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<mods:title id="49E8DA6A58B03648054C02FFBF32C6E9">Xoconochcothelphusa, a new genus for Ehecatusa chiapensis (Rodríguez &amp; Smalley, 1972), with notes on Spirothelphusa Pretzmann, 1965 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<emphasis id="2FE3EA91FFA5FFC8C8A19FF7FFF55CE3" bold="true" box="[151,428,402,429]" italics="true" pageId="8" pageNumber="179">Ehecatusa mixtepensis</emphasis>
(Rodríguez &amp; Smalley, in
<bibRefCitation id="79064B72FFA5FFC8CAC09FF7FDFA5CE2" author="Smalley, A. E." box="[758,931,402,428]" pageId="8" pageNumber="179" pagination="96 - 106" refId="ref16153" refString="Smalley, A. E. (1970) A new genus of freshwater crabs from Guatemala, with a key to the Middle American genera (Crustacea, Decapoda, Pseudothelphusidae). American Midland Naturalist, 83 (1), 96 - 106. https: // doi. org / 10.2307 / 2424009" type="journal article" year="1970">Smalley, 1970</bibRefCitation>
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(
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,
<figureCitation id="85AC2A06FFA5FFC8C8CD9FD2FF505C9F" box="[251,265,439,465]" captionStart="FIGURE 3" captionStartId="10.[152,255,1086,1110]" captionTargetBox="[444,1143,237,1063]" captionTargetId="figure-57@10.[444,1144,237,1063]" captionTargetPageId="10" captionText="FIGURE 3. Ehecatusa mixtepensis (Rodríguez &amp; Smalley, in Smalley, 1970), male holotype (cl 16.8 mm, cw 26.9 mm) (CNCR 309), distal portion of left G1. A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, distal view. Abbreviations: Ac = apical cavity; Apclb = accessory prominences of caudolateral border; Ccac = central crest apical cavity; Ccclb = crenate crest of caudo-lateral border; Cmp = caudo-marginal projection; Cslp = conical spines of lateral process; Dlcmp = distal lobe of caudo-marginal projection; Map = marginal plate; Mas = marginal suture; Mep = mesial process; Sfac = setae field of apical cavity; Spch = spermatic channel." figureDoi="http://doi.org/10.5281/zenodo.13934119" httpUri="https://zenodo.org/record/13934119/files/figure.png" pageId="8" pageNumber="179">3</figureCitation>
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<emphasis id="2FE3EA91FFA5FFC8C8A19F9EFFC75F5D" box="[151,414,507,531]" italics="true" pageId="8" pageNumber="179">Epithelphusa mixtepensis</emphasis>
Rodríguez &amp; Smalley
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, in
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(key).—
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.—
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<bibRefCitation id="79064B72FFA5FFC8CCD19F9EFEB45F7C" author="Pretzmann, G." pageId="8" pageNumber="179" pagination="294 - 302" refId="ref15618" refString="Pretzmann, G. (1974) Zur Systematik Der Pseudothelphusidae (Decapoda, Brachyura). Crustaceana, 23 (3), 294 - 302. https: // doi. org / 10.1163 / 156854074 X 00820" type="journal article" year="1974">Pretzmann 1974: 297</bibRefCitation>
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.—
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<bibRefCitation id="79064B72FFA5FFC8C93C9C7FFC385F7C" author="Rodriguez, G. &amp; Smalley, A. E." box="[266,609,538,562]" pageId="8" pageNumber="179" pagination="69 - 112" refId="ref16049" refString="Rodriguez, G. &amp; Smalley, A. E. (1972) Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, serie Ciencias del Mar y Limnologia, 40, 69 - 112." type="journal article" year="1972">Rodríguez &amp; Smalley 1972: 76</bibRefCitation>
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.—
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<bibRefCitation id="79064B72FFA5FFC8CA499C7FFD3A5F7C" author="Rodriguez, G." box="[639,867,538,562]" pageId="8" pageNumber="179" pagination="1 - 224" refId="ref15957" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Faune Tropicale, 22, 1 - 224." type="journal article" year="1982">Rodriguez 1982: 126</bibRefCitation>
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(key), 127.—
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<bibRefCitation id="79064B72FFA5FFC8CBCE9C7FFB195F7C" author="Villalobos-Figueroa, A." box="[1016,1344,538,562]" pageId="8" pageNumber="179" pagination="215 - 239" refId="ref16346" refString="Villalobos-Figueroa, A. (1982) Decapoda. In: Hurlbert, S. H. &amp; Villalobos-Figueroa, A. (Eds.), Aquatic biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, pp. 215 - 239." type="book chapter" year="1982">Villalobos-Figueroa 1982: 220</bibRefCitation>
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(list).—
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Villalobos-Hiriart
<emphasis id="2FE3EA91FFA5FFC8C9B59C5CFFE85F1E" box="[387,433,568,592]" italics="true" pageId="8" pageNumber="179">et al</emphasis>
. 1993: 287
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(list).—
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<bibRefCitation id="79064B72FFA5FFC8CA4F9C5DFDD05F1F" author="Villalobos-Hiriart, J. L." box="[633,905,568,593]" pageId="8" pageNumber="179" refId="ref16458" refString="Villalobos-Hiriart, J. L. (2005) Sistematica de los cangrejos de agua dulce de Mexico, Tribu Pseudothelphusini Ortmann, 1897 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae). Analisis filogenetico, biogeografico y descripcion de especies nuevas. Ph. D. Thesis, Posgrado en Ciencias Biologicas, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Mexico City, 363 pp." type="book" year="2005">Villalobos-Hiriart 2005: 3</bibRefCitation>
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(tabla 1), 7, 8, 17, 51, 52, 58, 59, 87-89, 105, 181, 183.—Villalobos-Hiriart &amp; Álvarez 2008: 248 (tabla 1), 285, 299 (list).—
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Ng
<emphasis id="2FE3EA91FFA5FFC8CBE29C3DFA595F21" box="[980,1024,599,623]" italics="true" pageId="8" pageNumber="179">et al</emphasis>
. 2008: 173
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(list).—
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<bibRefCitation id="79064B72FFA5FFC8CCF49C32FF765FC0" author="Villalobos, J. L. &amp; Alvarez, F." pageId="8" pageNumber="179" pagination="457 - 481" refId="ref16627" refString="Villalobos, J. L. &amp; Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. https: // doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos &amp; Alvarez 2010: 466</bibRefCitation>
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, 467 (table 2), 468 (fig.), 471 (fig.), 473 (fig.).
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<emphasis id="2FE3EA91FFA5FFC8C8A19CF1FF245FE2" box="[151,381,660,684]" italics="true" pageId="8" pageNumber="179">Ehecatusa mixtepensis</emphasis>
<bibRefCitation id="79064B72FFA5FFC8C9B29CF0FC0F5FE2" author="Ng, P. K. L. &amp; Low, M. E. Y." box="[388,598,660,685]" pageId="8" pageNumber="179" pagination="34 - 46" refId="ref15233" refString="Ng, P. K. L. &amp; Low, M. E. Y. (2010) On the generic nomenclature of nine brachyuran names, with four replacement names and two nomina protecta (Crustacea: Decapoda). Zootaxa, 2489 (1), 34 - 46. https: // doi. org / 10.11646 / zootaxa. 2489.1.2" type="journal article" year="2010">Ng &amp; Low, 2010: 35</bibRefCitation>
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[
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nom. nov. for
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<emphasis id="2FE3EA91FFA5FFC8CAC29CF1FD235FE2" box="[756,890,660,684]" italics="true" pageId="8" pageNumber="179">Epithelphusa</emphasis>
Rodríguez &amp; Smalley
</taxonomicName>
, in
<bibRefCitation id="79064B72FFA5FFC8CCB59CF1FB4D5FE2" author="Smalley, A. E." box="[1155,1300,660,684]" pageId="8" pageNumber="179" pagination="96 - 106" refId="ref16153" refString="Smalley, A. E. (1970) A new genus of freshwater crabs from Guatemala, with a key to the Middle American genera (Crustacea, Decapoda, Pseudothelphusidae). American Midland Naturalist, 83 (1), 96 - 106. https: // doi. org / 10.2307 / 2424009" type="journal article" year="1970">Smalley, 1970</bibRefCitation>
].—
<treatmentCitation id="9C361092FFA5FFC8CD0E9CF1FFC55F85" author="Guinot, D. &amp; Hendrickx, M. E." page="478" pageId="8" pageNumber="179" year="2014">
<bibRefCitation id="79064B72FFA5FFC8CD0E9CF1FFC55F85" author="Guinot, D. &amp; Hendrickx, M. E." pageId="8" pageNumber="179" pagination="476 - 484" refId="ref14249" refString="Guinot, D. &amp; Hendrickx, M. E. (2014) Correct date and authorship of taxa of Middle American freshwater crabs described by Rodriguez &amp; Smalley (1972) (not 1969) and included in Smalley (1970) (Crustacea: Decapoda: Brachyura: Pseudothelphusidae). Zootaxa, 321 (4), 476 - 484. https: // doi. org / 10.11646 / zootaxa. 3821.4.6" type="journal article" year="2014">Guinot &amp; Hendrickx 2014: 478</bibRefCitation>
</treatmentCitation>
(
<tableCitation id="50150338FFA5FFC8C99E9CD6FFB65F85" box="[424,495,691,715]" captionStart="TABLE 1" captionStartId="1.[152,245,1484,1508]" captionTargetBox="[163,1407,1668,1881]" captionTargetPageId="1" captionText="TABLE 1. Specimens examined for this work (CNCR—Colección Nacional de Crustáceos; CCDB—Coleção de Crustáceos do Departamento de Biología, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo; INPA—Instituto Nacional de Pesquisas da Amazônia; UCR-MZ—Museo de Zoología, Escuela de Biología, Universidad de Costa Rica; ZSMA—Zoologische Staatssammlung München)." pageId="8" pageNumber="179">Table 1</tableCitation>
).—
<treatmentCitation id="9C361092FFA5FFC8CA209CD6FD055F85" author="Alvarez, F. &amp; Villalobos, J. L." box="[534,860,691,715]" page="253" pageId="8" pageNumber="179" year="2016">
<bibRefCitation id="79064B72FFA5FFC8CA209CD6FD055F85" author="Alvarez, F. &amp; Villalobos, J. L." box="[534,860,691,715]" pageId="8" pageNumber="179" pagination="237 - 266" refId="ref13700" refString="Alvarez, F. &amp; Villalobos, J. L. (2016) Freshwater decapod diversity and conservation in Mexico. In: Kawai, T. &amp; Cumberlidge, N. (Eds.), A global overview of the conservation of freshwater decapod crustaceans. Springer, Cham, pp. 237 - 266. https: // doi. org / 10.1007 / 978 - 3 - 319 - 42527 - 6 _ 8" type="book chapter" year="2016">Alvarez &amp; Villalobos 2016: 253</bibRefCitation>
</treatmentCitation>
(table 8.1).—
<treatmentCitation id="9C361092FFA5FFC8CBD49CD6FA885F85" author="Ojeda-Escoto, J. C." box="[994,1233,691,715]" page="16" pageId="8" pageNumber="179" year="2017">
<bibRefCitation id="79064B72FFA5FFC8CBD49CD6FA885F85" author="Ojeda-Escoto, J. C." box="[994,1233,691,715]" pageId="8" pageNumber="179" refId="ref15299" refString="Ojeda-Escoto, J. C. (2017) Sistematica y biogeografia de los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea: Decapoda). Ph. D. Dissertation, Posgrado en Ciencias Biologicas, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Mexico City, 124 pp." type="book" year="2017">Ojeda-Escoto 2017: 16</bibRefCitation>
</treatmentCitation>
, 26 (fig.), 27, 38, 39 (fig.), 44, 50, 88 (apéndice 1), 90-91 (apéndice 3), 94 (apéndice 3).—
<treatmentCitation id="9C361092FFA5FFC8CB419CB7FA8A5FA4" author="Villalobos-Hiriart, J. L. &amp; Kamanli, S. A. &amp; Alvarez, F. &amp; Garbout, A. &amp; Clark, P. F." box="[887,1235,722,746]" page="156" pageId="8" pageNumber="179" year="2019">
<bibRefCitation id="79064B72FFA5FFC8CB419CB7FA8A5FA4" author="Villalobos-Hiriart, J. L. &amp; Kamanli, S. A. &amp; Alvarez, F. &amp; Garbout, A. &amp; Clark, P. F." box="[887,1235,722,746]" pageId="8" pageNumber="179" pagination="151 - 161" refId="ref16793" refString="Villalobos-Hiriart, J. L., Kamanli, S. A., Alvarez, F., Garbout, A. &amp; Clark, P. F. (2019) Lobithelphusa mexicana Rodriguez, 1982 (Decapoda: Pseudothelphusidae): a reassessment of key characters and systematics. Zootaxa, 4586 (1), 151 - 161. https: // doi. org / 10.11646 / zootaxa. 4586.1.8" type="journal article" year="2019">
Villalobos-Hiriart
<emphasis id="2FE3EA91FFA5FFC8CC009CB6FA3B5FA4" box="[1078,1122,722,746]" italics="true" pageId="8" pageNumber="179">et al</emphasis>
. 2019: 156
</bibRefCitation>
</treatmentCitation>
(table 1).—
<treatmentCitation id="9C361092FFA5FFC8CD709CB6FF145E46" author="Alvarez, F. &amp; Ojeda, J. C. &amp; Souza-Carvalho, E. &amp; Villalobos, J. L. &amp; Magalhaes, C. &amp; Wehrtmann, I. S. &amp; Mantelatto, F. L." page="6" pageId="8" pageNumber="179" year="2020">
<bibRefCitation id="79064B72FFA5FFC8CD709CB6FF145E46" author="Alvarez, F. &amp; Ojeda, J. C. &amp; Souza-Carvalho, E. &amp; Villalobos, J. L. &amp; Magalhaes, C. &amp; Wehrtmann, I. S. &amp; Mantelatto, F. L." pageId="8" pageNumber="179" pagination="973 - 1001" refId="ref13779" refString="Alvarez, F., Ojeda, J. C., Souza-Carvalho, E., Villalobos, J. L., Magalhaes, C., Wehrtmann, I. S. &amp; Mantelatto, F. L. (2020) Revision of the higher taxonomy of Neotropical freshwater crabs of the family Pseudothelphusidae, based on multi-gene and morphological analyses. Zoological Journal of the Linnean Society, 93 (3), 973 - 1001. https: // doi. org / 10.1093 / zoolinnean / zlaa 162" type="journal article" year="2020">
Álvarez
<emphasis id="2FE3EA91FFA5FFC8C8F19C94FEAD5E46" box="[199,244,752,776]" italics="true" pageId="8" pageNumber="179">et al</emphasis>
. 2020: 6
</bibRefCitation>
</treatmentCitation>
(table 1), 10 (fig.), 13 (fig.), 20, 23 (key), supplementary material (table 1).—
<treatmentCitation id="9C361092FFA5FFC8CC6B9C94FBC15E47" author="Moreno-Juarez, E. G. &amp; Villalobos, J. L. &amp; Alvarez, F." box="[1117,1432,752,777]" page="2" pageId="8" pageNumber="179" year="2022">
<bibRefCitation id="79064B72FFA5FFC8CC6B9C94FBC15E47" author="Moreno-Juarez, E. G. &amp; Villalobos, J. L. &amp; Alvarez, F." box="[1117,1432,752,777]" pageId="8" pageNumber="179" pagination="1 - 35" refId="ref14962" refString="Moreno-Juarez, E. G., Villalobos, J. L. &amp; Alvarez, F. (2022 a) Revision of the freshwater crabs of the genus Tehuana Rodriguez and Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the description of two new species. ZooKeys, 1117, 1 - 35. https: // doi. org / 10.3897 / zookeys. 1117.85362" type="journal article" year="2022">
Moreno-Juárez
<emphasis id="2FE3EA91FFA5FFC8CD339C94FB6B5E46" box="[1285,1330,752,776]" italics="true" pageId="8" pageNumber="179">et al</emphasis>
. 2022a: 2
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="558D6508FFA5FFC8C8A19D53FC965ED7" pageId="8" pageNumber="179" type="materials_examined">
<paragraph id="1D283683FFA5FFC8C8A19D53FC965ED7" blockId="8.[151,1436,822,1029]" pageId="8" pageNumber="179">
<emphasis id="2FE3EA91FFA5FFC8C8A19D53FFDA5E1E" bold="true" box="[151,387,822,848]" pageId="8" pageNumber="179">Material examined.</emphasis>
<materialsCitation id="ADFF3CDEFFA5FFC8C9BA9D52FC965ED7" collectingDate="1964-12-10" collectionCode="CNCR" collectorName="W. J. Schaldach" country="Mexico" elevation="858" latitude="16.345053" location="San Gabriel Mixtepec" longLatPrecision="1" longitude="-97.08934" municipality="San Gabriel Mixtepec" pageId="8" pageNumber="179" specimenCode="CNCR 309" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<typeStatus id="C22C8821FFA5FFC8C9BA9D52FFA05E1F" box="[396,505,823,849]" pageId="8" pageNumber="179" type="holotype">Holotype</typeStatus>
:
<specimenCount id="0B91FD0AFFA5FFC8CA339D52FC625E1F" box="[517,571,823,849]" pageId="8" pageNumber="179" type="male">male</specimenCount>
cl
<quantity id="DA6F9B66FFA5FFC8CA549D52FC935E1F" box="[610,714,823,849]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="179" unit="mm" value="16.8">16.8 mm</quantity>
, cw
<quantity id="DA6F9B66FFA5FFC8CB379D52FD3E5E1F" box="[769,871,823,849]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="8" pageNumber="179" unit="mm" value="26.9">26.9 mm</quantity>
(
<specimenCode id="4D319EF8FFA5FFC8CB4E9D52FDA45E1F" box="[888,1021,823,849]" collectionCode="CNCR" pageId="8" pageNumber="179">CNCR 309</specimenCode>
),
<locationDeviation id="0C9A2FA1FFA5FFC8CC249D52FA215E1F" box="[1042,1144,823,849]" pageId="8" pageNumber="179">
<quantity id="DA6F9B66FFA5FFC8CC249D52FA025E1F" box="[1042,1115,823,849]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="179" unit="km" value="36.0">36 km</quantity>
N
</locationDeviation>
of
<collectingMunicipality id="FD4CACF9FFA5FFC8CC949D52FBC55E1F" box="[1186,1436,823,849]" pageId="8" pageNumber="179">San Gabriel Mixtepec</collectingMunicipality>
(approximately
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,
<geoCoordinate id="78A35044FFA5FFC8CA269D3FFC9D5E3B" box="[528,708,858,885]" degrees="97" direction="west" minutes="05" orientation="longitude" pageId="8" pageNumber="179" precision="1" seconds="21.61" value="-97.08934">97°0521.61”W</geoCoordinate>
;
<elevation id="96BAD1B0FFA5FFC8CAE09D3FFD0B5E3B" box="[726,850,858,885]" metricMagnitude="2" metricUnit="m" metricValue="8.58" pageId="8" pageNumber="179" unit="m" value="858.0">
<quantity id="DA6F9B66FFA5FFC8CAE09D3FFD7A5E3B" box="[726,803,858,885]" metricMagnitude="2" metricUnit="m" metricValue="8.58" pageId="8" pageNumber="179" unit="m" value="858.0">858 m</quantity>
asl
</elevation>
), municipality of San Gabriel Mixtepec,
<collectingRegion id="DF53F861FFA5FFC8CD779D3EFBC15E3B" box="[1345,1432,859,885]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="179">Oaxaca</collectingRegion>
,
<collectingCountry id="65807613FFA5FFC8C8A19D1AFEB65ED7" box="[151,239,895,921]" name="Mexico" pageId="8" pageNumber="179">Mexico</collectingCountry>
, coll.
<collectorName id="B0625355FFA5FFC8C9049D1AFFBE5ED7" box="[306,487,895,921]" pageId="8" pageNumber="179">W. J. Schaldach</collectorName>
,
<date id="69291043FFA5FFC8C9C49D1AFC935ED7" box="[498,714,895,921]" pageId="8" pageNumber="179" value="1964-12-10">
<collectingDate id="796DE9ABFFA5FFC8C9C49D1AFC935ED7" box="[498,714,895,921]" pageId="8" pageNumber="179" value="1964-12-10">10 December 1964</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="558D6508FFA5FFC8C8F19DC6FCC7594B" pageId="8" pageNumber="179" type="discussion">
<paragraph id="1D283683FFA5FFC8C8F19DC6FCC7594B" blockId="8.[151,1436,822,1029]" pageId="8" pageNumber="179">
<emphasis id="2FE3EA91FFA5FFC8C8F19DC6FF625EF3" bold="true" box="[199,315,931,957]" pageId="8" pageNumber="179">Remarks.</emphasis>
The only specimen existent of
<taxonomicName id="DA974D00FFA5FFC8CAAE9DC6FD605EF2" authorityName="Ng &amp; Low" authorityYear="2010" box="[664,825,931,956]" class="Malacostraca" family="Pseudothelphusidae" genus="Ehecatusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="mixtepensis">
<emphasis id="2FE3EA91FFA5FFC8CAAE9DC6FD605EF2" box="[664,825,931,956]" italics="true" pageId="8" pageNumber="179">E. mixtepensis</emphasis>
</taxonomicName>
is the
<typeStatus id="C22C8821FFA5FFC8CBB79DC6FDBF5EF3" box="[897,998,931,957]" pageId="8" pageNumber="179" type="holotype">holotype</typeStatus>
, since its collection in 1964, there have been no new samplings in the type locality, although in some nearby streams, crabs of the species
<taxonomicName id="DA974D00FFA5FFC8CCEB9DA2FFEA594B" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="sulcifrons">
<emphasis id="2FE3EA91FFA5FFC8CCEB9DA2FF5D594B" italics="true" pageId="8" pageNumber="179">Pseudothelphusa sulcifrons</emphasis>
<bibRefCitation id="79064B72FFA5FFC8C93A9D8EFFEA594B" author="Rathbun, M. J." box="[268,435,1003,1029]" pageId="8" pageNumber="179" pagination="507 - 537" refId="ref15870" refString="Rathbun, M. J. (1898) A contribution to the knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21, 507 - 537. https: // doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
</taxonomicName>
, have been captured.
</paragraph>
</subSubSection>
</treatment>
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