treatments-xml/data/97/50/C3/9750C307FF8F4C3FFF07F5C4FBFEFCBC.xml

<document ID-DOI="10.1093/zoolinnean/zlac063" ID-ISSN="0024-4082" ID-Zenodo-Dep="7690954" approvalRequired="5" approvalRequired_for_treatments="5" checkinTime="1677744395091" checkinUser="plazi" docAuthor="Macaluso, Loredana, Wencker, Lukardis C M, Castrovilli, Maria, Carnevale, Giorgio &amp; Delfino, Massimo" docDate="2023" docId="9750C307FF8F4C3FFF07F5C4FBFEFCBC" docLanguage="en" docName="zlac063.pdf" docOrigin="Zoological Journal of the Linnean Society 197 (3)" docSource="https://academic.oup.com/zoolinnean/article/197/3/569/6693955" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Lyciasalamandra Veith &amp; Steinfartz 2004" docType="treatment" docVersion="2" lastPageNumber="595" masterDocId="6B69BB7FFF954C25FFBFF368FF87FFC2" masterDocTitle="A comparative atlas of selected skeletal elements of European urodeles (Amphibia: Urodela) for palaeontological investigations" masterLastPageNumber="619" masterPageNumber="569" pageNumber="595" updateTime="1677845860903" updateUser="ExternalLinkService">
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<mods:title>A comparative atlas of selected skeletal elements of European urodeles (Amphibia: Urodela) for palaeontological investigations</mods:title>
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<mods:namePart>Macaluso, Loredana</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wencker, Lukardis C M</mods:namePart>
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<mods:namePart>Castrovilli, Maria</mods:namePart>
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<mods:namePart>Carnevale, Giorgio</mods:namePart>
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<mods:namePart>Delfino, Massimo</mods:namePart>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-01</mods:number>
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<mods:number>197</mods:number>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<paragraph blockId="26.[184,721,1707,1732]" box="[184,721,1707,1732]" pageId="26" pageNumber="595">
<heading box="[184,721,1707,1732]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="595" reason="2">
<taxonomicName ID-CoL="5HJR" ID-ENA="297009" authorityName="Veith &amp; Steinfartz" authorityYear="2004" box="[184,721,1707,1732]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="genus">
<emphasis box="[184,403,1708,1731]" italics="true" pageId="26" pageNumber="595">LYCIASALAMANDRA</emphasis>
VEITH &amp; STEINFARTZ, 2004
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<subSubSection pageId="26" pageNumber="595" type="discussion">
<paragraph blockId="26.[145,760,1780,1832]" pageId="26" pageNumber="595">
<emphasis box="[145,240,1780,1801]" italics="true" pageId="26" pageNumber="595">Species:</emphasis>
<taxonomicName authority="(Steindachner, 1891)" baseAuthorityName="Steindachner" baseAuthorityYear="1891" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[254,577,1780,1801]" italics="true" pageId="26" pageNumber="595">Lyciasalamandra luschani</emphasis>
(Steindachner, 1891)
</taxonomicName>
and 
<taxonomicName authority="(Pieper, 1963)" baseAuthorityName="Pieper" baseAuthorityYear="1963" box="[264,751,1810,1832]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="helverseni">
<emphasis box="[264,585,1810,1831]" italics="true" pageId="26" pageNumber="595">Lyciasalamandra helverseni</emphasis>
(Pieper, 1963)
</taxonomicName>
.
</paragraph>
<paragraph blockId="26.[145,761,1857,1910]" lastBlockId="26.[809,1426,196,894]" pageId="26" pageNumber="595">
No specimen of this genus was available for study, but a few observations can be made based on the work of 
<bibRefCitation author="Ozeti N" box="[838,981,196,220]" pageId="26" pageNumber="595" pagination="287 - 298" refId="ref36861" refString="Ozeti N. 1967. The morphology of the salamander Mertensiella luschani (Steindachner) and the relationships of Mertensiella and Salamandra. Copeia 1967: 287 - 298." type="journal article" year="1967">Özeti (1967)</bibRefCitation>
, who compared 
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[1172,1425,197,218]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[1172,1425,197,218]" italics="true" pageId="26" pageNumber="595">Mertensiella luschani</emphasis>
</taxonomicName>
(now 
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[874,1191,228,249]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[874,1191,228,249]" italics="true" pageId="26" pageNumber="595">Lyciasalamandra luschani</emphasis>
</taxonomicName>
) with 
<taxonomicName authorityName="* (Waga" authorityYear="1876" box="[1271,1425,228,249]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="caucasica">
<emphasis box="[1271,1425,228,249]" italics="true" pageId="26" pageNumber="595">M. caucasica</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="* (Linnaeus" authorityYear="1758" box="[867,1070,258,280]" class="Amphibia" family="Salamandridae" genus="Salamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="salamandra">
<emphasis box="[867,1070,258,280]" italics="true" pageId="26" pageNumber="595">Sa. salamandra</emphasis>
</taxonomicName>
. According to 
<bibRefCitation author="Ozeti N" box="[1260,1418,257,281]" pageId="26" pageNumber="595" pagination="287 - 298" refId="ref36861" refString="Ozeti N. 1967. The morphology of the salamander Mertensiella luschani (Steindachner) and the relationships of Mertensiella and Salamandra. Copeia 1967: 287 - 298." type="journal article" year="1967">Özeti (1967)</bibRefCitation>
, the otic–occipitum complex of 
<taxonomicName authorityName="Veith &amp; Steinfartz" authorityYear="2004" box="[1205,1425,289,310]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="genus">
<emphasis box="[1205,1425,289,310]" italics="true" pageId="26" pageNumber="595">Lyciasalamandra</emphasis>
</taxonomicName>
more closely resembles that of 
<taxonomicName authorityName="* (Waga" authorityYear="1876" box="[1199,1358,320,341]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="caucasica">
<emphasis box="[1199,1358,320,341]" italics="true" pageId="26" pageNumber="595">M. caucasica</emphasis>
</taxonomicName>
than that of 
<taxonomicName authorityName="* (Linnaeus" authorityYear="1758" box="[896,1085,350,372]" class="Amphibia" family="Salamandridae" genus="Salamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="salamandra">
<emphasis box="[896,1085,350,372]" italics="true" pageId="26" pageNumber="595">Sa. salamandra</emphasis>
</taxonomicName>
, in having the prominentiae semicircularis anterioris and posterioris more visible on the dorsal surface than in the last species. Also, 
<bibRefCitation author="Ozeti N" box="[809,962,441,465]" pageId="26" pageNumber="595" pagination="287 - 298" refId="ref36861" refString="Ozeti N. 1967. The morphology of the salamander Mertensiella luschani (Steindachner) and the relationships of Mertensiella and Salamandra. Copeia 1967: 287 - 298." type="journal article" year="1967">Özeti (1967)</bibRefCitation>
stated that ‘the anterior part of the otic capsule appears strongly displaced laterally in 
<taxonomicName authorityName="* (Linnaeus" authorityYear="1758" box="[844,1043,504,525]" class="Amphibia" family="Salamandridae" genus="Salamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="salamandra">
<emphasis box="[844,1043,504,525]" italics="true" pageId="26" pageNumber="595">Sa. salamandra</emphasis>
</taxonomicName>
’, but not in the other species, and described the fenestra ovalis as more ventral in 
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[843,994,565,587]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[843,994,565,587]" italics="true" pageId="26" pageNumber="595">Ly. luschani</emphasis>
</taxonomicName>
than in 
<taxonomicName authorityName="Garsault" authorityYear="1764" box="[1106,1260,565,586]" class="Amphibia" family="Salamandridae" genus="Salamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="genus">
<emphasis box="[1106,1260,565,586]" italics="true" pageId="26" pageNumber="595">Salamandra</emphasis>
</taxonomicName>
. The atlas is described as short and posteriorly widened, with an aliform projection (herein inferior crests) on each side of the centrum. The general shape of this vertebra is similar to that of 
<taxonomicName authorityName="* (Waga" authorityYear="1876" box="[1016,1167,688,709]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="caucasica">
<emphasis box="[1016,1167,688,709]" italics="true" pageId="26" pageNumber="595">M. caucasica</emphasis>
</taxonomicName>
. The general shape of the vertebrae of 
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[991,1125,718,740]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[991,1125,718,740]" italics="true" pageId="26" pageNumber="595">Ly. luschani</emphasis>
</taxonomicName>
seems to appear similar to those of 
<taxonomicName authorityName="* (Linnaeus" authorityYear="1758" box="[902,1083,749,770]" class="Amphibia" family="Salamandridae" genus="Salamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="salamandra">
<emphasis box="[902,1083,749,770]" italics="true" pageId="26" pageNumber="595">Sa. salamandra</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="* (Waga" authorityYear="1876" box="[1137,1283,750,771]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="caucasica">
<emphasis box="[1137,1283,750,771]" italics="true" pageId="26" pageNumber="595">M. caucasica</emphasis>
</taxonomicName>
, but slightly wider and with a lower neural crest in 
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[1276,1419,780,801]" class="Amphibia" family="Salamandridae" genus="Lyciasalamandra" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="luschani">
<emphasis box="[1276,1419,780,801]" italics="true" pageId="26" pageNumber="595">Ly. luschani</emphasis>
</taxonomicName>
. The vertebrae of 
<taxonomicName authorityName="* (Waga" authorityYear="1876" box="[1008,1158,811,832]" class="Amphibia" family="Salamandridae" genus="Mertensiella" kingdom="Animalia" order="Caudata" pageId="26" pageNumber="595" phylum="Chordata" rank="species" species="caucasica">
<emphasis box="[1008,1158,811,832]" italics="true" pageId="26" pageNumber="595">M. caucasica</emphasis>
</taxonomicName>
are described by 
<bibRefCitation author="Ozeti N" pageId="26" pageNumber="595" pagination="287 - 298" refId="ref36861" refString="Ozeti N. 1967. The morphology of the salamander Mertensiella luschani (Steindachner) and the relationships of Mertensiella and Salamandra. Copeia 1967: 287 - 298." type="journal article" year="1967">Özeti (1967)</bibRefCitation>
as more slender and with smaller processes than in the other two species.
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