342 lines
36 KiB
XML
342 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.230.105878" ID-Pensoft-Pub="1314-2003-230-157" ID-Pensoft-UUID="2E95E2BCAF7955F8B2ABBD7F472DD9B0" ModsDocID="1314-2003-230-157" checkinTime="1691771898489" checkinUser="pensoft" docAuthor="Vigosa-Mercado, J. Luis, Delgado-Salinas, Alfonso, Alvarado Cardenas, Leonardo O. & Eguiarte, Luis E." docDate="2023" docId="9905255EFE665EF18248CB79C264A622" docLanguage="en" docName="PhytoKeys 230: 157-256" docOrigin="PhytoKeys 230" docPubDate="2023-08-11" docSource="http://dx.doi.org/10.3897/phytokeys.230.105878" docTitle="Agrostis pallens Trin., Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4 (3 – 4): 328. 1841." docType="treatment" docVersion="1" id="2E95E2BCAF7955F8B2ABBD7F472DD9B0" lastPageNumber="157" masterDocId="2E95E2BCAF7955F8B2ABBD7F472DD9B0" masterDocTitle="Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico" masterLastPageNumber="256" masterPageNumber="157" pageNumber="157" updateTime="1691771898489" updateUser="pensoft">
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<mods:titleInfo id="47AABB16ABA57B34B389B85B92E2899E">
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<mods:title id="142D4E19E4F3CE318298AE3103FC4506">Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico</mods:title>
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<mods:name id="7B823F3F29F878D6242C4A3573E83B5C" type="personal">
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<mods:namePart id="81B42AE201993F76F9E6367A199F61B7">Vigosa-Mercado, J. Luis</mods:namePart>
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<mods:nameIdentifier id="B32908D709620A3BCEDAE63A2B336BD3" type="ORCID">https://orcid.org/0000-0002-9999-0272</mods:nameIdentifier>
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<mods:affiliation id="62D141AC8E248698235DBDBCAC45F284">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:nameIdentifier id="63AFB81EBECB36E826CF481F9E808BD9" type="email">luis_vigosa@ciencias.unam.mx</mods:nameIdentifier>
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<mods:roleTerm id="6D9FC47C61955CF5135917B8AA5C7071">Author</mods:roleTerm>
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<mods:namePart id="F1BFC04B019F8D088486CBAF07C875CF">Delgado-Salinas, Alfonso</mods:namePart>
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<mods:nameIdentifier id="8B0C1955B092D2C46CADF3806E87FADE" type="ORCID">https://orcid.org/0000-0002-9322-9968</mods:nameIdentifier>
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<mods:affiliation id="A4724DEDBF6803349B762367DFDACF8A">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="74485807BAEE29FF48BE538C2CFB228F">Author</mods:roleTerm>
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<mods:namePart id="79D82961F7FB8A798B71CCE1127D38D9">Alvarado Cardenas, Leonardo O.</mods:namePart>
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<mods:nameIdentifier id="999728B6B704CDAA23D4AFC6B74CCF47" type="ORCID">https://orcid.org/0000-0002-4938-8339</mods:nameIdentifier>
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<mods:affiliation id="7AE41EE40ACA2BF6D4D5D7F3D80CE345">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:namePart id="8004797929A278DFB0A4C8A6B9C41D44">Eguiarte, Luis E.</mods:namePart>
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<mods:nameIdentifier id="A71DC992CFED753138F5AD5CF75CC679" type="ORCID">https://orcid.org/0000-0002-5906-9737</mods:nameIdentifier>
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<mods:affiliation id="0393F41A16EDF2D171333823CB1DEB63">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:title id="EBDD2EFE35083C149C4FB7E7B7CA9CC8">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="F7BED0AA1BF041FC028D48A0F08F9623">
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<mods:date id="FA62726C91E57F171E22EA3B3B48990B">2023</mods:date>
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<mods:number id="DC6703F87D12BE82B33D1D5A8DB5F0CC">2023-08-11</mods:number>
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<mods:number id="92E76DDDDAB1E913C7FAF82A0FF416A2">230</mods:number>
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<mods:start id="25654D41A8FD7E90427221FDF68A023D">157</mods:start>
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<mods:end id="2AA1104D549490D0D3CDFFAC7047E851">256</mods:end>
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<mods:url id="65756737A9E5F1596BB497DDCEF8DA70">http://dx.doi.org/10.3897/phytokeys.230.105878</mods:url>
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<mods:classification id="A85F1FEDFE0DCA1C84EA343AB9ABE878">journal article</mods:classification>
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<mods:identifier id="CB7DBD78FDDD1B9AAA96E01008D037DA" type="DOI">http://dx.doi.org/10.3897/phytokeys.230.105878</mods:identifier>
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<treatment id="9905255EFE665EF18248CB79C264A622" LSID="urn:lsid:plazi:treatment:9905255EFE665EF18248CB79C264A622" httpUri="http://treatment.plazi.org/id/9905255EFE665EF18248CB79C264A622" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection id="7209C9EE772ED4E25D7FE1759F62DB0F" pageId="0" pageNumber="157" type="nomenclature">
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<paragraph id="2311A14F650134ECBFE84FEB9BE19236" pageId="0" pageNumber="157">
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12.
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<taxonomicName id="09A416E164156E58E2C7CA3FAEDEA4B2" ID-CoL="B9YJ" ID-ENA="345919" LSID="9905255E-FE66-5EF1-8248-CB79C264A622" authority="Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4 (3 - 4): 328. 1841." authorityName="Trin., Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4 (3 – 4): 328. 1841." class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pallens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="pallens">
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Agrostis pallens Trin.,
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<normalizedToken id="9D8A5550285B81E308C4A0EEF6D78DFE" originalValue="Mém">Mem</normalizedToken>
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. Acad. Imp. Sci.
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<normalizedToken id="A882A099696F4C4A5FFAE24613BFE2D9" originalValue="Saint-Pétersbourg">Saint-Petersbourg</normalizedToken>
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,
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<normalizedToken id="91B3F81442A7F461C3443246F3B489F0" originalValue="Sér">Ser</normalizedToken>
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||
. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(3-4): 328. 1841.
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D68729ABB9703C512C8A47B349A1D54C" pageId="0" pageNumber="157" type="description">
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||
<paragraph id="3A1A11023AE92FA02FD4FC0CC8B85F8E" pageId="0" pageNumber="157">
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<figureCitation id="8913989C657F7BD823E94D33B345A212" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Florets of Agrostis species observed with SEM A A. bourgaei B A. calderoniae C A. capillaris D A. elliottiana E A. exarata F A. ghiesbreghtii G A. gigantea H A. hyemalis I A. laxissima J A. microphylla K A. pallens L A. perennans M A. scabra N A. stolonifera O A. subpatens P A. subrepens Q A. tolucensis R A. turrialbae S A. variabilis. Scale bars: 0.3 mm." figureDoi="10.3897/phytokeys.230.105878.figure4" httpUri="https://binary.pensoft.net/fig/891239" pageId="0" pageNumber="157">Figs 4K</figureCitation>
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<figureCitation id="FDB5B4C0877A72E46500AC7CD0E48101" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Calluses of Agrostis species observed with SEM A A. bourgaei B A. calderoniae C A. capillaris D A. elliottiana E A. exarata F A. ghiesbreghtii G A. gigantea H A. hyemalis I A. laxissima J A. microphylla K A. pallens L A. perennans. Scale bars: 50 μm." figureDoi="10.3897/phytokeys.230.105878.figure5" httpUri="https://binary.pensoft.net/fig/891240" pageId="0" pageNumber="157">, 5K</figureCitation>
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<figureCitation id="D5F5C8E9E92F0471E33F5941E530B09A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Agrostis pallens A whole plant B detail of the rhizome C ligular area D spikelet E floret, abaxial view F floret, adaxial view. Based on Hendrickson 1099 (SD). Scale bars: 3 cm (A); 2 cm (B); 2 mm (C); 0.5 mm (D-F)." figureDoi="10.3897/phytokeys.230.105878.figure24" httpUri="https://binary.pensoft.net/fig/891259" pageId="0" pageNumber="157">, 24</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="26A1FDFBC403F667A51CE29EBBB801EE" pageId="0" pageNumber="157" type="reference_group">
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<paragraph id="15D1E0D4179AAAA4E152CB085761FF77" pageId="0" pageNumber="157">
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<taxonomicName id="DA64C074E208D6DFD618E71F02E97CCA" authorityName="Vasey" authorityYear="1886" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis diegoensis" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="diegoensis">= Agrostis diegoensis</taxonomicName>
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Vasey, Bull. Torrey Bot. Club 13: 55. 1886.
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<taxonomicName id="3AF18A2B7D17B1E1190245B9B3D32CE8" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis multiculmis" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="multiculmis">Agrostis multiculmis</taxonomicName>
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Vasey ex Beal, Grass. N. Amer. 328. 1896, nom. inval., pro syn. Type: USA. California: San Diego, 1884, C. Orcutt s.n. (lectotype, designated by
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<bibRefCitation id="0BE74835543453AF8B8220C0D0ECBADC" author="Vigosa-Mercado, JL" journalOrPublisher="138) Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Mexico City" pageId="0" pageNumber="157" publicationUrl="https://www.phytoneuron.net/wp-content/uploads/2022/10/44PhytoN-PooideaeLectotypes.pdf" refId="B75" refString="Vigosa-Mercado, JL, 2022a. Typification of names in Mexican species of Pooideae (Poaceae). Phytoneuron 2022-44: 1-4. https://www.phytoneuron.net/wp-content/uploads/2022/10/44PhytoN-PooideaeLectotypes.pdf" title="Typification of names in Mexican species of Pooideae (Poaceae). Phytoneuron 2022 - 44: 1 - 4." url="https://www.phytoneuron.net/wp-content/uploads/2022/10/44PhytoN-PooideaeLectotypes.pdf" year="2022 a">Vigosa-Mercado (2022a</bibRefCitation>
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: 2): US (US00131740 [image!]); isolectotypes: GH (GH00022962 [image!]), K (K000838209 [image!]), NY (NY00327625 [image!]), OSC (OSC0001812 [image!]), W (W19160036561 [image!])).*
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</paragraph>
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</subSubSection>
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||
<subSubSection id="EB935D5D690A193153B614E2E57D07B9" pageId="0" pageNumber="157" type="materials_examined">
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<paragraph id="C1CD90D0AAC07183ACD52DBE8A901646" pageId="0" pageNumber="157">
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<typeStatus id="95B43E9C38DA90B42E16EDA705B798A3">Type</typeStatus>
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.
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</paragraph>
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<paragraph id="E98664267146FA222BEFCFB8AD929293" pageId="0" pageNumber="157">
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<materialsCitation id="E1EB55E46BB4F2CB828ABDE73FB711E3" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="J. D. Hooker" country="US" location="US" specimenCount="1" typeStatus="holotype">
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America borealis,
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<collectorName id="B36E51D56A98D6A7695A6950D5001CB6">J.D. Hooker</collectorName>
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243 (
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<typeStatus id="49E8843C7CCDF3F4903971895F4A2410">holotype</typeStatus>
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: LE-TRIN (LE-TRIN-1634.01); isotype:
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<collectingCountry id="1EBA2D2B6064A376E07F4E957AF059F3" name="United States of America">US</collectingCountry>
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[fragm. ex LE-TRIN] (
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<collectingCountry id="B0A2D83C28CFF188EE0A373AB96AB126" name="United States of America">US</collectingCountry>
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00156470 [image!]))
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D90083BEAB5EED45CD0F4FDB2571884" pageId="0" pageNumber="157" type="description">
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<paragraph id="AAE63A96AE899A9DA4FE06E0260C86C0" pageId="0" pageNumber="157">Description.</paragraph>
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<paragraph id="6E8CE0B4B7D39E4269F414540DBB02DE" pageId="0" pageNumber="157">
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<emphasis id="338DB2C9AD6E3540EEC9FE49C03A43D1" bold="true" italics="true" pageId="0" pageNumber="157">Plants</emphasis>
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perennial, rhizomatous.
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<emphasis id="794A7CC35E673F25CF679C1974777161" bold="true" italics="true" pageId="0" pageNumber="157">Tillers</emphasis>
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extravaginal, with cataphylls.
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<emphasis id="C247108D9397619294C0D1BF0605C6E4" bold="true" italics="true" pageId="0" pageNumber="157">Rhizomes</emphasis>
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up to 10 cm long.
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<emphasis id="9D398F8BDE928696E93FAC873219B0A6" bold="true" italics="true" pageId="0" pageNumber="157">Culms</emphasis>
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10-70 cm long, erect or decumbent at the base, nodes 2-7, glabrous, sometimes rooting at the lower nodes, internodes glabrous.
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<emphasis id="906CF621B322797B7893F6B7FF1F62E2" bold="true" italics="true" pageId="0" pageNumber="157">Leaves</emphasis>
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mostly cauline; sheaths 2.5-13 cm long, longer or shorter than the internodes, glabrous or scaberulous; ligules 1-5 mm long, longer than wide, dorsally scaberulous, apices acute, lacerate; blades 2.5-14 cm long, 1-4 mm wide, flat, involute when drying, scaberulous on both surfaces.
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<emphasis id="DCA13698F3F6DC2F91909228A1312C04" bold="true" italics="true" pageId="0" pageNumber="157">Panicles</emphasis>
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(4-)5-20 cm long, 0.4-3 cm wide, contracted to open, dense to lax, lanceolate to narrow ovate, sometimes spiciform, sometimes partially included in the upper sheaths; branches appressed to ascending, rebranching from about mid-length, scaberulous, without spikelets near their base, inferior branches 1.5-3 cm long; pedicels 0.5-4 mm long, appressed to ascending, scaberulous.
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<emphasis id="F0BB16444E46727F3217F56AA8C0170B" bold="true" italics="true" pageId="0" pageNumber="157">Spikelets</emphasis>
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2-3.5(-4) mm long, greenish to stramineous, sometimes tinged with purple; glumes equal to subequal, lanceolate, apices acute, 1-veined, scaberulous on the keel, sometimes also on the body, lower gluma 2-3.5(-4) mm long, upper glume 1.8-3.5(-4) mm long; callus pubescent, with two bunches of trichomes, sometimes short and inconspicuous; lemmas 1.5-2.5 mm long, elliptic, apices entire, acute, or toothed, 5-nerved, veins prominent throughout or only distally, unawned, sometimes awned subapically to about mid-length, awn 0.5-1.5(-2.5) mm long, straight; paleas absent or up to 0.2 mm, veinless, glabrous; anthers 3, 0.7-1.5 mm long.
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<emphasis id="6B44E5F2096D04442A19746D70742F15" bold="true" italics="true" pageId="0" pageNumber="157">Caryopsis</emphasis>
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1-1.5 mm long, elliptic; endosperm solid. 2n= 42, 56 (
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<bibRefCitation id="C6CA6E982F79576942B13DA3FF3CF509" author="Harvey, MJ" editor="Barkworth, ME" journalOrPublisher="24) Oxford University Press, New York" pageId="0" pageNumber="157" pagination="633 - 662" publicationUrl="http://floranorthamerica.org/Agrostis" refId="B22" refString="Harvey, MJ, 2007. Agrostis. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Flora of North America North of Mexico (Vol. 24) Oxford University Press, New York: 633 - 662, http://floranorthamerica.org/Agrostis" title="Agrostis." url="http://floranorthamerica.org/Agrostis" volumeTitle="Flora of North America North of Mexico (Vol." year="2007">Harvey 2007</bibRefCitation>
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).
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</paragraph>
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<caption id="0E09AB4B33A702DD964734B3D394D864" doi="10.3897/phytokeys.230.105878.figure24" httpUri="https://binary.pensoft.net/fig/891259" pageId="0" pageNumber="157" start="Figure 24" startId="F24">
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||
<paragraph id="D56FA1519AACAC5256DBFEF4AB2C19E1" pageId="0" pageNumber="157">
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<emphasis id="CDB35F438A09C13919C03947C9DE98CC" bold="true" pageId="0" pageNumber="157">Figure 24.</emphasis>
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<taxonomicName id="D5B6ECA237EF8C8E5B9C55DCE94203B2" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pallens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="pallens">
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<emphasis id="34EB29767BDD1B6886D413AB4D04C433" italics="true" pageId="0" pageNumber="157">Agrostis pallens</emphasis>
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</taxonomicName>
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<emphasis id="3EFED0BA02AAA634EC79DE40C6B86DCF" bold="true" pageId="0" pageNumber="157">A</emphasis>
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whole plant
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<emphasis id="DA9BCAC7405CD17016FA3D5EF0AC15C3" bold="true" pageId="0" pageNumber="157">B</emphasis>
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detail of the rhizome
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<emphasis id="C08D1E10ACC789171C8D67E72B5A8E01" bold="true" pageId="0" pageNumber="157">C</emphasis>
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ligular area
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<emphasis id="75255ACE9468A7DB1DA74012C2FFB531" bold="true" pageId="0" pageNumber="157">D</emphasis>
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spikelet
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<emphasis id="E24C519E89939933EE985586F42EB254" bold="true" pageId="0" pageNumber="157">E</emphasis>
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floret, abaxial view
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<emphasis id="2EA2A59451D0BB7309FEE95A506E9468" bold="true" pageId="0" pageNumber="157">F</emphasis>
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floret, adaxial view. Based on Hendrickson 1099 (SD). Scale bars: 3 cm (
|
||
<emphasis id="5417091EEF73110882B45D17B32D58CE" bold="true" pageId="0" pageNumber="157">A</emphasis>
|
||
); 2 cm (
|
||
<emphasis id="D14DE716C366BF8C6C335115577DDF25" bold="true" pageId="0" pageNumber="157">B</emphasis>
|
||
); 2 mm (
|
||
<emphasis id="2265518396384D9455A212F6667AA7AD" bold="true" pageId="0" pageNumber="157">C</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis id="E714EFE452EF0A4E0111A12BC5551760" bold="true" pageId="0" pageNumber="157">D-F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BD104F1661408C60759174B1014B6359" pageId="0" pageNumber="157" type="anatomy and micromorphology">
|
||
<paragraph id="DF33652FD813BF9BE82F767FF7273332" pageId="0" pageNumber="157">Anatomy and micromorphology.</paragraph>
|
||
<paragraph id="8AF753C934898B75BCFE02553AFD8077" pageId="0" pageNumber="157">
|
||
Leaf blades flat in transversal section; adaxial furrows deep, narrow; adaxial ribs rounded; keel absent; first order bundles circular in outline, sheath interrupted adaxially and abaxially, abaxial sclerenchyma in strands or girders, narrowing towards the bundle, adaxial sclerenchyma in strands or t-shaped girders; second order bundles circular in outline, sheath interrupted abaxially, abaxial and adaxial sclerenchyma in strands; intercostal sclerenchyma absent; leaf margins with well-developed sclerenchyma caps, rounded; colorless cells absent (Fig.
|
||
<figureCitation id="071773ADDCD7B6550EB79CDC54403757" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Leaf blade anatomy in transversal section of Agrostis species, in general view, and details of lateral bundles. A - C A. laxissima D - F A. microphylla G - I A. pallens J - L A. perennans. Scale bars: 0.1 mm." figureDoi="10.3897/phytokeys.230.105878.figure22" httpUri="https://binary.pensoft.net/fig/891257" pageId="0" pageNumber="157">22G-I</figureCitation>
|
||
). Lemmas with transversal thickenings polygonal, wider than the unthickened portions of the wall; prickle hairs abundant (Fig.
|
||
<figureCitation id="E7767CD679BE4C32C0A20F49996BA323" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lemma surface of Agrostis observed with SEM A A. bourgaei B A. calderoniae C A. capillaris D A. elliottiana E A. exarata F A. ghiesbreghtii G A. gigantea H A. hyemalis I A. laxissima J A. microphylla K A. pallens L A. perennans. Scale bars: 15 μm." figureDoi="10.3897/phytokeys.230.105878.figure7" httpUri="https://binary.pensoft.net/fig/891242" pageId="0" pageNumber="157">7K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A62E311C7679031BC2EEF4D5D3BF5C" pageId="0" pageNumber="157" type="distribution">
|
||
<paragraph id="A6A9309E3A337A065D7E33A2EB691C49" pageId="0" pageNumber="157">Distribution and habitat.</paragraph>
|
||
<paragraph id="37C81110C1C102E16B48635751712469" pageId="0" pageNumber="157">
|
||
<taxonomicName id="0557634929E81A50E4A48F719B3D33B6" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pallens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="pallens">
|
||
<emphasis id="7D8E15D1F2D3361313EF0705A87F3627" italics="true" pageId="0" pageNumber="157">Agrostis pallens</emphasis>
|
||
</taxonomicName>
|
||
is distributed from British Columbia, Canada, to Baja California, Mexico, and is also present in Montana and Utah (
|
||
<bibRefCitation id="7E3D7467F79CF9BA66050314F8391099" author="Harvey, MJ" editor="Barkworth, ME" journalOrPublisher="24) Oxford University Press, New York" pageId="0" pageNumber="157" pagination="633 - 662" publicationUrl="http://floranorthamerica.org/Agrostis" refId="B22" refString="Harvey, MJ, 2007. Agrostis. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Flora of North America North of Mexico (Vol. 24) Oxford University Press, New York: 633 - 662, http://floranorthamerica.org/Agrostis" title="Agrostis." url="http://floranorthamerica.org/Agrostis" volumeTitle="Flora of North America North of Mexico (Vol." year="2007">Harvey 2007</bibRefCitation>
|
||
). In the study zone, this species has been collected in southern California and the Mexican state of Baja California (Fig.
|
||
<figureCitation id="3477C64D4B42B7815B5852FD7DCDC669" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Map of known geographic distribution of Agrostis species, based on herbarium specimen data A A. gigantea B A. hyemalis C A. idahoensis D A. laxissima E A. microphylla F A. pallens." figureDoi="10.3897/phytokeys.230.105878.figure18" httpUri="https://binary.pensoft.net/fig/891253" pageId="0" pageNumber="157">18F</figureCitation>
|
||
). It grows in coastal sands, stream edges, temperate forests with
|
||
<taxonomicName id="11131970F626DF32310EB2D71B350184" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="157" phylum="Coniferophyta" rank="genus">
|
||
<emphasis id="98FC82A8C2731117E23EE3D38FA95CB9" italics="true" pageId="0" pageNumber="157">Pinus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="136FEBA615CE727A2F93E760CAE131A7" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="157" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="A74D43483C066AEEDF68AB5A8A6240AD" italics="true" pageId="0" pageNumber="157">Quercus</emphasis>
|
||
</taxonomicName>
|
||
, and xeric shrublands, between 40-1635 m a.s.l. (Fig.
|
||
<figureCitation id="DB26D75D4B71D0BEB03681AAA07F6D41" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Elevation histograms of Agrostis species A all records B native species C introduced species D A. bourgaei E A. calderoniae F A. capillaris G A. elliottiana H A. exarata I A. ghiesbreghtii J A. gigantea K A. hyemalis L A. idahoensis M A. laxissima N A. microphylla O A. pallens." figureDoi="10.3897/phytokeys.230.105878.figure9" httpUri="https://binary.pensoft.net/fig/891244" pageId="0" pageNumber="157">9O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BD4AB12C4868F24C89A12BC991DB4D6" pageId="0" pageNumber="157" type="phenology">
|
||
<paragraph id="29593B6D63DE659A002E810E2BDEC9C1" pageId="0" pageNumber="157">Phenology.</paragraph>
|
||
<paragraph id="B204F11E18A1379A9A56AAE15FFBB28F" pageId="0" pageNumber="157">
|
||
Specimens with spikelets have been collected from June to August (Fig.
|
||
<figureCitation id="0B7193EE231FB6EEADCC10ADE21AA0D3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Phenology histograms of Agrostis species A all records B native species C introduced species D A. bourgaei E A. calderoniae F A. capillaris G A. elliottiana H A. exarata I A. ghiesbreghtii J A. gigantea K A. hyemalis L A. idahoensis M A. laxissima N A. microphylla O A. pallens." figureDoi="10.3897/phytokeys.230.105878.figure10" httpUri="https://binary.pensoft.net/fig/891245" pageId="0" pageNumber="157">10O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8E521EDEB31191CE3D315E756F2B5D4C" pageId="0" pageNumber="157" type="comments">
|
||
<paragraph id="D601FB827D37C9F6EC2D0633B18D544B" pageId="0" pageNumber="157">Commentaries.</paragraph>
|
||
<paragraph id="7CA91555FDD3AEC5DD8D573CF267FC8D" pageId="0" pageNumber="157">
|
||
It has been reported for other regions that panicles can reach 6(-8) cm wide (
|
||
<bibRefCitation id="6E26BE20A09C56D29C194ADF7891990C" author="Harvey, MJ" editor="Barkworth, ME" journalOrPublisher="24) Oxford University Press, New York" pageId="0" pageNumber="157" pagination="633 - 662" publicationUrl="http://floranorthamerica.org/Agrostis" refId="B22" refString="Harvey, MJ, 2007. Agrostis. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Flora of North America North of Mexico (Vol. 24) Oxford University Press, New York: 633 - 662, http://floranorthamerica.org/Agrostis" title="Agrostis." url="http://floranorthamerica.org/Agrostis" volumeTitle="Flora of North America North of Mexico (Vol." year="2007">Harvey 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D4F4ACCEB21EDF90AD2152AD8D8E2493" pageId="0" pageNumber="157">
|
||
The Mexican populations of
|
||
<taxonomicName id="879CCF6D962B550682DA8924AD863ADE" family="Poaceae" kingdom="Plantae" lsidName="A. pallens" order="Poales" pageId="0" pageNumber="157" rank="species" species="pallens">
|
||
<emphasis id="F5C47829D1A52BED90F9D10CE86C8C10" italics="true" pageId="0" pageNumber="157">A. pallens</emphasis>
|
||
</taxonomicName>
|
||
have lemmas with longer awns, up to 2.5 mm long, than those of southern California, but the other characters are congruent with the previous descriptions of this species. The plants of lower elevations have more contracted panicles than those of higher elevations, as noted previously by
|
||
<bibRefCitation id="6FEB6E311E9543DA3EBFB9AED591C69D" author="Harvey, MJ" editor="Barkworth, ME" journalOrPublisher="24) Oxford University Press, New York" pageId="0" pageNumber="157" pagination="633 - 662" publicationUrl="http://floranorthamerica.org/Agrostis" refId="B22" refString="Harvey, MJ, 2007. Agrostis. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Flora of North America North of Mexico (Vol. 24) Oxford University Press, New York: 633 - 662, http://floranorthamerica.org/Agrostis" title="Agrostis." url="http://floranorthamerica.org/Agrostis" volumeTitle="Flora of North America North of Mexico (Vol." year="2007">Harvey (2007)</bibRefCitation>
|
||
. Plants of lower elevations are often confused with
|
||
<taxonomicName id="4FABEC3DF8B984475560736F9D753006" family="Poaceae" kingdom="Plantae" lsidName="A. exarata" order="Poales" pageId="0" pageNumber="157" rank="species" species="exarata">
|
||
<emphasis id="0C9394B38255B17467BC9119A145ABE4" italics="true" pageId="0" pageNumber="157">A. exarata</emphasis>
|
||
</taxonomicName>
|
||
, from which they differ in the rhizomatous habit, mostly cauline leaves, and palea absent or up to 0.2 mm long (vs. usually caespitose habit, basal and cauline leaves, palea often present, up to 0.8 mm long in
|
||
<taxonomicName id="DB96CA794069251A1F0E1E4C6E9AC642" family="Poaceae" kingdom="Plantae" lsidName="A. exarata" order="Poales" pageId="0" pageNumber="157" rank="species" species="exarata">
|
||
<emphasis id="A00BCD29E56FAD35C156240BBB3F9D7C" italics="true" pageId="0" pageNumber="157">A. exarata</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6640BA470AF46E78929C28BA308B486" pageId="0" pageNumber="157" type="conservation status">
|
||
<paragraph id="7BD98A9FF8DDA868E5CC8338C41F729D" pageId="0" pageNumber="157">Conservation status.</paragraph>
|
||
<paragraph id="AC77BD331AF22C5029AD552C9CEF1040" pageId="0" pageNumber="157">
|
||
<taxonomicName id="5DF5B6D58C0D9EF8CECFA0FF83761D67" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pallens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="pallens">
|
||
<emphasis id="E4843F2BC5FD40B5E01E7EB9DA5E2C75" italics="true" pageId="0" pageNumber="157">Agrostis pallens</emphasis>
|
||
</taxonomicName>
|
||
is apparently a rare species in the study zone. It is represented by 12 collections, with several populations occurring in four protected areas. The EOO is 10,902 km2 and the AOO is 48 km2. Following the IUCN criteria, the preliminary assessment category is Vulnerable (VU).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC99C1909C79AD9D9A7FE56839E4EF16" pageId="0" pageNumber="157" type="materials_examined">
|
||
<paragraph id="FE2EA3611BF0DD116EB290D240EC2587" pageId="0" pageNumber="157">Representative specimens examined.</paragraph>
|
||
<paragraph id="4DD221D0F6DA8DB9246AD37BDAB26DDC" pageId="0" pageNumber="157">
|
||
<materialsCitation id="7BA8ED9A4264871C23E89D11529B980E" collectingDate="1980-06-11" collectorName="Municipio Ensenada, S, R. Moran" country="Mexico" county="Municipio Ensenada" elevation="40" latitude="32.016735" location="Jatay" longLatPrecision="1" longitude="-116.866516" specimenCount="1" stateProvince="Baja California">
|
||
<emphasis id="0E2B680452D96FB5C1EF038DA328314E" bold="true" pageId="0" pageNumber="157">
|
||
<collectingCountry id="2FFC5085C840C81358439D136FC95090" name="Mexico">Mexico</collectingCountry>
|
||
.
|
||
<collectingRegion id="771FC47CF463C4E705245A8F563B59C1" country="Mexico" name="Baja California">Baja California</collectingRegion>
|
||
:
|
||
<collectorName id="BD0986210922040C2237655114FEB855">
|
||
<collectingCounty id="85F9DD8B0132D6589A93D0B8EEE04E8E">Municipio Ensenada</collectingCounty>
|
||
, S
|
||
</collectorName>
|
||
</emphasis>
|
||
bank of arroyo
|
||
<location id="3D3B5499977F90F327D716C47B24A154" LSID="urn:lsid:plazi:treatment:9905255EFE665EF18248CB79C264A622:3D3B5499977F90F327D716C47B24A154" country="Mexico" county="Municipio Ensenada" latitude="32.016735" longLatPrecision="1" longitude="-116.866516" name="Jatay" stateProvince="Baja California">Jatay</location>
|
||
,
|
||
<quantity id="0C6612FCE23BFFE8E79F522CB0F5626E" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="km" value="1.5">1.5 km</quantity>
|
||
from the mouth, [
|
||
<geoCoordinate id="1C0F192DBD56D82EAD1CA9B6E8D0CAA4" degrees="32.01673537" direction="north" orientation="latitude" precision="1" value="32.016735">32.01673537°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AEBCFE9725A07771019FDCA5D132C770" degrees="116.8665199" direction="west" orientation="longitude" precision="1" value="-116.866516">116.8665199°W</geoCoordinate>
|
||
],
|
||
<quantity id="8A49574ABB6C0AC545DA084762E7332A" metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">
|
||
<elevation id="A087AEE9C78EAB8A7FC45790B854C80D" metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">40 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<collectingDate id="5F715D1600BF2712ADEFEA4F89402C1E" value="1980-06-11">11 Jun 1980</collectingDate>
|
||
,
|
||
<collectorName id="3E334A6DF604BBB228B1E383FA8A4667">R. Moran</collectorName>
|
||
28770 (MEXU [*,**], SD)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="A3856213E85EFBE293FFCBB0A1397361" collectingDate="1971-08-14" collectorName="P. C. Baker" country="USA" county="Riverside County" latitude="33.746536" location="San Jacinto Mountains" longLatPrecision="1" longitude="-116.71529" municipality="Idyllwild" specimenCount="1" stateProvince="California">
|
||
<emphasis id="6BAC1EEE63D56E72ADCC6AA69CA3BDD7" bold="true" pageId="0" pageNumber="157">
|
||
<collectingCountry id="8D47A13FC1AF62CD432BD2959E2E6DDF" name="United States of America">USA</collectingCountry>
|
||
.
|
||
<collectingRegion id="9AF254095095139006943E615A5DC7A8" country="United States of America" name="California">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="44B4B0F19BFDABEBCAB7237C072E7F80">Riverside County</collectingCounty>
|
||
</emphasis>
|
||
, at
|
||
<collectingMunicipality id="AFEA02E79C0E67BE7B0EBA40506994D1">Idyllwild</collectingMunicipality>
|
||
, in
|
||
<location id="9E6F460C0C611B93CD065EA066F26B49" LSID="urn:lsid:plazi:treatment:9905255EFE665EF18248CB79C264A622:9E6F460C0C611B93CD065EA066F26B49" country="USA" county="Riverside County" latitude="33.746536" longLatPrecision="1" longitude="-116.71529" municipality="Idyllwild" name="San Jacinto Mountains" stateProvince="California">San Jacinto Mountains</location>
|
||
, [
|
||
<geoCoordinate id="C96B612B6D0F683A3CA295008C4460BA" degrees="33.746537" direction="north" orientation="latitude" precision="1" value="33.746536">33.746537°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5843852F0D2FC4506696BC194A9A05B6" degrees="116.715288" direction="west" orientation="longitude" precision="1" value="-116.71529">116.715288°W</geoCoordinate>
|
||
], 1635, m alt.
|
||
<collectingDate id="B1A04C740FBEB6341C3E6DE575DEBCDC" value="1971-08-14">14 Aug 1971</collectingDate>
|
||
,
|
||
<collectorName id="C72AF13C8E426B109CF63F792F54D865">P.C. Baker</collectorName>
|
||
7110 (LOB)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="78473590AA55F14D3D5E875AA7599422" collectingDate="2005-06-26" collectorName="L. Hendrickson, See Suppl." country="United States of America" county="San Diego County" elevation="100" latitude="32.9745" location="Los Caballos" longLatPrecision="46" longitude="-116.571" municipality="Stonewall Mine Road" specimenCount="1" stateProvince="California">
|
||
<emphasis id="9870D4B72B43F5485A7CDA8851D1CF11" bold="true" pageId="0" pageNumber="157">
|
||
<collectingCounty id="BFA016CBCDB37B15347EC98D7E8A16F6">San Diego County</collectingCounty>
|
||
</emphasis>
|
||
,
|
||
<collectingCounty id="15129AE21A95596CD98B32704E5F0018">Cuyamaca Rancho State Park</collectingCounty>
|
||
,
|
||
<quantity id="A365168063F7B39D3E91DAE96A070826" metricMagnitude="2" metricUnit="m" metricValue="6.437376" unit="mi" value="0.4">0.4 mile</quantity>
|
||
NE of intersection of state highway 79 and
|
||
<collectingMunicipality id="6E67FD875C861A2D0E34871F482A5FFC">Stonewall Mine Road</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="B59C3FBB4F30F7E8BB4214C0360B15AD" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
|
||
<elevation id="1F4E94BE36239EB510044E8E9413E458" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
NE of
|
||
<location id="1A1859DD1AE9F0A97F4ADEB831F63E7E" LSID="urn:lsid:plazi:treatment:9905255EFE665EF18248CB79C264A622:1A1859DD1AE9F0A97F4ADEB831F63E7E" country="United States of America" county="San Diego County" latitude="32.9745" longLatPrecision="46" longitude="-116.571" municipality="Stonewall Mine Road" name="Los Caballos" stateProvince="California">Los Caballos</location>
|
||
equestrian campground and
|
||
<quantity id="7D0310A1669AE264E6B8E8D9E23DC011" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
|
||
<elevation id="31E1D827C0C2DC7ED8DEEBE232D40934" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
south of
|
||
<location id="05306E58908E2850CA960FF3315A941F" LSID="urn:lsid:plazi:treatment:9905255EFE665EF18248CB79C264A622:05306E58908E2850CA960FF3315A941F" country="United States of America" county="San Diego County" latitude="32.9745" longLatPrecision="46" longitude="-116.571" municipality="Stonewall Mine Road" name="Stonewall Mine Road" stateProvince="California">Stonewall Mine Road</location>
|
||
, in north-treading drainage,
|
||
<geoCoordinate id="7882B94BFF9DF699904AA87F9B421299" degrees="32.9745" direction="north" orientation="latitude" precision="5" value="32.9745">32.9745°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6C61C17ED78DCF1996DE570CA9BEB2A4" degrees="116.571" direction="west" orientation="longitude" precision="55" value="-116.571">116.571°W</geoCoordinate>
|
||
,
|
||
<quantity id="8CE8B40D773919B73F355B1CAAC1891C" metricMagnitude="3" metricUnit="m" metricValue="1.44" unit="m" value="1440.0">
|
||
<elevation id="A98EE885461CB970ECCB5289F1324886" metricMagnitude="3" metricUnit="m" metricValue="1.44" unit="m" value="1440.0">1440 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<collectingDate id="A34F6566977353B704409E3922D5A29F" value="2005-06-26">26 Jun 2005</collectingDate>
|
||
,
|
||
<collectorName id="EBF62F1121A4FAFBFB950319B0616324">L. Hendrickson</collectorName>
|
||
1099 (SD [*, **]).
|
||
<collectorName id="E0DBADDCE6E12AFC3E10DA0EAC3D2E51">See Suppl.</collectorName>
|
||
materials 2, 3 for additional examined specimens
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |