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<document id="7E6C25815889AD89505F343CC6E6FE4E" ID-CLB-Dataset="36889" ID-DOI="10.11646/zootaxa.3774.1.1" ID-GBIF-Dataset="41d311ee-850e-4125-9a40-5a96a2aa9065" ID-ISSN="1175-5326" ID-Zenodo-Dep="285693" ID-ZooBank="B5A2EF46-2BF6-4ED3-A5F4-5F9951400545" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391910470" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="9A6687E64F00303EFF34F902FC8AF8E2" docLanguage="en" docName="zt03774p030.pdf" docOrigin="Zootaxa 3774 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Scydmomicrus) capitiseboraci Jałoszyński, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="21" masterDocId="665FFF9E4F13302AFFA3FFEEFFFEFFCA" masterDocTitle="The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="20" updateTime="1698298320569" updateUser="ExternalLinkService">
<mods:mods id="F31407350D9330D55ABDB33C9ECDC6DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F3148EC8FEFEF7C15545B3E7446ADB87">The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="E807032C899578C32F5AB17CA719BD8D">Author</mods:roleTerm>
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<mods:namePart id="003E329BBE1375456E56C51648F2F66A">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="C19154720EFFC46BCF328CEBC0DFD723">2014</mods:date>
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<mods:number id="72642B632A7CEF7165E0F47BDA08CAC5">3774</mods:number>
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<mods:classification id="2CFD9F1D020C39B3F2CF3538C3C090C5">journal article</mods:classification>
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<mods:identifier id="66EDDE9CBFCDD6E95A4B67A1A7CA1023" type="DOI">10.11646/zootaxa.3774.1.1</mods:identifier>
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<treatment id="9A6687E64F00303EFF34F902FC8AF8E2" ID-DOI="http://doi.org/10.5281/zenodo.5664972" ID-GBIF-Taxon="127667232" ID-Zenodo-Dep="5664972" LSID="urn:lsid:plazi:treatment:9A6687E64F00303EFF34F902FC8AF8E2" httpUri="http://treatment.plazi.org/id/9A6687E64F00303EFF34F902FC8AF8E2" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="5AD5657B4F003039FF34F902FDD4F8ED" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="127036F04F003039FF34F902FCF5F8CC" blockId="19.[151,779,1772,1831]" box="[151,779,1772,1798]" pageId="19" pageNumber="20">
<heading id="4938819C4F003039FF34F902FCF5F8CC" bold="true" box="[151,779,1772,1798]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="20BBEAE24F003039FF34F902FCF5F8CC" bold="true" box="[151,779,1772,1798]" pageId="19" pageNumber="20">
<taxonomicName id="D5CF4D734F003039FF34F902FD3CF8CC" ID-CoL="435TF" box="[151,706,1772,1798]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="capitiseboraci" status="sp. nov." subGenus="Scydmomicrus">
<emphasis id="20BBEAE24F003039FF34F902FEB9F8CC" bold="true" box="[151,327,1772,1798]" italics="true" pageId="19" pageNumber="20">Microscydmus</emphasis>
(
<emphasis id="20BBEAE24F003039FEFBF902FDF7F8CC" bold="true" box="[344,521,1772,1798]" italics="true" pageId="19" pageNumber="20">Scydmomicrus</emphasis>
)
<emphasis id="20BBEAE24F003039FDB9F902FD3CF8CC" bold="true" box="[538,706,1772,1798]" italics="true" pageId="19" pageNumber="20">capitiseboraci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B8857994F003039FD6AF902FCF5F8CC" box="[713,779,1772,1798]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="127036F04F003039FF34F8E0FDD4F8ED" blockId="19.[151,779,1772,1831]" box="[151,554,1806,1831]" pageId="19" pageNumber="20">
(
<figureCitation id="8AF42A754F003039FF3CF8E0FF02F8ED" box="[159,252,1806,1831]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="19" pageNumber="20">Figs. 22</figureCitation>
,
<figureCitation id="8AF42A754F003039FEABF8E1FED8F8ED" box="[264,294,1807,1831]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[157,1418,212,1473]" captionTargetId="figure@10.[151,1436,193,1492]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Body silhouette of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (29); M. (Sc.) nasicornis Franz, female (30); M. (Sc.) capitiseboraci sp. n., male (31); M. (Sc.) queenslandicus sp. n., male (32); M. (Sc.) styxianus sp. n., male (33); M. (Sc.) edithensis sp. n., male (34); M. (Sc.) tooloomensis sp. n., male (35); Penicillidmus masseyensis sp. n., male (36); P. unicolor sp. n., male (37)." httpUri="https://zenodo.org/record/285700/files/figure.png" pageId="19" pageNumber="20">31</figureCitation>
,
<figureCitation id="8AF42A754F003039FE91F8E1FEAFF8ED" box="[306,337,1807,1831]" captionStart="FIGURES 38 – 41" captionStartId="11.[151,264,1708,1731]" captionTargetBox="[166,1418,221,1674]" captionTargetId="figure@11.[151,1436,193,1687]" captionTargetPageId="11" captionText="FIGURES 38 – 41. Head capsule in dorsal (38, 40), dorso-posterior (39) and lateral (41) views. Microscydmus (Scydmomicrus) australiensis Franz, female (38); M. (Sc.) capitiseboraci sp. n., male (39); Penicillidmus sp., female (40 – 41). Abbreviations: cl, clypeus; fr, frons; gen, gena; lb, labrum; md, mandible; mn, mentum; mxp, maxillar palpus; nr, neck region; oc, occipital constriction; occ, occipital constriction; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285701/files/figure.png" pageId="19" pageNumber="20">39</figureCitation>
,
<figureCitation id="8AF42A754F003039FEFEF8E1FE85F8ED" box="[349,379,1807,1831]" captionStart="FIGURES 42 – 45" captionStartId="12.[151,264,1820,1843]" captionTargetBox="[182,1408,310,1798]" captionTargetId="figure@12.[171,1409,301,1808]" captionTargetPageId="12" captionText="FIGURES 42 – 45. Head capsule in ventral (42, 45), and antero-ventral (43 – 44) views. Microscydmus (Scydmomicrus) australiensis Franz, female (42); M. (Sc.) capitiseboraci sp. n., male (43); M. (Sc.) tooloomensis sp. n., male (44); Penicillidmus sp., female (45). Abbreviations: alc, anterior arm of longitudinal carina; bst, basistipes; cd, cardo; fo, foramen occipitale; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lb, labrum; lg, ligula; lp, labial palpus; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV; plc, posterior arm of longitudinal carina; ppf, palpifer; ptp, posterior tentorial pit; smn, submental carina; smn, submentum; tc, transverse carina." httpUri="https://zenodo.org/record/285702/files/figure.png" pageId="19" pageNumber="20">43</figureCitation>
,
<figureCitation id="8AF42A754F003039FE25F8E1FE5AF8ED" box="[390,420,1807,1831]" captionStart="FIGURES 46 – 49" captionStartId="13.[151,264,1431,1454]" captionTargetBox="[154,1421,208,1405]" captionTargetId="figure@13.[151,1436,193,1411]" captionTargetPageId="13" captionText="FIGURES 46 – 49. Prothorax in ventral view. Microscydmus (Scydmomicrus) australiensis Franz, female (46); M. (Sc.) capitiseboraci sp. n., male (47); M. (Sc.) tooloomensis sp. n., male (48); Penicillidmus sp., female (49). Abbreviations: hy, hypomeron; hyr, hypomeral ridge; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pf, profurcal fovea; plp, postero-lateral brush; psp, prosternal intercoxal process." httpUri="https://zenodo.org/record/285703/files/figure.png" pageId="19" pageNumber="20">47</figureCitation>
,
<figureCitation id="8AF42A754F003039FE13F8E0FE30F8ED" box="[432,462,1806,1831]" captionStart="FIGURES 50 – 53" captionStartId="15.[151,264,1823,1846]" captionTargetBox="[221,1330,199,1771]" captionTargetId="figure@15.[210,1365,193,1795]" captionTargetPageId="15" captionText="FIGURES 50 – 53. Pterothorax in ventral view. Microscydmus (Scydmomicrus) australiensis Franz, female (50); M. (Sc.) capitiseboraci sp. n., male (51); M. (Sc.) tooloomensis sp. n., male (52); Penicillidmus sp., female (53). Abbreviations: aest 2, mesanepisternum; ai, asetose impression of mesoventrite; ar, anterior ridge of mesoventrite; cl, coxal line; lmfa; lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pl, posterior lobe of mesocoxal projection; pre, prepectus; sp, spermatheca; v 3, metaventrite." httpUri="https://zenodo.org/record/285704/files/figure.png" pageId="19" pageNumber="20">51</figureCitation>
,
<figureCitation id="8AF42A754F003039FE79F8E0FE07F8ED" box="[474,505,1806,1831]" captionStart="FIGURES 54 – 61" captionStartId="16.[151,264,1561,1584]" captionTargetBox="[152,1427,213,1530]" captionTargetId="figure@16.[151,1436,193,1540]" captionTargetPageId="16" captionText="FIGURES 54 – 61. Aedeagi in ventral views. M. (Sc.) capitiseboraci sp. n. (54); M. (Sc.) queenslandicus sp. n. (55); M. (Sc.) styxianus sp. n. (56); M. (Sc.) edithensis sp. n. (57); M. (Sc.) tooloomensis sp. n. (58 – 59); Penicillidmus masseyensis sp. n. (60); P. unicolor sp. n. (61)." httpUri="https://zenodo.org/record/285705/files/figure.png" pageId="19" pageNumber="20">54</figureCitation>
,
<figureCitation id="8AF42A754F003039FDA7F8E0FDDFF8ED" box="[516,545,1806,1831]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="19" pageNumber="20">62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F003039FF34F8B8FBB4F87D" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="127036F04F003039FF34F8B8FBB4F87D" blockId="19.[151,1437,1878,1975]" pageId="19" pageNumber="20">
<emphasis id="20BBEAE24F003039FF34F8B8FEA1F8A5" bold="true" box="[151,351,1878,1903]" pageId="19" pageNumber="20">Material studied</emphasis>
.
<typeStatus id="CD7488524F003039FECDF8B8FE25F8A5" box="[366,475,1878,1903]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
: ♂: two labels: &quot;
<geoCoordinate id="77FB50374F003039FD39F8B8FD17F8A5" box="[666,745,1878,1903]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="-11.51">11.51S</geoCoordinate>
<geoCoordinate id="77FB50374F003039FD51F8B9FCAEF8A5" box="[754,848,1879,1903]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="142.38">142.38E</geoCoordinate>
QLD /
<quantity id="D5379B154F003039FC0FF8B8FC15F8A5" box="[940,1003,1878,1903]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="km" value="12.0">12km</quantity>
SSE Heathlands /
<date id="667110304F003039FB67F8B9FA91F8A5" box="[1220,1391,1879,1903]" pageId="19" pageNumber="20" value="1992-03-01" valueMax="1992-03-21" valueMin="1992-03-01">1-21 Mar 1992</date>
/ P. Feehney, closed / forest MALAISE#3#4&quot; [white, printed], &quot;
<taxonomicName id="D5CF4D734F003039FC33F895FB99F858" box="[912,1127,1915,1938]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="20BBEAE24F003039FC33F895FB99F858" box="[912,1127,1915,1938]" italics="true" pageId="19" pageNumber="20">MICROSCYDMUS</emphasis>
</taxonomicName>
/ (
<emphasis id="20BBEAE24F003039FB03F895FA89F858" box="[1184,1399,1915,1938]" italics="true" pageId="19" pageNumber="20">SCYDMOMICRUS</emphasis>
) /
<taxonomicName id="D5CF4D734F003039FF34F870FEC9F87C" box="[151,311,1950,1974]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F003039FF34F870FEC9F87C" box="[151,311,1950,1974]" italics="true" pageId="19" pageNumber="20">capitiseboraci</emphasis>
</taxonomicName>
m. / det. P. Jałoszyński, 2013 /
<typeStatus id="CD7488524F003039FD3BF871FCBCF87D" box="[664,834,1951,1975]" pageId="19" pageNumber="20" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="74DEAE354F003039FC57F871FBC1F87D" LSID="urn:lsid:biocol.org:col:32981" box="[1012,1087,1951,1975]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="19" pageNumber="20">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F07303EFF64FF79FD9DFE8A" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="127036F04F07303EFF64FF79FD9DFE8A" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FF79FEBFFF7A" bold="true" box="[199,321,151,176]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
Male: body uniformly light brown; frons subtriangular but with weakly marked antero-median projection; punctures on head fine but distinct, slightly more distinct than those on pronotum; BL &lt;
<quantity id="D5379B154F07303EFA8EFF52FA66FF1E" box="[1325,1432,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="mm" value="0.6">0.60 mm</quantity>
; pronotum only 1.1x broader than head, with ante-basal transverse groove; EI&gt; 1.70; aedeagus with lightly sclerotized and indistinct internal armature with large sub-cylindrical structure located near basal foramen. Females and their diagnostic characters unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F07303EFF64FEA2FE5CFBDA" pageId="20" pageNumber="21" type="description">
<paragraph id="127036F04F07303EFF64FEA2FEF5FE67" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FEA2FEA7FEAF" bold="true" box="[199,345,332,357]" pageId="20" pageNumber="21">Description.</emphasis>
Body of male (
<figureCitation id="8AF42A754F07303EFD8BFEA2FD7CFEAE" box="[552,642,332,357]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="20" pageNumber="21">Fig. 22</figureCitation>
) moderately convex, elongate and slender, with moderately long appendages, BL
<quantity id="D5379B154F07303EFEF7FE81FE34FE4D" box="[340,458,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.12" pageId="20" pageNumber="21" unit="mm" value="0.512">0.512 mm</quantity>
; cuticle glossy; body uniformly light brown with slightly lighter appendages; vestiture yellowish.
</paragraph>
<paragraph id="127036F04F07303EFF64FE59FC01FDAA" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
Head (
<figureCitation id="8AF42A754F07303EFEB7FE59FE96FE1A" box="[276,360,439,464]" captionStart="FIGURES 38 – 41" captionStartId="11.[151,264,1708,1731]" captionTargetBox="[166,1418,221,1674]" captionTargetId="figure@11.[151,1436,193,1687]" captionTargetPageId="11" captionText="FIGURES 38 – 41. Head capsule in dorsal (38, 40), dorso-posterior (39) and lateral (41) views. Microscydmus (Scydmomicrus) australiensis Franz, female (38); M. (Sc.) capitiseboraci sp. n., male (39); Penicillidmus sp., female (40 – 41). Abbreviations: cl, clypeus; fr, frons; gen, gena; lb, labrum; md, mandible; mn, mentum; mxp, maxillar palpus; nr, neck region; oc, occipital constriction; occ, occipital constriction; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285701/files/figure.png" pageId="20" pageNumber="21">Fig. 39</figureCitation>
) subtrapezoidal, broadest at eyes, HL
<quantity id="D5379B154F07303EFC86FE56FC65FE05" box="[805,923,440,464]" metricMagnitude="-5" metricUnit="m" metricValue="8.7" pageId="20" pageNumber="21" unit="mm" value="0.087">0.087 mm</quantity>
, HW
<quantity id="D5379B154F07303EFC41FE56FBA7FE05" box="[994,1113,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.37" pageId="20" pageNumber="21" unit="mm" value="0.137">0.137 mm</quantity>
; tempora much shorter than eyes, strongly convergent postero-mesally; vertex weakly convex; frons subtriangular but with indistinct anteromedian projection. Eyes large and strongly projecting laterally from the head silhouette, coarsely faceted. Punctures on head dorsum sparse and small but distinct; setae short, sparse and suberect. Antennae slender, AnL
<quantity id="D5379B154F07303EFF34FDA9FEF2FD95" box="[151,268,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="0.25">0.250 mm</quantity>
; antennomeres I–II elongate, III–X transverse; XI as long as broad.
</paragraph>
<paragraph id="127036F04F07303EFF64FD83FE5AFD3A" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
Pronotum in dorsal view oval, broadest near anterior third, PL
<quantity id="D5379B154F07303EFC33FD82FBFBFD4E" box="[912,1029,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="20" pageNumber="21" unit="mm" value="0.125">0.125 mm</quantity>
, PW
<quantity id="D5379B154F07303EFBE7FD82FB44FD4E" box="[1092,1210,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="mm" value="0.15">0.150 mm</quantity>
; anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with two pairs of small and shallow pits, internal pair connected by transverse impression. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect.
</paragraph>
<paragraph id="127036F04F07303EFF64FD12FAF8FC97" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
Elytra oval, slightly more convex than pronotum, broadest near middle, EL 0.300 mm, EW
<quantity id="D5379B154F07303EFB58FD12FA8CFCDE" box="[1275,1394,764,789]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="21" unit="mm" value="0.175">0.175 mm</quantity>
, EI 1.714; basal impressions short; elytral apices separately rounded. Punctures on elytral disc as fine as those on pronotum; setae short, sparse and suberect.
<collectingCountry id="6AD876604F07303EFDDCFCAAFD49FC97" box="[639,695,836,861]" name="India" pageId="20" pageNumber="21">Hind</collectingCountry>
wings well developed, about twice as long as elytra.
</paragraph>
<paragraph id="127036F04F07303EFF64FC86FCB0FC4A" blockId="20.[151,1437,151,1365]" box="[199,846,871,896]" pageId="20" pageNumber="21">Legs moderately long and slender, without modifications.</paragraph>
<paragraph id="127036F04F07303EFF64FC62FEB3FC26" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
Aedeagus (
<figureCitation id="8AF42A754F07303EFEE5FC62FE66FC6F" box="[326,408,908,933]" captionStart="FIGURES 54 – 61" captionStartId="16.[151,264,1561,1584]" captionTargetBox="[152,1427,213,1530]" captionTargetId="figure@16.[151,1436,193,1540]" captionTargetPageId="16" captionText="FIGURES 54 – 61. Aedeagi in ventral views. M. (Sc.) capitiseboraci sp. n. (54); M. (Sc.) queenslandicus sp. n. (55); M. (Sc.) styxianus sp. n. (56); M. (Sc.) edithensis sp. n. (57); M. (Sc.) tooloomensis sp. n. (58 – 59); Penicillidmus masseyensis sp. n. (60); P. unicolor sp. n. (61)." httpUri="https://zenodo.org/record/285705/files/figure.png" pageId="20" pageNumber="21">Fig. 54</figureCitation>
) extremely small, AeL only
<quantity id="D5379B154F07303EFD42FC62FCA9FC6E" box="[737,855,908,933]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="mm" value="0.06">0.060 mm</quantity>
; slender, with indistinct, lightly sclerotized internal armature containing in its sub-basal part large sub-cylindrical basally broadening structure; each paramere with long apical seta.
</paragraph>
<paragraph id="127036F04F07303EFF64FC19FE5CFBDA" blockId="20.[151,1437,151,1365]" box="[199,418,1015,1040]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FC19FEE0FBDA" bold="true" box="[199,286,1015,1040]" pageId="20" pageNumber="21">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F07303EFF64FBF2FCCEFBFF" box="[199,816,1052,1077]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="127036F04F07303EFF64FBF2FCCEFBFF" blockId="20.[151,1437,151,1365]" box="[199,816,1052,1077]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FBF2FEA7FBFF" bold="true" box="[199,345,1052,1077]" pageId="20" pageNumber="21">Distribution</emphasis>
(
<figureCitation id="8AF42A754F07303EFECBFBF2FE42FBFF" box="[360,444,1052,1077]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="20" pageNumber="21">Fig. 62</figureCitation>
c). NE
<collectingCountry id="6AD876604F07303EFDAAFBF2FD8EFBFF" box="[521,624,1052,1077]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
(N Queensland).
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F07303EFF64FBD1FEAEFBB6" pageId="20" pageNumber="21" type="etymology">
<paragraph id="127036F04F07303EFF64FBD1FEAEFBB6" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FBD1FEB0FB92" bold="true" box="[199,334,1087,1112]" pageId="20" pageNumber="21">Etymology.</emphasis>
The name
<taxonomicName id="D5CF4D734F07303EFE76FBAEFD8BFB92" box="[469,629,1088,1112]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFE76FBAEFD8BFB92" box="[469,629,1088,1112]" italics="true" pageId="20" pageNumber="21">capitiseboraci</emphasis>
</taxonomicName>
refers to the
<typeStatus id="CD7488524F07303EFCB5FBAEFCB8FB92" box="[790,838,1088,1112]" pageId="20" pageNumber="21">type</typeStatus>
locality, Cape York Peninsula, in Latin
<emphasis id="20BBEAE24F07303EFABDFBAEFEB7FBB6" italics="true" pageId="20" pageNumber="21">Paeninsula Capitis Eboraci</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F07303EFF64FB69FC8AF8E2" pageId="20" pageNumber="21" type="discussion">
<paragraph id="127036F04F07303EFF64FB69FD41FA9F" blockId="20.[151,1437,151,1365]" pageId="20" pageNumber="21">
<emphasis id="20BBEAE24F07303EFF64FB69FEC5FB6A" bold="true" box="[199,315,1159,1184]" pageId="20" pageNumber="21">Remarks.</emphasis>
<taxonomicName id="D5CF4D734F07303EFEE6FB66FD6CFB6A" box="[325,658,1160,1184]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFEE6FB66FD6CFB6A" box="[325,658,1160,1184]" italics="true" pageId="20" pageNumber="21">Microscydmus capitiseboraci</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="D5CF4D734F07303EFCD7FB67FBB4FB6A" box="[884,1098,1160,1184]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="20BBEAE24F07303EFCD7FB67FBB4FB6A" box="[884,1098,1160,1184]" italics="true" pageId="20" pageNumber="21">M. queenslandicus</emphasis>
</taxonomicName>
. Both species have similarly pigmented bodies, comparable body length and proportions of body parts, but clearly differ in punctures on the head dorsum: in
<taxonomicName id="D5CF4D734F07303EFEFBFB3FFDE1FB22" box="[344,543,1232,1256]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFEFBFB3FFDE1FB22" box="[344,543,1232,1256]" italics="true" pageId="20" pageNumber="21">M. capitiseboraci</emphasis>
</taxonomicName>
distinct but sparse and small, while those in
<taxonomicName id="D5CF4D734F07303EFB84FB3FFB05FB22" box="[1063,1275,1232,1256]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="20BBEAE24F07303EFB84FB3FFB05FB22" box="[1063,1275,1232,1256]" italics="true" pageId="20" pageNumber="21">M. queenslandicus</emphasis>
</taxonomicName>
punctures are strikingly large, deep and dense (adjacent to each other). These two species differ also clearly in the elytral index:
<quantity id="D5379B154F07303EFF34FAF6FF0BFAFA" box="[151,245,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="4.35356" pageId="20" pageNumber="21" unit="in" value="1.714">1.714 in</quantity>
<taxonomicName id="D5CF4D734F07303EFF5EFAF7FE3BFAFA" box="[253,453,1304,1328]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFF5EFAF7FE3BFAFA" box="[253,453,1304,1328]" italics="true" pageId="20" pageNumber="21">M. capitiseboraci</emphasis>
</taxonomicName>
vs. only
<quantity id="D5379B154F07303EFD91FAF9FD6EFAFA" box="[562,656,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="20" pageNumber="21" unit="in" value="1.5">1.500 in</quantity>
<taxonomicName id="D5CF4D734F07303EFD3AFAF7FC93FAFA" box="[665,877,1304,1328]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="20BBEAE24F07303EFD3AFAF7FC93FAFA" box="[665,877,1304,1328]" italics="true" pageId="20" pageNumber="21">M. queenslandicus</emphasis>
</taxonomicName>
.
<taxonomicName id="D5CF4D734F07303EFCDEFAF6FB36FAFA" box="[893,1224,1304,1328]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFCDEFAF6FB36FAFA" box="[893,1224,1304,1328]" italics="true" pageId="20" pageNumber="21">Microscydmus capitiseboraci</emphasis>
</taxonomicName>
differs from other Australian congeners in the following characters:
</paragraph>
<paragraph id="127036F04F07303EFF34FA6BFD5BF99A" blockId="20.[151,1436,1412,1832]" pageId="20" pageNumber="21">
- from
<taxonomicName id="D5CF4D734F07303EFEA7FA6BFE46FA56" box="[260,440,1412,1436]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F07303EFEA7FA6BFE46FA56" box="[260,440,1412,1436]" italics="true" pageId="20" pageNumber="21">M. australiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D5CF4D734F07303EFE4DFA6BFD76FA56" box="[494,648,1413,1436]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="20BBEAE24F07303EFE4DFA6BFD76FA56" box="[494,648,1413,1436]" italics="true" pageId="20" pageNumber="21">M. nasicornis</emphasis>
</taxonomicName>
in the shorter and anteriorly less narrowing frons covered with several distinct punctures (frons more distinctly narrowing anteriorly and covered with inconspicuous punctures in
<taxonomicName id="D5CF4D734F07303EFADEFA47FED8FA2E" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F07303EFADEFA47FED8FA2E" italics="true" pageId="20" pageNumber="21">M. australiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D5CF4D734F07303EFEFFFA23FE0BFA2E" box="[348,501,1485,1508]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="20BBEAE24F07303EFEFFFA23FE0BFA2E" box="[348,501,1485,1508]" italics="true" pageId="20" pageNumber="21">M. nasicornis</emphasis>
</taxonomicName>
), in different proportions of the pronotum (PI 0.833 vs.
<quantity id="D5379B154F07303EFBC4FA22FB3DFA2F" box="[1127,1219,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="1.0">1.000 in</quantity>
these two species), in much higher elytral index (EI&gt; 1.700 vs EI &lt;1.600), and a higher ratio EL/PL (2.400 vs. 2.000); - from
<taxonomicName id="D5CF4D734F07303EFEA4F9FBFE69F9E6" box="[263,407,1557,1580]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="styxianus" status="sp. nov.">
<emphasis id="20BBEAE24F07303EFEA4F9FBFE69F9E6" box="[263,407,1557,1580]" italics="true" pageId="20" pageNumber="21">M. styxianus</emphasis>
</taxonomicName>
<emphasis id="20BBEAE24F07303EFE01F9FAFE1AF9E7" bold="true" box="[418,484,1556,1581]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="3B8857994F07303EFE01F9FAFE1AF9E7" box="[418,484,1556,1581]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in the subtriangular frons (in
<taxonomicName id="D5CF4D734F07303EFCE0F9FBFC2DF9E6" box="[835,979,1557,1580]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F07303EFCE0F9FBFC2DF9E6" box="[835,979,1557,1580]" italics="true" pageId="20" pageNumber="21">M. styxianus</emphasis>
</taxonomicName>
the frons is rather subtrapezoidal); and much higher elytral index (EI 1.714 vs. 1.474);
</paragraph>
<paragraph id="127036F04F07303EFF34F9B3FDA4F92A" blockId="20.[151,1436,1412,1832]" pageId="20" pageNumber="21">
- from
<taxonomicName id="D5CF4D734F07303EFEA4F9B3FE61F9BE" box="[263,415,1628,1652]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="edithensis" status="sp. nov.">
<emphasis id="20BBEAE24F07303EFEA4F9B3FE61F9BE" box="[263,415,1628,1652]" italics="true" pageId="20" pageNumber="21">M. edithensis</emphasis>
</taxonomicName>
<emphasis id="20BBEAE24F07303EFE0AF9B2FE15F9BF" bold="true" box="[425,491,1628,1653]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="3B8857994F07303EFE0AF9B2FE15F9BF" box="[425,491,1628,1653]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in the uniformly yellowish-brown body pigmentation (
<taxonomicName id="D5CF4D734F07303EFBC9F9B3FAFCF9BE" box="[1130,1282,1628,1652]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="edithensis">
<emphasis id="20BBEAE24F07303EFBC9F9B3FAFCF9BE" box="[1130,1282,1628,1652]" italics="true" pageId="20" pageNumber="21">M. edithensis</emphasis>
</taxonomicName>
has the head distinctly darker than the pronotum and elytra); elytral sculpture (fine, inconspicuous punctures in
<taxonomicName id="D5CF4D734F07303EFADEF96FFEC9F976" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F07303EFADEF96FFEC9F976" italics="true" pageId="20" pageNumber="21">M. capitiseboraci</emphasis>
</taxonomicName>
vs. dense and fine punctures forming fine transverse wrinkles in
<taxonomicName id="D5CF4D734F07303EFBEBF94BFB1CF976" box="[1096,1250,1700,1724]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="edithensis">
<emphasis id="20BBEAE24F07303EFBEBF94BFB1CF976" box="[1096,1250,1700,1724]" italics="true" pageId="20" pageNumber="21">M. edithensis</emphasis>
</taxonomicName>
); and distinctly smaller body (
<quantity id="D5379B154F07303EFE94F929FE54F92A" box="[311,426,1735,1760]" metricMagnitude="-4" metricUnit="m" metricValue="5.12" pageId="20" pageNumber="21" unit="mm" value="0.512">0.512 mm</quantity>
vs.
<quantity id="D5379B154F07303EFE7BF929FDB0F92A" box="[472,590,1735,1760]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="20" pageNumber="21" unit="mm" value="0.65">0.650 mm</quantity>
);
</paragraph>
<paragraph id="127036F04F07303EFF34F903FC8AF8E2" blockId="20.[151,1436,1412,1832]" pageId="20" pageNumber="21">
- from
<taxonomicName id="D5CF4D734F07303EFEAAF903FE39F8CE" box="[265,455,1772,1796]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tooloomensis">
<emphasis id="20BBEAE24F07303EFEAAF903FE39F8CE" box="[265,455,1772,1796]" italics="true" pageId="20" pageNumber="21">M. tooloomensis</emphasis>
</taxonomicName>
in the presence of transverse ante-basal impression on the pronotum (absent in
<taxonomicName id="D5CF4D734F07303EFADEF903FED5F8E2" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tooloomensis">
<emphasis id="20BBEAE24F07303EFADEF903FED5F8E2" italics="true" pageId="20" pageNumber="21">M. tooloomensis</emphasis>
</taxonomicName>
); and a smaller body (BL
<quantity id="D5379B154F07303EFDF1F8E1FD3AF8E2" box="[594,708,1807,1832]" metricMagnitude="-4" metricUnit="m" metricValue="5.12" pageId="20" pageNumber="21" unit="mm" value="0.512">0.512 mm</quantity>
vs. 0.600 mm).
</paragraph>
</subSubSection>
</treatment>
</document>
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