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<document id="43B41F67F368B13CE7728BB4BF77DBF1" ID-DOI="10.11646/zootaxa.5040.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531583" ID-ZooBank="FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749724544" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo &amp; García Aldrete, Alfonso N." docDate="2021" docId="9946F564FFBBFFE0FF686FD9FF6BFCE1" docLanguage="en" docName="zootaxa.5040.4.1.pdf" docOrigin="Zootaxa 5040 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja risaraldensis Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="468" masterDocId="657F8D1CFFB5FFF1FFFF6A5AFF8BFFBD" masterDocTitle="New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico" masterLastPageNumber="481" masterPageNumber="451" pageNumber="465" updateTime="1699415585501" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9DF7DA31E7BC10A3A288F1EFFE7D244F">New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico</mods:title>
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<mods:namePart id="47110472B1FA3C8AB961A9EA2D477E87">Carrejo, Nancy</mods:namePart>
<mods:affiliation id="9C1D9B9F3DD6519C0792C1BEAE8ADFD8">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
<mods:nameIdentifier id="0D0BEA712A399711DAB73D17C8912E91" type="email">nancy.carrejo@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="DAD3135D12D00D83CF215D5FB9FF32B4">Obando, Ranulfo González</mods:namePart>
<mods:affiliation id="750E239858385A457053368BA5803ED7">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
<mods:nameIdentifier id="AD5351963D7ADE7052BDADE6F6433304" type="email">ranulfo.gonzález@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="94127317206219A7AD53EB667332976A">Casasola-González, José Arturo</mods:namePart>
<mods:nameIdentifier id="FC1FE36F6EF2CAA6E86142C2D4B1A4A7" type="ORCID">0000-0002-4765-6707</mods:nameIdentifier>
<mods:affiliation id="B7F1BF8C30E1DEAFEB073E0B5D998880">casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707</mods:affiliation>
<mods:nameIdentifier id="7F4717F79654C7BB71EA8B59D5502810" type="email">casasola@unsij.edu.mx</mods:nameIdentifier>
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<mods:namePart id="F0821DEF3F6376F3BE8A86594D886B12">García Aldrete, Alfonso N.</mods:namePart>
<mods:affiliation id="8D1AC19CC2EAECA1F37CB91A86BFEA8B">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. &amp; anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<mods:title id="D170B572521820B6D7DEEA60A747767A">Zootaxa</mods:title>
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<mods:date id="C8AF4CCAD0B4CA31580360AAA9D4344D">2021</mods:date>
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<mods:number id="1E67F68BAF526746CBC4A00BC1184D7B">2021-09-23</mods:number>
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<treatment id="9946F564FFBBFFE0FF686FD9FF6BFCE1" ID-DOI="http://doi.org/10.5281/zenodo.5531597" ID-Zenodo-Dep="5531597" LSID="urn:lsid:plazi:treatment:9946F564FFBBFFE0FF686FD9FF6BFCE1" httpUri="http://treatment.plazi.org/id/9946F564FFBBFFE0FF686FD9FF6BFCE1" lastPageId="17" lastPageNumber="468" pageId="14" pageNumber="465">
<subSubSection id="59F517F9FFBBFFFFFF686FD9FD8EFA23" box="[151,517,1411,1438]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="11504472FFBBFFFFFF686FD9FD8EFA23" blockId="14.[151,517,1411,1474]" box="[151,517,1411,1438]" pageId="14" pageNumber="465">
<heading id="4A18F31EFFBBFFFFFF686FD9FD8EFA23" bold="true" box="[151,517,1411,1438]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<emphasis id="239B9860FFBBFFFFFF686FD9FD8EFA23" bold="true" box="[151,517,1411,1438]" pageId="14" pageNumber="465">
<taxonomicName id="D6EF3FF1FFBBFFFFFF686FD9FEFFFA23" ID-CoL="9J3VL" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[151,372,1411,1438]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="risaraldensis" status="sp. nov.">
<emphasis id="239B9860FFBBFFFFFF686FD9FEFFFA23" bold="true" box="[151,372,1411,1438]" italics="true" pageId="14" pageNumber="465">Goja risaraldensis</emphasis>
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<taxonomicNameLabel id="38A8251BFFBBFFFFFE846FDEFE35FA23" box="[379,446,1412,1438]" pageId="14" pageNumber="465" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
</heading>
</paragraph>
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<subSubSection id="59F517F9FFBBFFFFFF686FF2FEACFA7F" box="[151,295,1448,1474]" pageId="14" pageNumber="465" type="description">
<paragraph id="11504472FFBBFFFFFF686FF2FEACFA7F" blockId="14.[151,517,1411,1474]" box="[151,295,1448,1474]" pageId="14" pageNumber="465">
(
<figureCitation id="89D458F7FFBBFFFFFF606FF2FE94FA7F" box="[159,287,1448,1474]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="14" pageNumber="465">Figs 1924</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFBBFFE0FF686FB5FE24FD39" lastPageId="17" lastPageNumber="468" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="11504472FFBBFFFFFF686FB5FCACF9CB" blockId="14.[151,1437,1519,2014]" pageId="14" pageNumber="465">
<emphasis id="239B9860FFBBFFFFFF686FB5FE99F9B4" bold="true" box="[151,274,1519,1545]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
Similar to
<emphasis id="239B9860FFBBFFFFFE706FAAFE72F9B7" box="[399,505,1520,1546]" italics="true" pageId="14" pageNumber="465">
P.
<taxonomicName id="D6EF3FF1FFBBFFFFFE546FAAFE72F9B7" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[427,505,1520,1546]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="andina" status="sp. nov.">andina</taxonomicName>
</emphasis>
<emphasis id="239B9860FFBBFFFFFDFF6FAAFDB4F9B7" bold="true" box="[512,575,1520,1546]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="38A8251BFFBBFFFFFDFF6FAAFDB4F9B7" box="[512,575,1520,1546]" pageId="14" pageNumber="465" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, from which it differs in the wing venation, in having larger external parameres, in the head pattern, and in the endophallic sclerites (Compare
<figureCitation id="89D458F7FFBBFFFFFC6F6C4EFC7AF993" box="[912,1009,1556,1582]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="14" pageNumber="465">Figs 12</figureCitation>
with 1920, 3 with 21, and 6 with
<quantity id="D617E997FFBBFFFFFA7F6C4EFF26F9EF" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="14" pageNumber="465" unit="in" value="24.0">24 in</quantity>
this paper). Apex of the aedeagal arch robust, with the posterior border concave and a pair of the left endophallic sclerites more widened distally than in
<emphasis id="239B9860FFBBFFFFFDB16C06FD3CF9CB" box="[590,695,1628,1654]" italics="true" pageId="14" pageNumber="465">
P.
<taxonomicName id="D6EF3FF1FFBBFFFFFD966C06FD3CF9CB" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[617,695,1628,1654]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="andina">andina</taxonomicName>
</emphasis>
(
<figureCitation id="89D458F7FFBBFFFFFD396C06FC90F9CB" box="[710,795,1628,1654]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="14" pageNumber="465">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="11504472FFBBFFFFFF386C25FE49F863" blockId="14.[151,1437,1519,2014]" pageId="14" pageNumber="465">
<emphasis id="239B9860FFBBFFFFFF386C25FE80F924" bold="true" box="[199,267,1663,1689]" pageId="14" pageNumber="465">Color</emphasis>
(in 80% ethanol). Body brown with light spots. Head pattern (
<figureCitation id="89D458F7FFBBFFFFFC416CDAFB9BF927" box="[958,1040,1664,1690]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="14" pageNumber="465">Fig. 21</figureCitation>
); forehead brown, with broad ochre band between the compound eyes, enclosing the ocellar group. Compound eyes dark brown; ocelli hyaline, with ochre centripetal crescents. Anteclypeus brown, with ochre striae converging mesally as illustrated; labrum dark brown; genae cream to light brown, with ochre anterior areas; pre- and postgenitalia cream. Base of maxillae cream, with brown spots, maxillary palps brown, Mx4 darker.Antennae light brown. Thorax light brown, pronotum cream, tergal lobes of meso- and metathorax brown; pleura cream, with ochre spots. Legs light brown, coxae with basal ochre spots. Wings hyaline, veins light brown. Forewings (
<figureCitation id="89D458F7FFBBFFFFFCC86D02FC01F8CF" box="[823,906,1880,1906]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="14" pageNumber="465">Fig. 19</figureCitation>
) with abundant brown setae. Abdomen cream, with ochre brown subcuticular spots. Clunium, epiproct and paraprocts light brown, with brown spots. Hypandrium light brown. Phallosome proximally creamy, distally brown; aedeagal arch and endophallic sclerites dark brown, external parameres brown.
</paragraph>
<caption id="459014FAFFBAFFFEFF686D3FFD69F81C" ID-DOI="http://doi.org/10.5281/zenodo.5531589" ID-Zenodo-Dep="5531589" httpUri="https://zenodo.org/record/5531589/files/figure.png" pageId="15" pageNumber="466" startId="15.[151,264,1893,1917]" targetBox="[163,1423,195,1856]" targetPageId="15">
<paragraph id="11504472FFBAFFFEFF686D3FFD69F81C" blockId="15.[151,1437,1893,1953]" pageId="15" pageNumber="466">
<emphasis id="239B9860FFBAFFFEFF686D3FFEDEF8C0" bold="true" box="[151,341,1893,1917]" pageId="15" pageNumber="466">FIGURES 1318.</emphasis>
<taxonomicName id="D6EF3FF1FFBAFFFEFEA36D3FFD8CF8C0" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[348,519,1893,1917]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="meremberg" status="sp. nov.">
<emphasis id="239B9860FFBAFFFEFEA36D3FFD8CF8C0" box="[348,519,1893,1917]" italics="true" pageId="15" pageNumber="466">Goja meremberg</emphasis>
</taxonomicName>
<emphasis id="239B9860FFBAFFFEFDF06D3FFDC2F8C0" bold="true" box="[527,585,1893,1917]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="38A8251BFFBAFFFEFDF06D3FFDC2F8C0" box="[527,585,1893,1917]" pageId="15" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male.
<emphasis id="239B9860FFBAFFFEFD6D6D3FFD20F8C0" bold="true" box="[658,683,1893,1917]" pageId="15" pageNumber="466">13</emphasis>
. Forewing.
<emphasis id="239B9860FFBAFFFEFCD86D3FFCCBF8C0" bold="true" box="[807,832,1893,1917]" pageId="15" pageNumber="466">14</emphasis>
. Hindwing.
<emphasis id="239B9860FFBAFFFEFC3E6D3FFC51F8C0" bold="true" box="[961,986,1893,1917]" pageId="15" pageNumber="466">15</emphasis>
. Front view of head.
<emphasis id="239B9860FFBAFFFEFB476D3FFB5AF8C0" bold="true" box="[1208,1233,1893,1917]" pageId="15" pageNumber="466">16</emphasis>
. Left paraproct and epiproct.
<emphasis id="239B9860FFBAFFFEFF086DD3FE9BF81C" bold="true" box="[247,272,1929,1953]" pageId="15" pageNumber="466">17</emphasis>
. Hypandrium.
<emphasis id="239B9860FFBAFFFEFE556DD3FE48F81C" bold="true" box="[426,451,1929,1953]" pageId="15" pageNumber="466">18</emphasis>
. Phallosome. Scales in mm.
</paragraph>
</caption>
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<paragraph id="11504472FFA5FFE1FF686D09FD69F832" blockId="16.[151,1437,1875,1936]" pageId="16" pageNumber="467">
<emphasis id="239B9860FFA5FFE1FF686D09FEDFF8D1" bold="true" box="[151,340,1875,1900]" pageId="16" pageNumber="467">FIGURES 1924.</emphasis>
<taxonomicName id="D6EF3FF1FFA5FFE1FEA56D0EFD9CF8D1" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[346,535,1876,1900]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="risaraldensis" status="sp. nov.">
<emphasis id="239B9860FFA5FFE1FEA56D0EFD9CF8D1" box="[346,535,1876,1900]" italics="true" pageId="16" pageNumber="467">Goja risaraldensis</emphasis>
</taxonomicName>
<emphasis id="239B9860FFA5FFE1FDE16D0EFDDCF8D1" bold="true" box="[542,599,1876,1900]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="38A8251BFFA5FFE1FDE16D0EFDDCF8D1" box="[542,599,1876,1900]" pageId="16" pageNumber="467" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male.
<emphasis id="239B9860FFA5FFE1FD626D0EFD3DF8D1" bold="true" box="[669,694,1876,1900]" pageId="16" pageNumber="467">19</emphasis>
. Forewing.
<emphasis id="239B9860FFA5FFE1FCCE6D0EFCC1F8D1" bold="true" box="[817,842,1876,1900]" pageId="16" pageNumber="467">20</emphasis>
. Hindwing.
<emphasis id="239B9860FFA5FFE1FC366D0EFC69F8D1" bold="true" box="[969,994,1876,1900]" pageId="16" pageNumber="467">21</emphasis>
. Front view of head.
<emphasis id="239B9860FFA5FFE1FB446D0EFB5FF8D1" bold="true" box="[1211,1236,1876,1900]" pageId="16" pageNumber="467">22</emphasis>
. Left paraproct and epiproct.
<emphasis id="239B9860FFA5FFE1FF086D22FE9BF82D" bold="true" box="[247,272,1912,1936]" pageId="16" pageNumber="467">23</emphasis>
. Hypandrium.
<emphasis id="239B9860FFA5FFE1FE556D22FE48F82D" bold="true" box="[426,451,1912,1936]" pageId="16" pageNumber="467">24</emphasis>
. Phallosome. Scales in mm.
</paragraph>
</caption>
<paragraph id="11504472FFA4FFE0FF386ACCFE24FD39" blockId="17.[151,1437,150,860]" pageId="17" pageNumber="468">
<emphasis id="239B9860FFA4FFE0FF386ACCFEEAFF0D" bold="true" box="[199,353,150,176]" pageId="17" pageNumber="468">Morphology.</emphasis>
Head: H/MxW: 1.48; compound eyes small, H/d: 3.75; IO/MxW: 0.61. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.15. Labrum elongate, lower margin with a pair of projections with rounded apex, with five distal sensillae, one central placoid, flanked by a pair of trichoid-placoid. Forewings (
<figureCitation id="89D458F7FFA4FFE0FA906A84FF3DFEA1" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 19</figureCitation>
): FW/W: 2.81; pterostigma elongate, wider in the middle: lp/wp: 6.04; Rs with seven branches; areola postica elongate: al/ah: 4.32; M with six branches, M5 and M6 forked. Hindwings (
<figureCitation id="89D458F7FFA4FFE0FC0E6B7CFBCCFEFD" box="[1009,1095,294,320]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 20</figureCitation>
): HW/w: 3.12. Rs and M with four branches. Legs: Fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium (
<figureCitation id="89D458F7FFA4FFE0FA906B10FF3FFE35" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 23</figureCitation>
) broad, setose, posterior border slightly concave in the middle. Phallosome (
<figureCitation id="89D458F7FFA4FFE0FBE36B34FBF9FE35" box="[1052,1138,366,392]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 24</figureCitation>
): deeply obtusely concave anteriorly, side struts slender, straight; aedeagal arch robust, apically with two short projections, directed outward; external parameres well developed, inner border almost straight, outer border rounded, right one distally acuminate, left one blunt distally. Three pairs of asymmetric endophallic sclerites, as illustrated (
<figureCitation id="89D458F7FFA4FFE0FBA76B80FB26FE49" box="[1112,1197,474,500]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 24</figureCitation>
). Clunium projecting posteriorly over the area of the epiproct. Paraprocts large, broadly elliptic (
<figureCitation id="89D458F7FFA4FFE0FC206BA4FBB8FDA5" box="[991,1075,510,536]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 22</figureCitation>
), setae on posterior border as illustrated with setae and macrosetae; large elliptic sensory fields with 32-33 trichobothria in basal rosettes. Epiproct convex anteriorly, sides converging to almost straight posterior border; four macrosetae mesally, other setae as illustrated (
<figureCitation id="89D458F7FFA4FFE0FEF86830FED3FD39" box="[263,344,618,644]" captionStart="FIGURES 1924" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 1924. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="17" pageNumber="468">Fig. 22</figureCitation>
) broad.
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFA4FFE0FF3868D4FB2BFCA9" pageId="17" pageNumber="468" type="description">
<paragraph id="11504472FFA4FFE0FF3868D4FC98FD70" blockId="17.[151,1437,150,860]" pageId="17" pageNumber="468">
<emphasis id="239B9860FFA4FFE0FF3868D4FEF5FD15" bold="true" box="[199,382,654,680]" pageId="17" pageNumber="468">Measurements.</emphasis>
FW: 4500, HW: 3275, F: 1080, T: 1980, t1: 995, t2: 280, ctt1: 39, f1: 1020, f2: 960, f3: 780, Mx4: 390, IO: 295, D: 252, d: 315, IO/d: 1.55, PO: 0.85.
</paragraph>
<paragraph id="11504472FFA4FFE0FF38688CFB2BFCA9" blockId="17.[151,1437,150,860]" pageId="17" pageNumber="468">
<emphasis id="239B9860FFA4FFE0FF38688CFE10FD4D" bold="true" box="[199,411,726,752]" pageId="17" pageNumber="468">Specimen studied</emphasis>
.
<materialsCitation id="A1874E2FFFA4FFE0FE53688CFB2BFCA9" collectingDate="2015-07-18" collectionCode="R" collectorName="Gonzalez, N &amp; O. Saenz." country="Colombia" elevation="2099" latitude="5.1256666" location="Planes de San Rafael" longLatPrecision="1" longitude="-76.009" municipality="Santuario" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Risaralda" typeStatus="holotype">
<typeStatus id="CE54FAD0FFA4FFE0FE53688CFD9FFD4D" box="[428,532,726,752]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="69F804E2FFA4FFE0FD9A688CFD75FD4D" box="[613,766,726,752]" name="Colombia" pageId="17" pageNumber="468">COLOMBIA</collectingCountry>
.
<collectingRegion id="D32B8A90FFA4FFE0FCF4688CFCF3FD4D" box="[779,888,726,752]" country="Colombia" name="Risaralda" pageId="17" pageNumber="468">Risaralda</collectingRegion>
,
<collectingMunicipality id="F134DE08FFA4FFE0FC79688CFC7FFD4D" box="[902,1012,726,752]" pageId="17" pageNumber="468">Santuario</collectingMunicipality>
,
<location id="143012A9FFA4FFE0FBFD688CFB72FD4D" LSID="urn:lsid:plazi:treatment:9946F564FFBBFFE0FF686FD9FF6BFCE1:143012A9FFA4FFE0FBFD688CFB72FD4D" box="[1026,1273,726,752]" country="Colombia" latitude="5.1256666" longLatPrecision="1" longitude="-76.009" municipality="Santuario" name="Planes de San Rafael" pageId="17" pageNumber="468" stateProvince="Risaralda">Planes de San Rafael</location>
,
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:
<geoCoordinate id="74DB22B5FFA4FFE0FF6868A0FEB1FCA9" box="[151,314,762,788]" degrees="76" direction="west" minutes="00" orientation="longitude" pageId="17" pageNumber="468" precision="1" seconds="32.4" value="-76.009">76º0032.4”W</geoCoordinate>
,
<quantity id="D617E997FFA4FFE0FEBB68A0FE17FCA8" box="[324,412,762,789]" metricMagnitude="3" metricUnit="m" metricValue="2.099" pageId="17" pageNumber="468" unit="m" value="2099.0">
<elevation id="9AC2A341FFA4FFE0FEBB68A0FE17FCA8" box="[324,412,762,789]" metricMagnitude="3" metricUnit="m" metricValue="2.099" pageId="17" pageNumber="468" unit="m" value="2099.0">2099 m</elevation>
</quantity>
.
<date id="655162B2FFA4FFE0FE5868A0FDB8FCA8" box="[423,563,762,789]" pageId="17" pageNumber="468" value="2015-07-18">
<collectingDate id="75159B5AFFA4FFE0FE5868A0FDB8FCA8" box="[423,563,762,789]" pageId="17" pageNumber="468" value="2015-07-18">18.VII.2015</collectingDate>
</date>
, on tree trunk.
<collectionCode id="77FEDCB7FFA4FFE0FD2468A1FD7AFCA8" box="[731,753,763,789]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="468">R</collectionCode>
.
<collectorName id="BC1A21A4FFA4FFE0FD0368A0FC00FCA8" box="[764,907,762,789]" pageId="17" pageNumber="468">González, N</collectorName>
. Calderón and
<collectorName id="BC1A21A4FFA4FFE0FBCC68A0FB2BFCA9" box="[1075,1184,762,788]" pageId="17" pageNumber="468">O. Saenz.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFA4FFE0FF386944FF6BFCE1" pageId="17" pageNumber="468" type="etymology">
<paragraph id="11504472FFA4FFE0FF386944FF6BFCE1" blockId="17.[151,1437,150,860]" pageId="17" pageNumber="468">
<emphasis id="239B9860FFA4FFE0FF386944FEC5FC85" bold="true" box="[199,334,798,824]" pageId="17" pageNumber="468">Etymology.</emphasis>
The specific epithet refers to the department of
<collectingRegion id="D32B8A90FFA4FFE0FC746944FC73FC85" box="[907,1016,798,824]" country="Colombia" name="Risaralda" pageId="17" pageNumber="468">Risaralda</collectingRegion>
,
<collectingCountry id="69F804E2FFA4FFE0FBF86944FBF3FC85" box="[1031,1144,798,824]" name="Colombia" pageId="17" pageNumber="468">Colombia</collectingCountry>
, where the
<typeStatus id="CE54FAD0FFA4FFE0FAFB6944FAEDFC85" box="[1284,1382,798,824]" pageId="17" pageNumber="468" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>