309 lines
54 KiB
XML
309 lines
54 KiB
XML
<document id="DBA1B8C8E485A741B0D1943A88301FEF" ID-CLB-Dataset="22156" ID-DOI="10.11646/zootaxa.4778.2.4" ID-GBIF-Dataset="64190d5b-0175-4254-8d8f-c3657b2aceaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3856586" ID-ZooBank="5ACCAA9D-0C9B-4435-B665-15A0C79E2D6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589444738901" checkinUser="plazi" docAuthor="Lan, Tianqi, Yao, Zhiyuan, Zheng, Guo, Wongprom, Prasit & Li, Shuqiang" docDate="2020" docId="971C0A2AFFD7F473FF2AE3B6FCAFB486" docLanguage="en" docName="zootaxa.4778.2.4.pdf" docOrigin="Zootaxa 4778 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Savarna bannang Lan & Yao & Zheng & Wongprom & Li 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="315" masterDocId="6B257252FFD6F477FFBDE71EFFF5B705" masterDocTitle="Five new species of the spider genus Savarna Huber, 2005 (Araneae: Pholcidae) from Thailand" masterLastPageNumber="328" masterPageNumber="311" pageNumber="312" updateTime="1698839606252" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="33C0D4DB028A4331B2C7FDD8DCE35746">
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<mods:title id="E0D8AA1A0826A1661805D1B9C31B8B9F">Five new species of the spider genus Savarna Huber, 2005 (Araneae: Pholcidae) from Thailand</mods:title>
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</mods:titleInfo>
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<mods:name id="7D9B3E04CC25F6E2DCF09839B576AA4E" type="personal">
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<mods:roleTerm id="A7255C1FE4635869268E0C603DEA898F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E4527ACDDADDC0CFB82A854D81626260">Lan, Tianqi</mods:namePart>
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<mods:affiliation id="E6EF700B678864A3A3BD9D72E9776783">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China & ltq 0217 @ outlook. com; https: // orcid. org / 0000 - 0002 - 9491 - 9813</mods:affiliation>
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</mods:name>
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<mods:name id="BD6C2916F8DA9091F9D051A9152AB916" type="personal">
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<mods:roleTerm id="5F92AC1C0D510B6337D7569CA64455B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A30FDDA0BF101EDFFBD8AB693ED35101">Yao, Zhiyuan</mods:namePart>
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</mods:name>
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<mods:name id="0727433BA867EECABD9880F1535013A4" type="personal">
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<mods:roleTerm id="280E1EB8B02AC5304E035DBF43B00FFC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B51CFA2D33E43C9F78C790E0D392023B">Zheng, Guo</mods:namePart>
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</mods:name>
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<mods:name id="84623D93C0D982F37ED76F7C011366FA" type="personal">
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<mods:roleTerm id="9A01BBB2721FD76583E23398913B7F9D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77FDE4DB6E9C17FCD2B9CA09B86A1720">Wongprom, Prasit</mods:namePart>
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<mods:affiliation id="C3E2293DA645F2A7975E019CC4EFB66A">Faculty of Forestry, Kasetsart University, Ladyao, Jatujak, Bangkok 10903, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="9CB916FAEC3B3267970475CC5F5DFE73" type="personal">
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<mods:roleTerm id="937A209DEAE78184D33767B74D07F3A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ECD8ADD11E49DFB75854113C87711E92">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="3D06AF178EDDCE91A0C11AD52BDBC906">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:typeOfResource id="F94B72CB93E8C32E209AF59F42731D39">text</mods:typeOfResource>
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<mods:title id="B6190E16F4CB74BF96D0B50400569377">Zootaxa</mods:title>
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<mods:part id="3CB16C474421BFF6815B0F0D7FD69475">
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<mods:date id="456D71A34770B4D7943A6D45B3E10761">2020</mods:date>
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<mods:number id="C10F0E4DC1CD517957A7ADCAAFFC12A5">2020-05-14</mods:number>
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<mods:detail id="C6608383760F5ACEBD92AC96ABDC8313" type="volume">
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<mods:number id="6FC32675BD871C6C17370CB704EEB019">4778</mods:number>
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<mods:detail id="C06918E7727B3A7294BCB46329CE29B8" type="issue">
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<mods:classification id="693CB1E9E7CA20709D29910DDA18BC7E">journal article</mods:classification>
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<mods:identifier id="353B0268DF3FECDB14BB9997A4283EA5" type="CLB-Dataset">22156</mods:identifier>
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<mods:identifier id="20D48825599D18B8434AEDAC03D3A2F6" type="DOI">10.11646/zootaxa.4778.2.4</mods:identifier>
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<mods:identifier id="9C5794170F5B068DB7F9BFFCD422C540" type="GBIF-Dataset">64190d5b-0175-4254-8d8f-c3657b2aceaa</mods:identifier>
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<mods:identifier id="90A1FEDF6779601B571392AB38DC23B7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0EC93B38A5DB03979492E87A1ED52D63" type="Zenodo-Dep">3856586</mods:identifier>
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<mods:identifier id="33741248FBFAC2436FAC91D08F1CAE58" type="ZooBank">5ACCAA9D-0C9B-4435-B665-15A0C79E2D6E</mods:identifier>
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</mods:mods>
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<treatment id="971C0A2AFFD7F473FF2AE3B6FCAFB486" ID-DOI="http://doi.org/10.5281/zenodo.3856750" ID-GBIF-Taxon="164133559" ID-Zenodo-Dep="3856750" LSID="urn:lsid:plazi:treatment:971C0A2AFFD7F473FF2AE3B6FCAFB486" httpUri="http://treatment.plazi.org/id/971C0A2AFFD7F473FF2AE3B6FCAFB486" lastPageId="4" lastPageNumber="315" pageId="1" pageNumber="312">
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<subSubSection id="57AFE8B7FFD7F476FF2AE3B6FDB9B3C7" box="[151,588,1191,1218]" pageId="1" pageNumber="312" type="nomenclature">
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<paragraph id="1F0ABB3CFFD7F476FF2AE3B6FDB9B3C7" blockId="1.[151,588,1191,1254]" box="[151,588,1191,1218]" pageId="1" pageNumber="312">
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<heading id="44420C50FFD7F476FF2AE3B6FDB9B3C7" bold="true" box="[151,588,1191,1218]" fontSize="11" level="1" pageId="1" pageNumber="312" reason="1">
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<emphasis id="2DC1672EFFD7F476FF2AE3B6FDB9B3C7" bold="true" box="[151,588,1191,1218]" pageId="1" pageNumber="312">
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<taxonomicName id="D8B5C0BFFFD7F476FF2AE3B6FE1DB3C4" ID-CoL="948Q8" authority="Yao & Li" authorityName="Lan & Yao & Zheng & Wongprom & Li" authorityYear="2020" box="[151,488,1191,1218]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="bannang" status="sp. nov.">
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<emphasis id="2DC1672EFFD7F476FF2AE3B6FE85B3C7" bold="true" box="[151,368,1191,1218]" italics="true" pageId="1" pageNumber="312">Savarna bannang</emphasis>
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Yao & Li
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</taxonomicName>
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<taxonomicNameLabel id="36F2DA55FFD7F476FE52E3B6FDB9B3C7" box="[495,588,1192,1218]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="57AFE8B7FFD7F476FF2AE3D5FF0DB3E3" box="[151,248,1227,1254]" pageId="1" pageNumber="312" type="description">
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<paragraph id="1F0ABB3CFFD7F476FF2AE3D5FF0DB3E3" blockId="1.[151,588,1191,1254]" box="[151,248,1227,1254]" pageId="1" pageNumber="312">
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<figureCitation id="878EA7B9FFD7F476FF2AE3D5FF0DB3E3" box="[151,248,1227,1254]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1897,1922]" captionStartId-1="3.[151,250,1900,1925]" captionTargetBox-0="[217,1370,181,1867]" captionTargetBox-1="[218,1370,181,1874]" captionTargetId-0="figure@2.[217,1371,176,1872]" captionTargetId-1="figure@3.[218,1370,181,1875]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.3856588" figureDoi-1="http://doi.org/10.5281/zenodo.3856590" httpUri-0="https://zenodo.org/record/3856588/files/figure.png" httpUri-1="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Figs 1–2</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="57AFE8B7FFD7F476FF2AE20AFCD1B270" pageId="1" pageNumber="312" type="materials_examined">
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<paragraph id="1F0ABB3CFFD7F476FF2AE20AFCD1B270" blockId="1.[151,1437,1299,2010]" pageId="1" pageNumber="312">
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<emphasis id="2DC1672EFFD7F476FF2AE20AFEB2B22B" bold="true" box="[151,327,1299,1326]" pageId="1" pageNumber="312">Type material.</emphasis>
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||
<materialsCitation id="AFDDB161FFD7F476FEEDE20AFC73B254" ID-GBIF-Occurrence="2618695301" collectingDate="2015-10-22" collectorName="P. Wongprom" country="Thailand" elevation="52" latitude="6.274361" location="Rad Cave" longLatPrecision="1" longitude="101.25078" municipality="District" pageId="1" pageNumber="312" specimenCount="1" specimenCount-male="1" stateProvince="Yala" typeStatus="holotype">
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||
<emphasis id="2DC1672EFFD7F476FEEDE20AFE33B22B" bold="true" box="[336,454,1300,1326]" pageId="1" pageNumber="312">
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||
<typeStatus id="C00E059EFFD7F476FEEDE20AFE34B22B" box="[336,449,1300,1326]" pageId="1" pageNumber="312" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
<specimenCount id="09B370B5FFD7F476FE72E20AFE11B22B" box="[463,484,1300,1326]" pageId="1" pageNumber="312" type="male">♂</specimenCount>
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(IZCAS-Ar39725),
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<location id="1A6AEDE7FFD7F476FD70E20DFCC8B22B" LSID="urn:lsid:plazi:treatment:971C0A2AFFD7F473FF2AE3B6FCAFB486:1A6AEDE7FFD7F476FD70E20DFCC8B22B" box="[717,829,1299,1326]" country="Thailand" latitude="6.274361" longLatPrecision="1" longitude="101.25078" municipality="District" name="Rad Cave" pageId="1" pageNumber="312" stateProvince="Yala">Rad Cave</location>
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||
(
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||
<geoCoordinate id="7A81DDFBFFD7F476FCF3E20DFC13B228" box="[846,998,1299,1325]" degrees="6" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="312" precision="1" seconds="27.7" value="6.274361">6°16’27.7’’N</geoCoordinate>
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,
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<geoCoordinate id="7A81DDFBFFD7F476FC4EE20DFB61B22B" box="[1011,1172,1299,1326]" degrees="101" direction="east" minutes="15" orientation="longitude" pageId="1" pageNumber="312" precision="1" seconds="2.8" value="101.25078">101°15’2.8’’E</geoCoordinate>
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||
,
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<elevation id="94985C0FFFD7F476FB1FE20DFAA5B22B" box="[1186,1360,1299,1326]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="312" unit="m" value="52.0">
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elevation
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<quantity id="D84D16D9FFD7F476FAAFE20DFAA5B22B" box="[1298,1360,1299,1326]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="312" unit="m" value="52.0">52 m</quantity>
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</elevation>
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), Ban- nang
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<collectingMunicipality id="FF6E2146FFD7F476FF68E229FED9B254" box="[213,300,1335,1361]" pageId="1" pageNumber="312">District</collectingMunicipality>
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,
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<emphasis id="2DC1672EFFD7F476FE8BE226FE9EB257" bold="true" box="[310,363,1336,1362]" pageId="1" pageNumber="312">
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||
<collectingRegion id="DD7175DEFFD7F476FE8BE226FE9EB257" box="[310,363,1336,1362]" country="Thailand" name="Yala" pageId="1" pageNumber="312">Yala</collectingRegion>
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</emphasis>
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,
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<emphasis id="2DC1672EFFD7F476FEC4E229FE13B257" bold="true" box="[377,486,1335,1362]" pageId="1" pageNumber="312">
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||
<collectingCountry id="67A2FBACFFD7F476FEC4E229FE13B257" box="[377,486,1335,1362]" name="Thailand" pageId="1" pageNumber="312">Thailand</collectingCountry>
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</emphasis>
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,
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<date id="6B0B9DFCFFD7F476FE49E226FD46B257" box="[500,691,1335,1362]" pageId="1" pageNumber="312" value="2015-10-22">
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<collectingDate id="7B4F6414FFD7F476FE49E226FD46B257" box="[500,691,1335,1362]" pageId="1" pageNumber="312" value="2015-10-22">22 October 2015</collectingDate>
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</date>
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,
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<collectorName id="B240DEEAFFD7F476FD03E226FCA0B257" box="[702,853,1336,1362]" pageId="1" pageNumber="312">P. Wongprom</collectorName>
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leg.
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||
</materialsCitation>
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||
<emphasis id="2DC1672EFFD7F476FC31E226FBF8B257" bold="true" box="[908,1037,1336,1362]" pageId="1" pageNumber="312">
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<typeStatus id="C00E059EFFD7F476FC31E226FBFDB257" box="[908,1032,1336,1362]" pageId="1" pageNumber="312" type="paratype">Paratypes</typeStatus>
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||
:
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</emphasis>
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<specimenCount id="09B370B5FFD7F476FBA9E226FBB8B257" box="[1044,1101,1336,1362]" pageId="1" pageNumber="312" type="male">2♂♂</specimenCount>
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(IZCAS-Ar39726–Ar39727),
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<materialsCitation id="AFDDB161FFD7F476FF2AE242FCD4B270" ID-GBIF-Occurrence="2618695304" box="[151,801,1371,1398]" collectingDate="2015-10-22" collectorName="P. Wongprom" country="Thailand" elevation="52" latitude="6.274361" location="Rad Cave" longLatPrecision="1" longitude="101.25078" municipality="District" pageId="1" pageNumber="312" specimenCount="2" specimenCount-female="2" stateProvince="Yala" typeStatus="paratype">
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<specimenCount id="09B370B5FFD7F476FF2AE242FF3AB273" box="[151,207,1372,1398]" pageId="1" pageNumber="312" type="female">2♀♀</specimenCount>
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(IZCAS-Ar39728–Ar39729), same data as holotype
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="57AFE8B7FFD7F476FF7AE29EFB8EB29C" box="[199,1147,1407,1434]" pageId="1" pageNumber="312" type="etymology">
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||
<paragraph id="1F0ABB3CFFD7F476FF7AE29EFB8EB29C" blockId="1.[151,1437,1299,2010]" box="[199,1147,1407,1434]" pageId="1" pageNumber="312">
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||
<emphasis id="2DC1672EFFD7F476FF7AE29EFEBBB29F" bold="true" box="[199,334,1408,1434]" pageId="1" pageNumber="312">Etymology.</emphasis>
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The specific name is a noun in apposition and refers to the
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<typeStatus id="C00E059EFFD7F476FC57E29EFBEFB29F" box="[1002,1050,1408,1434]" pageId="1" pageNumber="312">type</typeStatus>
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locality.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="57AFE8B7FFD7F476FF7AE2BDFC71B1BF" pageId="1" pageNumber="312" type="diagnosis">
|
||
<paragraph id="1F0ABB3CFFD7F476FF7AE2BDFC71B1BF" blockId="1.[151,1437,1299,2010]" pageId="1" pageNumber="312">
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<emphasis id="2DC1672EFFD7F476FF7AE2BDFEB7B2BB" bold="true" box="[199,322,1443,1470]" pageId="1" pageNumber="312">Diagnosis.</emphasis>
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The species resembles
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<taxonomicName id="D8B5C0BFFFD7F476FDF8E2BAFD23B2BB" authority="Lan & Yao & Zheng & Wongprom & Li, 2020" authorityName="Lan & Yao & Zheng & Wongprom & Li" authorityYear="2020" box="[581,726,1444,1470]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="chiangmai" status="sp. nov.">
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<emphasis id="2DC1672EFFD7F476FDF8E2BAFD23B2BB" box="[581,726,1444,1470]" italics="true" pageId="1" pageNumber="312">S. chiangmai</emphasis>
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</taxonomicName>
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<emphasis id="2DC1672EFFD7F476FD66E2BAFCC1B2BB" bold="true" box="[731,820,1444,1470]" pageId="1" pageNumber="312">
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<taxonomicNameLabel id="36F2DA55FFD7F476FD66E2BAFCC1B2BB" box="[731,820,1444,1470]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
|
||
<figureCitation id="878EA7B9FFD7F476FCFCE2BDFC51B2BB" box="[833,932,1443,1470]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1899,1924]" captionStartId-1="6.[151,250,1902,1927]" captionTargetBox-0="[226,1369,181,1864]" captionTargetBox-1="[217,1370,181,1874]" captionTargetId-0="figure@5.[218,1369,181,1873]" captionTargetId-1="figure@6.[217,1370,181,1876]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Savarna chiangmai sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 4. Savarna chiangmai sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.3856592" figureDoi-1="http://doi.org/10.5281/zenodo.3856594" httpUri-0="https://zenodo.org/record/3856592/files/figure.png" httpUri-1="https://zenodo.org/record/3856594/files/figure.png" pageId="1" pageNumber="312">Figs 3–4</figureCitation>
|
||
) with similar embolus (
|
||
<figureCitation id="878EA7B9FFD7F476FB17E2BDFAF4B2BB" box="[1194,1281,1443,1470]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2C</figureCitation>
|
||
), male chelicerae (
|
||
<figureCitation id="878EA7B9FFD7F476FF69E2D9FEDEB2E7" box="[212,299,1479,1506]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2D</figureCitation>
|
||
) and epigynum (
|
||
<figureCitation id="878EA7B9FFD7F476FE5AE2D9FDCBB2E7" box="[487,574,1479,1506]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2A</figureCitation>
|
||
) and
|
||
<taxonomicName id="D8B5C0BFFFD7F476FDC7E2D6FC6BB2E7" authority="(Simon, 1901)" baseAuthorityName="Simon" baseAuthorityYear="1901" box="[634,926,1479,1506]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="tesselata">
|
||
<emphasis id="2DC1672EFFD7F476FDC7E2D6FD02B2E7" box="[634,759,1480,1506]" italics="true" pageId="1" pageNumber="312">S. tesselata</emphasis>
|
||
(
|
||
<bibRefCitation id="7B24C6CDFFD7F476FCB8E2D9FC63B2E7" author="Simon, E." box="[773,918,1479,1506]" pageId="1" pageNumber="312" pagination="45 - 84" refId="ref7968" refString="Simon, E. (1901) On the Arachnida collected during the Skeat expedition to the Malay Peninsula. Proceedings of the Zoological Society of London, 71 (1), 45 - 84. https: // doi. org / 10.1111 / j. 1469 - 7998.1901. tb 08164. x" type="journal article" year="1901">Simon, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="7B24C6CDFFD7F476FC65E2D9FB7DB2E7" author="Huber, B. A." box="[984,1160,1479,1506]" pageId="1" pageNumber="312" pagination="61 - 114" refId="ref7384" refString="Huber, B. A. (2005) Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen, 79, 61 - 114." type="journal article" year="2005">Huber 2005: 78</bibRefCitation>
|
||
, figs 129–130, 138–140;
|
||
<bibRefCitation id="7B24C6CDFFD7F476FF2AE2F5FE6DB103" author="Huber, B. A. & Petcharad, B. & Bumrungsri, S." box="[151,408,1515,1542]" pageId="1" pageNumber="312" pagination="1 - 23" refId="ref7708" refString="Huber, B. A., Petcharad, B. & Bumrungsri, S. (2015 b) Revision of the enigmatic Southeast Asian spider genus Savarna (Araneae, Pholcidae). European Journal of Taxonomy, 160, 1 - 23. https: // doi. org / 10.5852 / ejt. 2015.160" type="journal article" year="2015">
|
||
Huber
|
||
<emphasis id="2DC1672EFFD7F476FF58E2F2FEEBB103" box="[229,286,1516,1542]" italics="true" pageId="1" pageNumber="312">et al.</emphasis>
|
||
2015b: 12
|
||
</bibRefCitation>
|
||
, figs 11–13, 32–33, 40–41, 63–73) with male chelicerae (
|
||
<figureCitation id="878EA7B9FFD7F476FB9EE2F5FB8EB103" box="[1059,1147,1515,1542]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2D</figureCitation>
|
||
) but can be distinguished by presence of long, spine-shaped subdistal sclerite dorsally (arrow
|
||
<quantity id="D84D16D9FFD7F476FC2CE10EFC49B12C" box="[913,956,1551,1578]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="312" unit="in" value="1.0">1 in</quantity>
|
||
<figureCitation id="878EA7B9FFD7F476FC7EE111FBEDB12F" box="[963,1048,1551,1578]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="1" pageNumber="312">Fig. 1C</figureCitation>
|
||
), curved distal apophysis prolaterally (arrow
|
||
<quantity id="D84D16D9FFD7F476FEA7E12AFEB0B148" box="[282,325,1587,1614]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="312" unit="in" value="2.0">2 in</quantity>
|
||
<figureCitation id="878EA7B9FFD7F476FEF1E12DFE55B14B" box="[332,416,1587,1614]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="1" pageNumber="312">Fig. 1C</figureCitation>
|
||
) and two large, distal membranous processes (arrows 3 and
|
||
<quantity id="D84D16D9FFD7F476FBFBE12AFB84B148" box="[1094,1137,1587,1614]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="1" pageNumber="312" unit="in" value="4.0">4 in</quantity>
|
||
<figureCitation id="878EA7B9FFD7F476FBC5E12DFB39B14B" box="[1144,1228,1587,1614]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="1" pageNumber="312">Fig. 1C</figureCitation>
|
||
) on procursus and by presence of large, elliptic vulval pore plates (
|
||
<figureCitation id="878EA7B9FFD7F476FD0CE149FCFCB177" box="[689,777,1623,1650]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2B</figureCitation>
|
||
); also distinguised from
|
||
<taxonomicName id="D8B5C0BFFFD7F476FBA3E146FB45B177" authority="Lan & Yao & Zheng & Wongprom & Li, 2020" authorityName="Lan & Yao & Zheng & Wongprom & Li" authorityYear="2020" box="[1054,1200,1624,1650]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="chiangmai" status="sp. nov.">
|
||
<emphasis id="2DC1672EFFD7F476FBA3E146FB45B177" box="[1054,1200,1624,1650]" italics="true" pageId="1" pageNumber="312">S. chiangmai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2DC1672EFFD7F476FB0AE146FAE5B177" bold="true" box="[1207,1296,1624,1650]" pageId="1" pageNumber="312">
|
||
<taxonomicNameLabel id="36F2DA55FFD7F476FB0AE146FAE5B177" box="[1207,1296,1624,1650]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by presence of ridge-shaped vulval anterior arch (
|
||
<figureCitation id="878EA7B9FFD7F476FD8BE165FD7BB193" box="[566,654,1659,1686]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2B</figureCitation>
|
||
); also distinguished from
|
||
<taxonomicName id="D8B5C0BFFFD7F476FC12E162FBD8B193" baseAuthorityName="Simon" baseAuthorityYear="1901" box="[943,1069,1660,1686]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="tesselata">
|
||
<emphasis id="2DC1672EFFD7F476FC12E162FBD8B193" box="[943,1069,1660,1686]" italics="true" pageId="1" pageNumber="312">S. tesselata</emphasis>
|
||
</taxonomicName>
|
||
by short and wide embolus (
|
||
<figureCitation id="878EA7B9FFD7F476FAD2E165FF4DB1BF" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2C</figureCitation>
|
||
) and by absence of medial posterior lip on epigynum (
|
||
<figureCitation id="878EA7B9FFD7F476FCA0E181FC8DB1BF" box="[797,888,1695,1722]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AFE8B7FFD7F473FF7AE1DDFB46B45A" lastPageId="4" lastPageNumber="315" pageId="1" pageNumber="312" type="description">
|
||
<paragraph id="1F0ABB3CFFD7F473FF7AE1DDFDF9B51E" blockId="1.[151,1437,1299,2010]" lastBlockId="4.[151,1437,153,899]" lastPageId="4" lastPageNumber="315" pageId="1" pageNumber="312">
|
||
<emphasis id="2DC1672EFFD7F476FF7AE1DDFE6AB1DB" bold="true" box="[199,415,1731,1758]" pageId="1" pageNumber="312">Description. Male</emphasis>
|
||
(
|
||
<emphasis id="2DC1672EFFD7F476FE0FE1DAFDECB1DB" bold="true" box="[434,537,1732,1758]" pageId="1" pageNumber="312">
|
||
<typeStatus id="C00E059EFFD7F476FE0FE1DAFDECB1DB" box="[434,537,1732,1758]" pageId="1" pageNumber="312" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
, IZCAS-Ar39725): Total length 2.78 (2.96 with clypeus), prosoma 0.86 long, 1.02 wide, opisthosoma 1.92 long, 1.28 wide. Leg I: 30.39 (7.51 + 0.43 + 7.24 + 12.30 + 2.91), leg II: 21.46 (5.93 + 0.42 + 5.19 + 8.08 + 1.84), leg III: 14.16 (4.22 + 0.35 + 3.34 + 5.03 + 1.22), leg IV: 18.33 (5.32 + 0.36 + 4.36 + 7.04 + 1.25); tibia I L/d: 59; leg formula: I-II-IV-III. Distance PME-PME 0.18, diameter PME 0.15, distance PME-ALE 0.03, AME absent. Sternum slightly wider than long (0.72/0.66). Habitus as in
|
||
<figureCitation id="878EA7B9FFD7F476FB95E04DFB9AB06B" box="[1064,1135,1875,1902]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="1" pageNumber="312">Figs 2</figureCitation>
|
||
E–F. Dorsal shield of prosoma yellowish, with black margins and wide, dark median stripe behind ocular area; ocular area and clypeus dark brown; sternum black. Legs brown, but slightly whitish on distal parts of femora and tibiae, with darker rings on subdistal parts of femora and proximal and subdistal parts of tibiae. Opisthosoma yellowish, with black subcuticular spots lying above deeper white spots dorsally and laterally and wide black stripe ventrally. Ocular area elevated, each eye triad on top of short, laterally directed hump. Thoracic furrow deep. Chelicerae as in
|
||
<figureCitation id="878EA7B9FFD2F473FB6DE7A3FADCB7D2" box="[1232,1321,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2D</figureCitation>
|
||
, with pair of proximo-lateral apophyses. Palps as in
|
||
<figureCitation id="878EA7B9FFD2F473FDC4E7FFFD49B7FE" box="[633,700,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="4" pageNumber="315">Figs 1</figureCitation>
|
||
A–B, trochanter with small dorsal apophysis (arrow in
|
||
<figureCitation id="878EA7B9FFD2F473FA86E7FFFA66B7FE" box="[1339,1427,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="4" pageNumber="315">Fig. 1A</figureCitation>
|
||
) and long ventral apophysis proximally attached to femur; procursus simple proximally but complex distally, with long, spine-shaped subdistal sclerite dorsally (arrow
|
||
<quantity id="D84D16D9FFD2F473FD49E637FCD4B646" box="[756,801,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="315" unit="in" value="1.0">1 in</quantity>
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<figureCitation id="878EA7B9FFD2F473FC97E637FC77B646" box="[810,898,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="4" pageNumber="315">Fig. 1C</figureCitation>
|
||
), curved distal apophysis prolaterally (arrow
|
||
<quantity id="D84D16D9FFD2F473FA33E637FF58B662" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="315" unit="in" value="2.0">2 in</quantity>
|
||
<figureCitation id="878EA7B9FFD2F473FF08E653FEFCB662" box="[181,265,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="4" pageNumber="315">Fig. 1C</figureCitation>
|
||
) and two large, distal membranous processes (arrows 3 and
|
||
<quantity id="D84D16D9FFD2F473FC08E653FC15B662" box="[949,992,333,359]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="315" unit="in" value="4.0">4 in</quantity>
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||
<figureCitation id="878EA7B9FFD2F473FC55E653FBCBB662" box="[1000,1086,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1922]" captionTargetBox="[217,1370,181,1867]" captionTargetId="figure@2.[217,1371,176,1872]" captionTargetPageId="2" captionText="FIGURE 1. Savarna bannang sp. nov., holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="4" pageNumber="315">Fig. 1C</figureCitation>
|
||
); bulb (
|
||
<figureCitation id="878EA7B9FFD2F473FB2AE653FB18B662" box="[1175,1261,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2C</figureCitation>
|
||
) with proximal sclerite and sclerotized embolus (provided with large, sclerotized distal apophysis, arrow
|
||
<quantity id="D84D16D9FFD2F473FB31E66FFB4DB68E" box="[1164,1208,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="315" unit="in" value="1.0">1 in</quantity>
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||
<figureCitation id="878EA7B9FFD2F473FB7DE66FFAE0B68E" box="[1216,1301,369,395]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2C</figureCitation>
|
||
) apparently containing sperm duct (sperm duct opening visible, arrow
|
||
<quantity id="D84D16D9FFD2F473FC8BE68BFC96B6AA" box="[822,867,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="315" unit="in" value="2.0">2 in</quantity>
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||
<figureCitation id="878EA7B9FFD2F473FCD1E68BFC33B6AA" box="[876,966,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2C</figureCitation>
|
||
). Retrolateral trichobothrium of tibia I at 6% proximally; prolateral trichobothrium absent on tibia I, present on other tibiae; legs with short spines in single ventral row on femur I, with vertical hairs in higher than usual density on all tibiae, without curved hairs; tarsus I with 14 distinct pseudosegments.
|
||
</paragraph>
|
||
<caption id="4BCAEBB4FFD4F475FF2AE077FAFAB0EB" ID-DOI="http://doi.org/10.5281/zenodo.3856588" ID-Zenodo-Dep="3856588" httpUri="https://zenodo.org/record/3856588/files/figure.png" pageId="2" pageNumber="313" startId="2.[151,250,1897,1922]" targetBox="[217,1370,181,1867]" targetPageId="2">
|
||
<paragraph id="1F0ABB3CFFD4F475FF2AE077FAFAB0EB" blockId="2.[151,1437,1897,2030]" pageId="2" pageNumber="313">
|
||
<emphasis id="2DC1672EFFD4F475FF2AE077FEE0B087" bold="true" box="[151,277,1897,1922]" pageId="2" pageNumber="313">FIGURE 1.</emphasis>
|
||
<taxonomicName id="D8B5C0BFFFD4F475FEA3E075FE21B087" authority="Lan & Yao & Zheng & Wongprom & Li, 2020" authorityName="Lan & Yao & Zheng & Wongprom & Li" authorityYear="2020" box="[286,468,1898,1922]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="313" phylum="Arthropoda" rank="species" species="bannang" status="sp. nov.">
|
||
<emphasis id="2DC1672EFFD4F475FEA3E075FE21B087" box="[286,468,1898,1922]" italics="true" pageId="2" pageNumber="313">Savarna bannang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2DC1672EFFD4F475FE61E074FDDAB087" bold="true" box="[476,559,1898,1922]" pageId="2" pageNumber="313">
|
||
<taxonomicNameLabel id="36F2DA55FFD4F475FE61E074FDDAB087" box="[476,559,1898,1922]" pageId="2" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male. A–B. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (A–B), 0.05 (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4BCAEBB4FFD5F474FF2AE072FB8BB0F4" ID-DOI="http://doi.org/10.5281/zenodo.3856590" ID-Zenodo-Dep="3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="3" pageNumber="314" startId="3.[151,250,1900,1925]" targetBox="[218,1370,181,1874]" targetPageId="3">
|
||
<paragraph id="1F0ABB3CFFD5F474FF2AE072FB8BB0F4" blockId="3.[151,1436,1900,2033]" pageId="3" pageNumber="314">
|
||
<emphasis id="2DC1672EFFD5F474FF2AE072FEE6B080" bold="true" box="[151,275,1900,1925]" pageId="3" pageNumber="314">FIGURE 2.</emphasis>
|
||
<taxonomicName id="D8B5C0BFFFD5F474FEA4E073FE38B080" authority="Lan & Yao & Zheng & Wongprom & Li, 2020" authorityName="Lan & Yao & Zheng & Wongprom & Li" authorityYear="2020" box="[281,461,1901,1925]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="314" phylum="Arthropoda" rank="species" species="bannang" status="sp. nov.">
|
||
<emphasis id="2DC1672EFFD5F474FEA4E073FE38B080" box="[281,461,1901,1925]" italics="true" pageId="3" pageNumber="314">Savarna bannang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2DC1672EFFD5F474FE6FE073FDD6B080" bold="true" box="[466,547,1901,1925]" pageId="3" pageNumber="314">
|
||
<taxonomicNameLabel id="36F2DA55FFD5F474FE6FE073FDD6B080" box="[466,547,1901,1925]" pageId="3" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1F0ABB3CFFD2F473FF7AE53BFC67B5F6" blockId="4.[151,1437,153,899]" pageId="4" pageNumber="315">
|
||
<emphasis id="2DC1672EFFD2F473FF7AE53BFEEBB53A" bold="true" box="[199,286,549,575]" pageId="4" pageNumber="315">Female</emphasis>
|
||
(
|
||
<emphasis id="2DC1672EFFD2F473FE8DE53BFE6EB53A" bold="true" box="[304,411,549,575]" pageId="4" pageNumber="315">
|
||
<typeStatus id="C00E059EFFD2F473FE8DE53BFE6EB53A" box="[304,411,549,575]" pageId="4" pageNumber="315" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
, IZCAS-Ar39728): Similar to male, habitus as in
|
||
<figureCitation id="878EA7B9FFD2F473FC68E53BFBEEB53A" box="[981,1051,549,575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Figs 2</figureCitation>
|
||
G–H. Total length 2.90 (3.08 with clypeus), prosoma 0.74 long, 0.92 wide, opisthosoma 2.16 long, 1.48 wide. Leg I: – (7.01 + 0.43 + 6.54 + 10.79 + –), femur II: 5.22 (other segments missing), leg III: 13.05 (3.83 + 0.38 + 3.17 + 4.43 + 1.24), leg IV: 17.25 (5.14 + 0.38 + 4.16 + 6.33 + 1.24); tibia I L/d: 55. Distance PME-PME 0.16, diameter PME 0.10, distance PME-ALE 0.04, AME absent. Sternum slightly wider than long (0.64/0.52). Epigynum (
|
||
<figureCitation id="878EA7B9FFD2F473FC10E5ABFBF1B5CA" box="[941,1028,693,719]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2A</figureCitation>
|
||
) slightly protruding. Vulva (
|
||
<figureCitation id="878EA7B9FFD2F473FA82E5ABFA61B5CA" box="[1343,1428,693,719]" captionStart="FIGURE 2" captionStartId="3.[151,250,1900,1925]" captionTargetBox="[218,1370,181,1874]" captionTargetId="figure@3.[218,1370,181,1875]" captionTargetPageId="3" captionText="FIGURE 2. Savarna bannang sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A–D), 0.50 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3856590" httpUri="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Fig. 2B</figureCitation>
|
||
) with ridge-shaped anterior arch and pair of large elliptic pore plates.
|
||
</paragraph>
|
||
<paragraph id="1F0ABB3CFFD2F473FF7AE5E2FCF5B43E" blockId="4.[151,1437,153,899]" pageId="4" pageNumber="315">
|
||
<emphasis id="2DC1672EFFD2F473FF7AE5E2FEB4B412" bold="true" box="[199,321,764,791]" pageId="4" pageNumber="315">Variation:</emphasis>
|
||
Tibia I in male
|
||
<typeStatus id="C00E059EFFD2F473FE40E5E3FD9DB412" box="[509,616,765,791]" pageId="4" pageNumber="315" type="paratype">paratypes</typeStatus>
|
||
(n = 2): 7.06 (IZCAS-Ar39726), 7.21 (IZCAS-Ar39727). Legs I and II missing in another female
|
||
<typeStatus id="C00E059EFFD2F473FE7DE43FFDD5B43E" box="[448,544,801,827]" pageId="4" pageNumber="315" type="paratype">paratype</typeStatus>
|
||
(IZCAS-Ar39729).
|
||
</paragraph>
|
||
<paragraph id="1F0ABB3CFFD2F473FF7AE45BFB46B45A" blockId="4.[151,1437,153,899]" box="[199,1203,836,863]" pageId="4" pageNumber="315">
|
||
<emphasis id="2DC1672EFFD2F473FF7AE45BFE70B45A" bold="true" box="[199,389,836,863]" pageId="4" pageNumber="315">Natural history.</emphasis>
|
||
The species was found in the twilight zone of the Rad Cave (karst area).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AFE8B7FFD2F473FF7AE476FCAFB486" box="[199,858,872,899]" pageId="4" pageNumber="315" type="distribution">
|
||
<paragraph id="1F0ABB3CFFD2F473FF7AE476FCAFB486" blockId="4.[151,1437,153,899]" box="[199,858,872,899]" pageId="4" pageNumber="315">
|
||
<emphasis id="2DC1672EFFD2F473FF7AE476FE94B486" bold="true" box="[199,353,872,899]" pageId="4" pageNumber="315">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="C00E059EFFD2F473FDDDE477FD65B486" box="[608,656,873,899]" pageId="4" pageNumber="315">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="878EA7B9FFD2F473FD46E477FCB9B486" box="[763,844,873,899]" captionStart="FIGURE 11" captionStartId="16.[151,250,1318,1343]" captionTargetBox="[151,1436,181,1293]" captionTargetId="figure@16.[151,1436,181,1293]" captionTargetPageId="16" captionText="FIGURE 11. Known distribution records of new Savarna species from Thailand. 1. Savarna bannang sp. nov.; 2. S. chiangmai sp. nov.; 3. S. huahin sp. nov.; 4. S. satun sp. nov.; 5. S. thungsong sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3856608" httpUri="https://zenodo.org/record/3856608/files/figure.png" pageId="4" pageNumber="315">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |