">Leg 5 (Fig. 14G) with general appearance and integumentary ornamentation as in the female, but with outer setiform element on distal segment vestige longer than inner spiniform element.</paragraph>
|
||
<paragraph id="73E1483D3C9B3257371056B1F50B7994" pageId="19" pageNumber="20">Leg 6 vestige (Fig. 14G) large, with protruded outer distal edge, bearing 2 elements: outer one setiform, half as long as inner one. The latter spiniform. Surface of valve unadorned.</paragraph>
|
||
<paragraph id="43C35A8F829EC3BE9CBA206F873259D7" pageId="19" pageNumber="20">Variability: Apart from slight variation in body length and L/W-ratio of the caudal rami and the setae on the caudal rami, no structural variability was observed.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |