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<document id="48F2BBC3E0F415919F956882B0836815" ID-CLB-Dataset="39777" ID-DOI="10.11646/zootaxa.4020.3.4" ID-GBIF-Dataset="e2caeaf4-fac2-4d17-8362-5cc42ab03ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="399931" ID-ZooBank="893ACB72-A8EF-4330-BA59-5D88F46B88D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461270650673" checkinUser="plazi" docAuthor="Pauwels, Olivier S. G., Sumontha, Montri, Kunya, Kirati, Ul, Awat Ni Tik, Samphanthamit, Phamon, Wood, Perry L. &amp; Grismer, Lee L." docDate="2015" docId="915C87B59513FF93DAC0FDF08DEAF801" docLanguage="en" docName="zt04020p494.pdf" docOrigin="Zootaxa 4020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthosaura phuketensis Pauwels, Sumontha, Kunya, Ul, Samphanthamit, Wood &amp; Grismer, 2015, sp. nov." docType="treatment" docVersion="11" lastPageNumber="484" masterDocId="6D65FFCD9511FF98DA57FFD0887DFFE4" masterDocTitle="Acanthosaura phuketensis (Squamata: Agamidae), a new long-horned tree agamid from southwestern Thailand" masterLastPageNumber="494" masterPageNumber="473" pageNumber="475" updateTime="1732489655468" updateUser="ExternalLinkService">
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<mods:title id="83C099119C42EA02CAAD6BD2C255130E">Acanthosaura phuketensis (Squamata: Agamidae), a new long-horned tree agamid from southwestern Thailand</mods:title>
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<mods:namePart id="6C7ACF48727C2C221BC962D8FA63768B">Pauwels, Olivier S. G.</mods:namePart>
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<emphasis id="2B81EAB19513FF9ADAC0FDF089A5FDDE" bold="true" box="[151,472,544,570]" italics="true" pageId="2" pageNumber="475">Acanthosaura phuketensis</emphasis>
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<figureCitation id="81CE2A269513FF9ADAC0FD948957FDB8" box="[151,298,580,605]" captionStart-0="FIGURE 1 a" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3 a" captionStart-3="FIGURE 4 a" captionStart-4="FIGURE 5 a" captionStart-5="FIGURE 6 a" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1533,1555]" captionStartId-1="4.[151,250,1441,1463]" captionStartId-10="15.[151,250,1953,1975]" captionStartId-11="16.[151,250,1039,1061]" captionStartId-2="5.[151,250,1673,1695]" captionStartId-3="6.[151,250,1569,1591]" captionStartId-4="7.[151,250,1885,1907]" captionStartId-5="8.[151,250,1359,1381]" captionStartId-6="9.[151,250,1112,1134]" captionStartId-7="12.[151,250,1969,1991]" captionStartId-8="13.[151,250,1965,1987]" captionStartId-9="14.[151,250,1967,1989]" captionTargetBox-0="[151,1436,200,1501]" captionTargetBox-1="[151,1436,522,1419]" captionTargetBox-10="[210,1377,193,1932]" captionTargetBox-2="[151,1436,558,1625]" captionTargetBox-3="[151,1436,202,1528]" captionTargetBox-6="[151,1436,176,1090]" captionTargetBox-7="[151,1436,1126,1948]" captionTargetBox-8="[201,1372,193,1937]" captionTargetBox-9="[206,1370,193,1932]" captionTargetId-0="figure@3.[151,1436,195,1509]" captionTargetId-1="figure@4.[151,1436,522,1419]" captionTargetId-10="figure@15.[210,1377,193,1932]" captionTargetId-2="figure@5.[151,1436,553,1651]" captionTargetId-3="figure@6.[151,1436,193,1548]" captionTargetId-6="figure@9.[151,1436,127,1091]" captionTargetId-7="figure@12.[151,1436,1094,1948]" captionTargetId-8="figure@13.[201,1372,193,1938]" captionTargetId-9="figure@14.[206,1370,193,1932]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="15" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-6="9" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="FIGURE 1 a. Lateral view of preserved Acanthosaura phuketensis sp. nov. adult male holotype. Photo by O. S. G. Pauwels. FIGURE 1 b. Ventral view of preserved Acanthosaura phuketensis sp. nov. adult male holotype. Photo by O. S. G. Pauwels." captionText-1="FIGURE 2. Left head side of preserved adult male paratype Acanthosaura phuketensis sp. nov. IRSNB 2681. Photo by O. S. G. Pauwels." captionText-10="FIGURE 11. Live subadult female Acanthosaura phuketensis sp. nov. in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." captionText-11="FIGURE 12. Live juvenile Acanthosaura phuketensis sp. nov. in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." captionText-2="FIGURE 3 a. Dorsal view of preserved Acanthosaura phuketensis sp. nov. adult female paratype PSUZC-RT 2012.5. Photo by O. S. G. Pauwels." captionText-3="FIGURE 4 a. Dorsal view of preserved Acanthosaura phuketensis sp. nov. adult female paratype THNHM 22543. Photo by O. S. G. Pauwels." captionText-4="FIGURE 5 a. Dorsal view of preserved Acanthosaura phuketensis sp. nov. subadult male paratype QSMI 1166. Photo by O. S. G. Pauwels." captionText-5="FIGURE 6 a. Dorsal view of preserved Acanthosaura phuketensis sp. nov. subadult male paratype PSUZC-RT 2012.6. Photo by O. S. G. Pauwels." captionText-6="FIGURE 7. Live adult male Acanthosaura phuketensis sp. nov. in situ at the type locality of Cyrtodactylus ranongensis in Suk Samran District, Ranong Province, southwestern Thailand. Photo by M. Sumontha." captionText-7="FIGURE 8. Live adult male Acanthosaura phuketensis sp. nov., asleep on a branch in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." captionText-8="FIGURE 9. Live subadult male Acanthosaura phuketensis sp. nov., eating a grasshopper on a tree in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." captionText-9="FIGURE 10. Live adult female Acanthosaura phuketensis sp. nov. in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." httpUri-0="https://zenodo.org/record/399932/files/figure.png" httpUri-1="https://zenodo.org/record/399933/files/figure.png" httpUri-10="https://zenodo.org/record/399942/files/figure.png" httpUri-2="https://zenodo.org/record/399934/files/figure.png" httpUri-3="https://zenodo.org/record/399935/files/figure.png" httpUri-6="https://zenodo.org/record/399938/files/figure.png" httpUri-7="https://zenodo.org/record/399939/files/figure.png" httpUri-8="https://zenodo.org/record/399940/files/figure.png" httpUri-9="https://zenodo.org/record/399941/files/figure.png" pageId="2" pageNumber="475">Figures 112</figureCitation>
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<emphasis id="2B81EAB19513FF9ADAC0FD5C8909FD46" box="[151,372,652,674]" italics="true" pageId="2" pageNumber="475">Acanthosaura armata</emphasis>
:
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: 38
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.
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<emphasis id="2B81EAB19513FF9ADAC0FD7A8958FD24" box="[151,293,682,704]" italics="true" pageId="2" pageNumber="475">Acanthosaura</emphasis>
cf.
<emphasis id="2B81EAB19513FF9ADB1BFD7B89D3FD24" box="[332,430,683,704]" italics="true" pageId="2" pageNumber="475">crucigera</emphasis>
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:
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<bibRefCitation id="7D644B529513FF9ADBECFD7A8AF3FD24" author="Chan-ard" box="[443,654,682,704]" pageId="2" pageNumber="475" refString="Chan-ard, T., Grossmann, W., Gumprecht, A. &amp; Schulz, K. - D. (1999) Amphibians and reptiles of Peninsular Malaysia and Thailand. An illustrated checklist. Amphibien und reptilien der Halbinsel Malaysia und Thailands. Eine illustrierte Checkliste. Bushmaster Publications, Wuerselen, 240 pp." type="book" year="1999">
Chan-ard
<emphasis id="2B81EAB19513FF9AD877FD7B8A31FD24" box="[544,588,682,704]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. 1999
</bibRefCitation>
: 90
</treatmentCitation>
, left.
<taxonomicName id="DEF54D209513FF9ADAC0FD198AABFD3B" authority="Grossmann &amp; Tillack 2000: 30" authorityName="Grossmann &amp; Tillack" authorityPageNumber="30" authorityYear="2000" box="[151,726,713,735]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FD1989F0FD3B" box="[151,397,713,735]" italics="true" pageId="2" pageNumber="475">Acanthosaura crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBCCFD198AABFD3B" author="Grossmann" box="[411,726,713,735]" page="30" pageId="2" pageNumber="475" year="2000">
<bibRefCitation id="7D644B529513FF9ADBCCFD198ACEFD3B" author="Grossmann" box="[411,691,713,735]" pageId="2" pageNumber="475" refString="Grossmann, W. &amp; Tillack, F. (2000) Bemerkungen zur Herpetofauna des Khao Lak, Phang Nga, thailandische Halbinsel. Teil I: Einfuhrung; Amphibia; Reptilia: Sauria. Sauria, 22 (4), 23 - 38." type="journal article" year="2000">Grossmann &amp; Tillack 2000</bibRefCitation>
: 30
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</taxonomicName>
31, 36, 38.
<taxonomicName id="DEF54D209513FF9ADAC0FD388AE8FD1A" authority="Pauwels et al. 2000: 132" authorityName="Pauwels et al." authorityPageNumber="132" authorityYear="2000" box="[151,661,744,766]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FD3889F0FD1A" box="[151,397,744,766]" italics="true" pageId="2" pageNumber="475">Acanthosaura crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBCCFD388AE8FD1A" author="Pauwels" box="[411,661,744,766]" page="132" pageId="2" pageNumber="475" year="2000">
<bibRefCitation id="7D644B529513FF9ADBCCFD388A18FD1A" author="Pauwels" box="[411,613,744,766]" pageId="2" pageNumber="475" refString="Pauwels, O. S. G., Laohawat, O. - A., David, P., Bour, R., Dangsee, P., Puangjit, C. &amp; Chimsunchart, C. (2000) Herpetological investigations in Phang-Nga Province, southern peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilia, 4 (2), 123 - 154." type="journal article" year="2000">
Pauwels
<emphasis id="2B81EAB19513FF9ADBA2FD398A5FFD1A" box="[501,546,744,766]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. 2000
</bibRefCitation>
: 132
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</taxonomicName>
133.
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</paragraph>
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<treatmentCitationGroup id="39E5118D9513FF9ADAC0FCD68AF2FCF8" box="[151,655,774,797]" pageId="2" pageNumber="475">
<taxonomicName id="DEF54D209513FF9ADAC0FCD68AF7FCF8" ID-CoL="8ZVK" authority="Pauwels et al. 2002: 27" authorityName="Pauwels et al." authorityPageNumber="27" authorityYear="2002" box="[151,650,774,797]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FCD689F0FCF9" box="[151,397,774,797]" italics="true" pageId="2" pageNumber="475">Acanthosaura crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBCCFCD68AF7FCF8" author="Pauwels" box="[411,650,774,797]" page="27" pageId="2" pageNumber="475" year="2002">
<bibRefCitation id="7D644B529513FF9ADBCCFCD68A18FCF8" author="Pauwels" box="[411,613,774,797]" pageId="2" pageNumber="475" refString="Pauwels, O. S. G., Laohawat, O. - A., Naaktae, W., Puangjit, C., Wisutharom, T., Chimsunchart, C. &amp; David, P. (2002) Reptile and amphibian diversity in Phang-Nga Province, southern Thailand. Natural History Journal of Chulalongkorn University, 2 (1), 25 - 30." type="journal article" year="2002">
Pauwels
<emphasis id="2B81EAB19513FF9ADBA2FCD88A5FFCF8" box="[501,546,774,797]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. 2002
</bibRefCitation>
: 27
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.
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<paragraph id="194A36A39513FF9ADAC0FCF58ABCFCBE" blockId="2.[151,1040,652,888]" pageId="2" pageNumber="475">
<treatmentCitationGroup id="39E5118D9513FF9ADAC0FCF58ABCFCBE" pageId="2" pageNumber="475">
<taxonomicName id="DEF54D209513FF9ADAC0FCF58A25FCDF" ID-CoL="8ZVK" authority="Manthey 2008: 24" authorityName="Manthey" authorityPageNumber="24" authorityYear="2008" box="[151,600,805,827]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FCF589F0FCDF" box="[151,397,805,827]" italics="true" pageId="2" pageNumber="475">Acanthosaura crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBCCFCF58A25FCDF" author="Manthey" box="[411,600,805,827]" page="24" pageId="2" pageNumber="475" year="2008">
<bibRefCitation id="7D644B529513FF9ADBCCFCF58A4CFCDF" author="Manthey" box="[411,561,805,827]" pageId="2" pageNumber="475" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey 2008</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
: RA00044-4, RA00046-3; 25: RA00047-4.
<taxonomicName id="DEF54D209513FF9ADAC0FC948AC0FCBE" authority="Sumontha et al. 2012: 69" authorityName="Sumontha et al." authorityPageNumber="69" authorityYear="2012" box="[151,701,836,858]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FC948958FCBE" box="[151,293,836,858]" italics="true" pageId="2" pageNumber="475">Acanthosaura</emphasis>
cf.
<emphasis id="2B81EAB19513FF9ADB1BFC9589D3FCBE" box="[332,430,837,858]" italics="true" pageId="2" pageNumber="475">crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBECFC948AC0FCBE" author="Sumontha" box="[443,701,836,858]" page="69" pageId="2" pageNumber="475" year="2012">
<bibRefCitation id="7D644B529513FF9ADBECFC948AE5FCBE" author="Sumontha" box="[443,664,836,858]" pageId="2" pageNumber="475" refString="Sumontha, M., Pauwels, O. S. G., Kunya, K., Nitikul, A., Samphanthamit, P. &amp; Grismer, L. L. (2012) A new forest-dwelling gecko from Phuket Island, Southern Thailand, related to Cyrtodactylus macrotuberculatus (Squamata: Gekkonidae). Zootaxa, 3522, 61 - 72." type="journal article" year="2012">
Sumontha
<emphasis id="2B81EAB19513FF9AD87FFC958A28FCBE" box="[552,597,836,858]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. 2012
</bibRefCitation>
: 69
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="194A36A39513FF9ADAC0FCB28AB0FC9C" blockId="2.[151,1040,652,888]" box="[151,717,866,888]" pageId="2" pageNumber="475">
<treatmentCitationGroup id="39E5118D9513FF9ADAC0FCB28AB0FC9C" box="[151,717,866,888]" pageId="2" pageNumber="475">
<taxonomicName id="DEF54D209513FF9ADAC0FCB28AB4FC9C" ID-CoL="8ZVK" authority="Sumontha et al. 2015: 110" authorityName="Sumontha et al." authorityPageNumber="110" authorityYear="2015" box="[151,713,866,888]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="475" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19513FF9ADAC0FCB28958FC9C" box="[151,293,866,888]" italics="true" pageId="2" pageNumber="475">Acanthosaura</emphasis>
cf.
<emphasis id="2B81EAB19513FF9ADB1BFCB389D3FC9C" box="[332,430,867,888]" italics="true" pageId="2" pageNumber="475">crucigera</emphasis>
:
<treatmentCitation id="985410B29513FF9ADBECFCB28AB4FC9C" author="Sumontha" box="[443,713,866,888]" page="110" pageId="2" pageNumber="475" year="2015">
<bibRefCitation id="7D644B529513FF9ADBECFCB28AEAFC9C" author="Sumontha" box="[443,663,866,888]" pageId="2" pageNumber="475" refString="Sumontha, M., Pauwels, O. S. G., Panitvong, N., Kunya, K. &amp; Grismer, L. L. (2015) A new lowland forest Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Ranong Province, peninsular Thailand. Zootaxa, 3911 (1), 106 - 118. http: // dx. doi. org / 10.11646 / zootaxa. 3911.1.6" type="journal article" year="2015">
Sumontha
<emphasis id="2B81EAB19513FF9AD87FFCB38A28FC9C" box="[552,597,866,888]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. 2015
</bibRefCitation>
: 110
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="51EF65289513FF9ADAC0FC758957FAE3" pageId="2" pageNumber="475" type="materials_examined">
<paragraph id="194A36A39513FF9ADAC0FC758A0BFBCB" blockId="2.[151,1437,933,2010]" pageId="2" pageNumber="475">
<emphasis id="2B81EAB19513FF9ADAC0FC758976FC59" bold="true" box="[151,267,933,957]" pageId="2" pageNumber="475">
<typeStatus id="C64E88019513FF9ADAC0FC75897BFC59" box="[151,262,933,957]" pageId="2" pageNumber="475" type="holotype">Holotype</typeStatus>
.
</emphasis>
Ethanol-preserved whole adult male individual,
<collectionCode id="7FE4AE669513FF9AD910FC768BD0FC59" box="[839,941,934,957]" pageId="2" pageNumber="475">THNHM</collectionCode>
22542 (formerly Montri Sumonthas field number MS 560) from Ban Bangrong (coordinates
<collectionCode id="7FE4AE669513FF9AD941FC1D8B29FC00" box="[790,852,973,996]" pageId="2" pageNumber="475">UTM</collectionCode>
<collectionCode id="7FE4AE669513FF9AD932FC1C8BE4FC00" LSID="urn:lsid:biocol.org:col:15542" box="[869,921,972,996]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15542" name="Uppsala University, Museum of Evolution, Botany Section (Fytoteket)" pageId="2" pageNumber="475">UPS</collectionCode>
ca. 47 p0433130 0 888891 =
<geoCoordinate id="7CC150649513FF9ADF40FC1C8D47FC32" box="[1303,1338,968,996]" direction="west" orientation="longitude" pageId="2" pageNumber="475" precision="925" value="-0.13333334">0 8o</geoCoordinate>
<geoCoordinate id="7CC150649513FF9ADF6CFC1C8DE1FC01" box="[1339,1436,972,997]" direction="north" orientation="latitude" pageId="2" pageNumber="475" precision="55" value="2.465">02.465N</geoCoordinate>
98
<geoCoordinate id="7CC150649513FF9ADAE4FC3F8976FBE8" box="[179,267,1007,1036]" direction="west" orientation="longitude" pageId="2" pageNumber="475" precision="55" value="-23.588">
o
<geoCoordinate id="7CC150649513FF9ADAECFC238976FBE8" box="[187,267,1011,1036]" direction="east" orientation="longitude" pageId="2" pageNumber="475" precision="55" value="23.588">23.588</geoCoordinate>
</geoCoordinate>
E; altitude ca.
<quantity id="DE0D9B469513FF9ADBEEFC23898AFBEF" box="[441,503,1011,1035]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="475" unit="m" value="41.0">41 m</quantity>
asl), Thalang District, Phuket Island, Phuket Province, southwestern
<collectingCountry id="61E276339513FF9ADF66FC238DE5FBE8" box="[1329,1432,1011,1036]" name="Thailand" pageId="2" pageNumber="475">Thailand</collectingCountry>
. Collected by Awat Nitikul on
<date id="6D4B10639513FF9ADBB0FBC78A0FFBCB" box="[487,626,1047,1071]" pageId="2" pageNumber="475" value="2008-06-02">2 June 2008</date>
.
</paragraph>
<paragraph id="194A36A39513FF9ADA90FBEB8957FAE3" blockId="2.[151,1437,933,2010]" pageId="2" pageNumber="475">
<emphasis id="2B81EAB19513FF9ADA90FBEB893BFBB7" bold="true" box="[199,326,1083,1107]" pageId="2" pageNumber="475">
<typeStatus id="C64E88019513FF9ADA90FBEB893CFBB7" box="[199,321,1083,1107]" pageId="2" pageNumber="475" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Five ethanol-preserved whole individuals, one adult male (hemipenes everted)
<collectionCode id="7FE4AE669513FF9ADF53FBEB8D2AFBB7" box="[1284,1367,1083,1107]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="475">IRSNB</collectionCode>
2681 (formerly
<collectionCode id="7FE4AE669513FF9ADB47FB8F891FFB93" box="[272,354,1119,1143]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="475">IRSNB</collectionCode>
15141) from Phang-Nga Wildlife Breeding Station, Muang District, Phang-Nga Province, southwestern
<collectingCountry id="61E276339513FF9ADB64FB5389E4FB78" box="[307,409,1155,1180]" name="Thailand" pageId="2" pageNumber="475">Thailand</collectingCountry>
, collected by O. S. G. Pauwels on
<date id="6D4B10639513FF9AD975FB538BCAFB7F" box="[802,951,1155,1180]" pageId="2" pageNumber="475" value="1998-07-26">26 July 1998</date>
, and two adult females PSUZC-RT 2012.5 and
<collectionCode id="7FE4AE669513FF9ADA91FB788950FB5B" box="[198,301,1192,1215]" pageId="2" pageNumber="475">THNHM</collectionCode>
22543 (formerly MS 559 and MS 225 respectively) and two subadult males, PSUZC-RT 2012.6 and
<collectionCode id="7FE4AE669513FF9ADAC0FB1B88A0FB07" box="[151,221,1227,1251]" pageId="2" pageNumber="475">QSMI</collectionCode>
1166 (formerly MS 558 and MS 557 respectively) from the same locality, collecting date and collector as for the
<typeStatus id="C64E88019513FF9ADA97FB3E895BFAE3" box="[192,294,1262,1287]" pageId="2" pageNumber="475" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="51EF65289513FF9ADA90FAC28C65F9E1" pageId="2" pageNumber="475" type="diagnosis">
<paragraph id="194A36A39513FF9ADA90FAC28C65F9E1" blockId="2.[151,1437,933,2010]" pageId="2" pageNumber="475">
<emphasis id="2B81EAB19513FF9ADA90FAC2893CFACF" bold="true" box="[199,321,1298,1323]" pageId="2" pageNumber="475">Diagnosis.</emphasis>
A medium-sized species (maximum SVL
<quantity id="DE0D9B469513FF9AD967FAC38BDBFACF" box="[816,934,1299,1324]" metricMagnitude="-1" metricUnit="m" metricValue="1.2349999999999999" pageId="2" pageNumber="475" unit="mm" value="123.5">123.5 mm</quantity>
) with a single long cylindrical spine above posterior margin of eye; large spine on occiput between tympanum and nuchal crest; tympanum naked, roundish, large; moderately developed gular pouch, black in females; scales on flanks randomly intermixed with medium and large scales; nuchal crest present and strongly developed; 12-17 scales in the diastema between the nuchal and vertebral crests; vertebral crest strongly developed, composed of enlarged, pointed scales beginning at shoulder region and continuing till first 6th of tail while regularly decreasing in size; tail 1.41.7 times SVL; black collar present; black eye patch present, extending posteriorly to reach the nuchal crest.
</paragraph>
</subSubSection>
<subSubSection id="51EF65289513FF9CDA90F9C18CE3F966" lastPageId="4" lastPageNumber="477" pageId="2" pageNumber="475" type="description">
<paragraph id="194A36A39513FF9CDA90F9C18D64FE32" blockId="2.[151,1437,933,2010]" lastBlockId="4.[151,1437,151,470]" lastPageId="4" lastPageNumber="477" pageId="2" pageNumber="475">
<emphasis id="2B81EAB19513FF9ADA90F9C18A64F9CE" bold="true" box="[199,537,1553,1578]" pageId="2" pageNumber="475">
Description of the
<typeStatus id="C64E88019513FF9ADBFCF9C28A69F9CE" box="[427,532,1554,1578]" pageId="2" pageNumber="475" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male. SVL
<quantity id="DE0D9B469513FF9AD8A6F9C18B14F9CD" box="[753,873,1553,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.2349999999999999" pageId="2" pageNumber="475" unit="mm" value="123.5">123.5 mm</quantity>
. TaL
<quantity id="DE0D9B469513FF9AD9FAF9C18C59F9CD" box="[941,1060,1553,1578]" metricMagnitude="-1" metricUnit="m" metricValue="2.056" pageId="2" pageNumber="475" unit="mm" value="205.6">205.6 mm</quantity>
, tail complete. Head length
<quantity id="DE0D9B469513FF9ADF3CF9C288BBF9A9" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="2" pageNumber="475" unit="mm" value="31.4">31.4 mm</quantity>
; head moderately long (HL/SVL 25%), somewhat narrow (HW/SVL 18%), not tall (HD/HL 0.59), triangular in dorsal and lateral profile. Snout short (SL/HL 35%); interorbital and frontal regions and rostrum wide (RW/RH 238%), steepy sloping anteriorly. Canthus rostralis prominent, forming a large projecting shelf extending above eye, composed of 14/14 enlarged scales; shelf terminates with a notch anterior to the postorbital spine. Rostral moderate in size, rectangular, bordered laterally by first supralabials and posteriorly by five smaller scales; nasal concave, surrounded by 1/1 prenasal anteriorly and 3/3 postnasals posteriorly; seven scales between nasal scales; supranasals somewhat elongate; enlarged scales above orbit weakly keeled; no enlarged row of keeled scales below orbit. Eye relatively large (EYE/HL 20%), orbit very large (ORBIT/HL 33%). Prefrontal and frontal scales slightly keeled and larger than scales between orbit and supralabials; scales on occiput weakly keeled; no enlarged parietal eyespot. Extremely elongate epidermal spine above posterior margin of eye, posteriorly recurved, surrounded by 6/ 5 enlarged scales. Suborbitals small, slightly keeled, extending by a row of 6/6 enlarged keeled scales from below posterior margin of eye to anterior margin of tympanum, increasing in size posteriorly. Elongate epidermal spine on lateral margin of nape, anteriorly recurved (the left one broken near its base and healed), surrounded by a rosette of 5/5 much shorter spiny scales, themselves surrounded by a row of pointed scales. Tympanum exposed, oblong, about as wide as tall, surrounded by small conical scales. SL 11/11, rectangular, slightly decreasing in size posteriorly; mental squarish above, becoming triangular below, larger than 1st pair of
<collectingCountry id="61E276339512FF9BDEF3F9198CBFF904" box="[1188,1218,1737,1760]" name="Iceland" pageId="3" pageNumber="476">IL</collectingCountry>
; 2 postmentals of comparable size, widely in contact behind mental; 4 scales contacting the mental; chinshields enlarged, extending posteriorly past angle of jaw, separated from infralabials by one scale row anteriorly and three at angle of jaw;
<collectingCountry id="61E276339512FF9BDFD5F8C18DE1F8CC" box="[1410,1436,1809,1832]" name="Iceland" pageId="3" pageNumber="476">IL</collectingCountry>
12/12, rectangular, scales in center of series largest; gulars sharply keeled and spinose with 3 enlarged midventral rows. Dewlap very slightly extensible, gular pouch moderate. Nuchal crest composed of 7 very elongate, lanceolate scales bordered on each side by 23 rows of enlarged, spinose scales; nuchal crest followed by a diastema of 17 scales at base of nape. Dorsal body crest very developed, extending from posterior margin of diastema onto first sixth of tail; vertebral crest composed of enlarged, epidermal, laterally compressed, spinose scales, bordered by a single row of smaller paravertebral spinose scales; vertebral crest tapers slightly to base of tail, then fades progressively. Body moderately developed, triangular in cross-section. Dorsal scales small, medium-sized and large mixed without regular pattern, with keels projecting posteriorwards; scales of pectoral region and abdomen larger than dorsals, keeled, more or less arranged in transverse rows; keeled scales anterior to vent not enlarged. Limbs relatively long; dorsal and ventral scales of forelimbs keeled, spinose, about the same size. Five digits on manus; subdigital scales keeled, subdigital lamellae under 4th finger 17/16. Scales of hind limbs keeled and spinose; postfemoral scales small, interspersed with larger spinose scales. Five digits on pes; subdigital scales keeled, subdigital lamellae under 4th toe 22/23. Tail length 1.7 times SVL, tail covered with keeled spinose scales, keels on subcaudals directed posteriorly; subcaudals much longer than supracaudals; base of tail
<quantity id="DE0D9B469515FF9CDE27FE6E8CA9FE31" box="[1136,1236,446,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="4" pageNumber="477" unit="mm" value="14.3">14.3 mm</quantity>
wide.
</paragraph>
<caption id="4D8A662B9512FF9BDAC0FA2D8DFFF9D6" httpUri="https://zenodo.org/record/399932/files/figure.png" pageId="3" pageNumber="476" targetBox="[151,1436,200,1501]" targetPageId="3">
<paragraph id="194A36A39512FF9BDAC0FA2D8DFFF9D6" blockId="3.[151,1410,1533,1586]" pageId="3" pageNumber="476">
<emphasis id="2B81EAB19512FF9BDAC0FA2D8962F9F7" bold="true" box="[151,287,1533,1555]" pageId="3" pageNumber="476">FIGURE 1a.</emphasis>
Lateral view of preserved
<taxonomicName id="DEF54D209512FF9BD878FA2D8B47F9F7" box="[559,826,1533,1555]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="476" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19512FF9BD878FA2D8B47F9F7" box="[559,826,1533,1555]" italics="true" pageId="3" pageNumber="476">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9512FF9BD917FA2D8BEDF9F7" box="[832,912,1533,1555]" pageId="3" pageNumber="476" rank="species">
<emphasis id="2B81EAB19512FF9BD917FA2D8BEDF9F7" bold="true" box="[832,912,1533,1555]" pageId="3" pageNumber="476">sp. nov.</emphasis>
</taxonomicNameLabel>
adult male holotype. Photo by O. S. G. Pauwels.
<emphasis id="2B81EAB19512FF9BDAC0F9CC895DF9D6" bold="true" box="[151,288,1564,1586]" pageId="3" pageNumber="476">FIGURE 1b.</emphasis>
Ventral view of preserved
<taxonomicName id="DEF54D209512FF9BD865F9CC8B41F9D6" box="[562,828,1564,1586]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="476" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19512FF9BD865F9CC8B41F9D6" box="[562,828,1564,1586]" italics="true" pageId="3" pageNumber="476">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9512FF9BD914F9CC8BEEF9D6" box="[835,915,1564,1586]" pageId="3" pageNumber="476" rank="species">
<emphasis id="2B81EAB19512FF9BD914F9CC8BEEF9D6" bold="true" box="[835,915,1564,1586]" pageId="3" pageNumber="476">sp. nov.</emphasis>
</taxonomicNameLabel>
adult male holotype. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9515FF9CDAC0FA718973FA32" httpUri="https://zenodo.org/record/399933/files/figure.png" pageId="4" pageNumber="477" targetBox="[151,1436,522,1419]" targetPageId="4">
<paragraph id="194A36A39515FF9CDAC0FA718973FA32" blockId="4.[151,1436,1441,1494]" pageId="4" pageNumber="477">
<emphasis id="2B81EAB19515FF9CDAC0FA71896EFA53" bold="true" box="[151,275,1441,1463]" pageId="4" pageNumber="477">FIGURE 2.</emphasis>
Left head side of preserved adult male paratype
<taxonomicName id="DEF54D209515FF9CD95FFA718C6EFA53" box="[776,1043,1441,1463]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19515FF9CD95FFA718C6EFA53" box="[776,1043,1441,1463]" italics="true" pageId="4" pageNumber="477">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<emphasis id="2B81EAB19515FF9CDE4CFA718C11FA53" bold="true" box="[1051,1132,1441,1463]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="30B257CA9515FF9CDE4CFA718C11FA53" box="[1051,1132,1441,1463]" pageId="4" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
IRSNB 2681. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<paragraph id="194A36A39515FF9CDA90FA2D8CE3F966" blockId="4.[151,1437,1533,2026]" pageId="4" pageNumber="477">
<emphasis id="2B81EAB19515FF9CDA90FA2D8946F9F2" bold="true" box="[199,315,1533,1558]" pageId="4" pageNumber="477">Variation.</emphasis>
Morphometrical and meristic data for the
<typeStatus id="C64E88019515FF9CD942FA2E8B38F9F2" box="[789,837,1534,1558]" pageId="4" pageNumber="477">type</typeStatus>
series are shown in
<tableCitation id="547703189515FF9CDE71FA2D8C04F9F2" box="[1062,1145,1533,1558]" captionStart="TABLE 1" captionStartId="11.[151,239,406,429]" captionTargetBox="[159,1431,488,1980]" captionTargetPageId="11" captionText="TABLE 1. Morphometrical (in mm) and meristic data for the type series of Acanthosaura phuketensis sp. nov. For character abbreviations see material and methods. Paired meristic characters are given left / right. NA = not applicable." httpUri="http://table.plazi.org/id/4D8A662B951AFF93DAC0FE468D02FE29" pageId="4" pageNumber="477" tableUuid="4D8A662B951AFF93DAC0FE468D02FE29">Table 1</tableCitation>
. All
<typeStatus id="C64E88019515FF9CDEE7FA2E8C9DF9F2" box="[1200,1248,1534,1558]" pageId="4" pageNumber="477">type</typeStatus>
specimens show 2 postmentals. They also all show a nuchal crest composed of 7 lanceolate scales, although the size of these scales is much larger in the
<typeStatus id="C64E88019515FF9CDBD5F9958999F9BA" box="[386,484,1605,1630]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
/adult males than in the adult female or subadults. The throat of the two adult females is blackish, while it is beige with black longitudinal reticulations in the adult male
<typeStatus id="C64E88019515FF9CDE63F9B98CE7F966" box="[1076,1178,1641,1666]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="51EF65289515FF9FDA90F95D896BFD29" lastPageId="7" lastPageNumber="480" pageId="4" pageNumber="477" type="distribution">
<paragraph id="194A36A39515FF9DDA90F95D8D48FE10" blockId="4.[151,1437,1533,2026]" lastBlockId="5.[151,1437,151,501]" lastPageId="5" lastPageNumber="478" pageId="4" pageNumber="477">
<emphasis id="2B81EAB19515FF9CDA90F95D891DF942" bold="true" box="[199,352,1677,1702]" pageId="4" pageNumber="477">Distribution.</emphasis>
Within Phuket Island,
<taxonomicName id="DEF54D209515FF9CD837F95D8BFAF942" box="[608,903,1677,1702]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19515FF9CD837F95D8BFAF942" box="[608,903,1677,1702]" italics="true" pageId="4" pageNumber="477">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9515FF9CD9DAF95E8B9AF942" box="[909,999,1678,1702]" pageId="4" pageNumber="477" rank="species">
<emphasis id="2B81EAB19515FF9CD9DAF95E8B9AF942" bold="true" box="[909,999,1678,1702]" pageId="4" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
was recorded so far only from the
<typeStatus id="C64E88019515FF9CDF3CF95E8DE1F942" box="[1387,1436,1678,1702]" pageId="4" pageNumber="477">type</typeStatus>
locality and nearby in Khao Phra Thaeo Non-hunting Area (
<figureCitation id="81CE2A269515FF9CD96BF9618BC7F92E" box="[828,954,1713,1738]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="16.[151,250,1864,1886]" captionStartId-1="17.[151,250,1158,1180]" captionTargetBox-0="[346,1239,1147,1835]" captionTargetBox-1="[151,1436,176,1136]" captionTargetId-0="figure@16.[340,1247,1141,1843]" captionTargetId-1="figure@17.[151,1436,173,1137]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 13. Map of Thailand showing the type locality of Acanthosaura phuketensis sp. nov. (black dot). Courtesy of W. Sodob." captionText-1="FIGURE 14. Biotope of Acanthosaura phuketensis sp. nov. in Khao Phra Thaeo Non-hunting Area, Phuket Island, southwestern Thailand. Photo by O. S. G. Pauwels." httpUri-0="https://zenodo.org/record/399943/files/figure.png" httpUri-1="https://zenodo.org/record/399944/files/figure.png" pageId="4" pageNumber="477">Fig. 1314</figureCitation>
), but it probably occurs in other remaining forested areas of the island.
<bibRefCitation id="7D644B529515FF9CDB99F9058ABDF90A" author="Chan-ard" box="[462,704,1749,1774]" pageId="4" pageNumber="477" refString="Chan-ard, T., Grossmann, W., Gumprecht, A. &amp; Schulz, K. - D. (1999) Amphibians and reptiles of Peninsular Malaysia and Thailand. An illustrated checklist. Amphibien und reptilien der Halbinsel Malaysia und Thailands. Eine illustrierte Checkliste. Bushmaster Publications, Wuerselen, 240 pp." type="book" year="1999">
Chan-ard
<emphasis id="2B81EAB19515FF9CD86AF9068A13F90A" box="[573,622,1749,1774]" italics="true" pageId="4" pageNumber="477">et al</emphasis>
. (1999
</bibRefCitation>
: 90) illustrated two individuals, including an adult male, that they identified as
<taxonomicName id="DEF54D209515FF9CDB7DF9298A1FF8F5" box="[298,610,1785,1810]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19515FF9CDB7DF92989BAF8F6" box="[298,455,1785,1810]" italics="true" pageId="4" pageNumber="477">Acanthosaura</emphasis>
cf.
<emphasis id="2B81EAB19515FF9CDBA2F92A8A1FF8F5" box="[501,610,1786,1809]" italics="true" pageId="4" pageNumber="477">crucigera</emphasis>
</taxonomicName>
, from Khao Lak National Park in Phang-Nga Province. That same adult male and an adult female were illustrated by
<bibRefCitation id="7D644B529515FF9CD8C6F8CE8C7DF8D2" author="Grossmann" box="[657,1024,1821,1846]" pageId="4" pageNumber="477" refString="Grossmann, W. &amp; Tillack, F. (2000) Bemerkungen zur Herpetofauna des Khao Lak, Phang Nga, thailandische Halbinsel. Teil I: Einfuhrung; Amphibia; Reptilia: Sauria. Sauria, 22 (4), 23 - 38." type="journal article" year="2000">Grossmann &amp; Tillack (2000: 31)</bibRefCitation>
under
<taxonomicName id="DEF54D209515FF9CDE19F8CD8D22F8D1" box="[1102,1375,1821,1846]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19515FF9CDE19F8CD8D22F8D1" box="[1102,1375,1821,1846]" italics="true" pageId="4" pageNumber="477">Acanthosaura crucigera</emphasis>
</taxonomicName>
. One of the individuals already illustrated by
<bibRefCitation id="7D644B529515FF9CD833F8918B23F8BE" author="Chan-ard" box="[612,862,1857,1882]" pageId="4" pageNumber="477" refString="Chan-ard, T., Grossmann, W., Gumprecht, A. &amp; Schulz, K. - D. (1999) Amphibians and reptiles of Peninsular Malaysia and Thailand. An illustrated checklist. Amphibien und reptilien der Halbinsel Malaysia und Thailands. Eine illustrierte Checkliste. Bushmaster Publications, Wuerselen, 240 pp." type="book" year="1999">
Chan-ard
<emphasis id="2B81EAB19515FF9CD882F8928B75F8BE" box="[725,776,1857,1882]" italics="true" pageId="4" pageNumber="477">et al</emphasis>
. (1999
</bibRefCitation>
: 90) plus an additional individual from Khao Lak were illustrated by
<bibRefCitation id="7D644B529515FF9CDB3BF8B58A33F89A" author="Manthey" box="[364,590,1893,1918]" pageId="4" pageNumber="477" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey (2008: 24)</bibRefCitation>
. Khao Lak park is situated along the sea on the mainland, at ca.
<quantity id="DE0D9B469515FF9CDF43F8B58D26F89A" box="[1300,1371,1893,1918]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="477" unit="km" value="65.0">65 km</quantity>
north of the
<typeStatus id="C64E88019515FF9CDAB2F85A8968F846" box="[229,277,1930,1954]" pageId="4" pageNumber="477">type</typeStatus>
locality of
<taxonomicName id="DEF54D209515FF9CDBF7F8598AB7F846" box="[416,714,1929,1954]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19515FF9CDBF7F8598AB7F846" box="[416,714,1929,1954]" italics="true" pageId="4" pageNumber="477">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9515FF9CD882F8598B4FF845" box="[725,818,1929,1953]" pageId="4" pageNumber="477" rank="species">
<emphasis id="2B81EAB19515FF9CD882F8598B4FF845" bold="true" box="[725,818,1929,1953]" pageId="4" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
Based on the coloration and scalation characteristics visible on these photographs, these individuals are referrable to
<taxonomicName id="DEF54D209515FF9CD933F87D8CF6F822" box="[868,1163,1965,1990]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19515FF9CD933F87D8CF6F822" box="[868,1163,1965,1990]" italics="true" pageId="4" pageNumber="477">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9515FF9CDEC5F87E8C91F822" box="[1170,1260,1966,1990]" pageId="4" pageNumber="477" rank="species">
<emphasis id="2B81EAB19515FF9CDEC5F87E8C91F822" bold="true" box="[1170,1260,1966,1990]" pageId="4" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
The phylogeny presented by
<bibRefCitation id="7D644B529515FF9CDB64F8018AFFF80E" box="[307,642,2001,2026]" pageId="4" pageNumber="477" refString="Kalyabina-Hauf, S., Ananjeva, N. B., Joger, U., Lenk, P., Murphy, R. W., Stuart, L., Orlov, N. L., Ho, T. C. &amp; Wink, M. (2004) Molecular phylogeny of the genus Acanthosaura (Agamidae). Current Herpetology, 23 (1), 7 - 16. http: // dx. doi. org / 10.5358 / hsj. 23.7" type="journal article">
Kalyabina-Hauf
<emphasis id="2B81EAB19515FF9CDBA6F8028A5BF80E" box="[497,550,2001,2026]" italics="true" pageId="4" pageNumber="477">et al</emphasis>
. (2004)
</bibRefCitation>
and
<bibRefCitation id="7D644B529515FF9CD8E9F8018BB8F80E" box="[702,965,2001,2026]" pageId="4" pageNumber="477" refString="Ananjeva, N. B., Orlov, N. L. &amp; Kalyabina-Hauf, S. A. (2008) Species of Acanthosaura Gray, 1831 (Agamidae: Sauria, Reptilia) of Vietnam: results of molecular and morphological study. Biology Bulletin, 35 (2), 178 - 186. http: // dx. doi. org / 10.1134 / s 106235900802012 x" type="journal article">
Ananjeva
<emphasis id="2B81EAB19515FF9CD963F8028B14F80E" box="[820,873,2001,2026]" italics="true" pageId="4" pageNumber="477">et al</emphasis>
. (2008)
</bibRefCitation>
groups three
<taxonomicName id="DEF54D209515FF9CDE30F8028C8AF80D" box="[1127,1271,2002,2025]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19515FF9CDE30F8028C8AF80D" box="[1127,1271,2002,2025]" italics="true" pageId="4" pageNumber="477">A. crucigera</emphasis>
</taxonomicName>
samples from Takua Pa District with a sample (IRSNB 15141, here renumbered IRSNB 2681 and designated
<typeStatus id="C64E88019514FF9DDEA0FF488D2AFF54" box="[1271,1367,152,176]" pageId="5" pageNumber="478" type="paratype">paratype</typeStatus>
of
<taxonomicName id="DEF54D209514FF9DDFD3FF488967FF31" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DDFD3FF488967FF31" italics="true" pageId="5" pageNumber="478">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9514FF9DDB77FF6C8907FF30" box="[288,378,188,212]" pageId="5" pageNumber="478" rank="species">
<emphasis id="2B81EAB19514FF9DDB77FF6C8907FF30" bold="true" box="[288,378,188,212]" pageId="5" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
) from
<collectingCountry id="61E276339514FF9DDB9BFF6C8A79FF31" box="[460,516,188,213]" name="Thailand" pageId="5" pageNumber="478">Thai</collectingCountry>
Muang (in fact Muang) District, all from Phang-Nga Province. On the map these authors provided, the Takua Pa locality is erroneously shown as if it was located on Phuket Island.
<bibRefCitation id="7D644B529514FF9DDEBFFF0F88C1FEF9" author="Manthey" pageId="5" pageNumber="478" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey (2008: 25)</bibRefCitation>
also showed the photograph of an adult male from Khao Sok in
<collectingCountry id="61E276339514FF9DD9F2FED48B9DFEF8" box="[933,992,260,284]" name="India" pageId="5" pageNumber="478">Surat</collectingCountry>
Thani Province. The mention of
<taxonomicName id="DEF54D209514FF9DDF09FED48979FEA4" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19514FF9DDF09FED48D0AFEFF" box="[1374,1399,260,283]" italics="true" pageId="5" pageNumber="478">A.</emphasis>
cf.
<emphasis id="2B81EAB19514FF9DDAC0FEF98979FEA4" box="[151,260,297,320]" italics="true" pageId="5" pageNumber="478">crucigera</emphasis>
</taxonomicName>
from the
<typeStatus id="C64E88019514FF9DDB23FEF889D9FEA4" box="[372,420,296,320]" pageId="5" pageNumber="478">type</typeStatus>
locality of
<taxonomicName id="DEF54D209514FF9DD870FEF788B2FE81" authority="Sumontha, Pauwels, Panitvong, Kunya &amp; Grismer, 2015" authorityName="Sumontha, Pauwels, Panitvong, Kunya &amp; Grismer" authorityYear="2015" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="ranongensis">
<emphasis id="2B81EAB19514FF9DD870FEF78B28FEA4" box="[551,853,295,320]" italics="true" pageId="5" pageNumber="478">Cyrtodactylus ranongensis</emphasis>
<bibRefCitation id="7D644B529514FF9DD90AFEF788B2FE81" author="Sumontha" pageId="5" pageNumber="478" refString="Sumontha, M., Pauwels, O. S. G., Panitvong, N., Kunya, K. &amp; Grismer, L. L. (2015) A new lowland forest Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Ranong Province, peninsular Thailand. Zootaxa, 3911 (1), 106 - 118. http: // dx. doi. org / 10.11646 / zootaxa. 3911.1.6" type="journal article" year="2015">Sumontha, Pauwels, Panitvong, Kunya &amp; Grismer, 2015</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="7D644B529514FF9DDAAEFE9C8986FE81" box="[249,507,332,357]" pageId="5" pageNumber="478" refString="Sumontha, M., Pauwels, O. S. G., Panitvong, N., Kunya, K. &amp; Grismer, L. L. (2015) A new lowland forest Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Ranong Province, peninsular Thailand. Zootaxa, 3911 (1), 106 - 118. http: // dx. doi. org / 10.11646 / zootaxa. 3911.1.6" type="journal article">
Sumontha
<emphasis id="2B81EAB19514FF9DDB25FE9D89DEFE81" box="[370,419,332,357]" italics="true" pageId="5" pageNumber="478">et al</emphasis>
. (2015)
</bibRefCitation>
in Suk Samran District in Ranong Province is referrable to
<taxonomicName id="DEF54D209514FF9DDECEFE9C8D41FE81" box="[1177,1340,332,357]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DDECEFE9C8D41FE81" box="[1177,1340,332,357]" italics="true" pageId="5" pageNumber="478">A. phuketensis</emphasis>
</taxonomicName>
<emphasis id="2B81EAB19514FF9DDF15FE9C8DE1FE80" bold="true" box="[1346,1436,332,356]" pageId="5" pageNumber="478">
<taxonomicNameLabel id="30B257CA9514FF9DDF15FE9C8DE1FE80" box="[1346,1436,332,356]" pageId="5" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The known localities of
<taxonomicName id="DEF54D209514FF9DDBF9FEBF8AAAFE6C" box="[430,727,367,392]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DDBF9FEBF8AAAFE6C" box="[430,727,367,392]" italics="true" pageId="5" pageNumber="478">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9514FF9DD888FEA08B47FE6C" box="[735,826,368,392]" pageId="5" pageNumber="478" rank="species">
<emphasis id="2B81EAB19514FF9DD888FEA08B47FE6C" bold="true" box="[735,826,368,392]" pageId="5" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
, spreading over Phuket, Phang-Nga,
<collectingCountry id="61E276339514FF9DDEB2FEA08D5DFE6C" box="[1253,1312,368,392]" name="India" pageId="5" pageNumber="478">Surat</collectingCountry>
Thani and Ranong provinces, are all located on the Phuket Range, a
<quantity id="DE0D9B469514FF9DD975FE448B05FE49" box="[802,888,404,429]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="478" unit="km" value="200.0">200 km</quantity>
long continuation of the Tenasserim Range. The range of the new species is surrounded by other populations of the
<taxonomicName id="DEF54D209514FF9DD9C3FE698C7CFE34" box="[916,1025,441,464]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19514FF9DD9C3FE698C7CFE34" box="[916,1025,441,464]" italics="true" pageId="5" pageNumber="478">crucigera</emphasis>
</taxonomicName>
group with shorter horns, as is well illustrated through the photographs from various populations and localities presented by
<bibRefCitation id="7D644B529514FF9DDE2FFE0C8D4DFE10" author="Manthey" box="[1144,1328,476,501]" pageId="5" pageNumber="478" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey (2008)</bibRefCitation>
.
</paragraph>
<caption id="4D8A662B9514FF9DDAC0F9598935F959" httpUri="https://zenodo.org/record/399934/files/figure.png" pageId="5" pageNumber="478" targetBox="[151,1436,558,1625]" targetPageId="5">
<paragraph id="194A36A39514FF9DDAC0F9598935F959" blockId="5.[151,1436,1673,1786]" pageId="5" pageNumber="478">
<emphasis id="2B81EAB19514FF9DDAC0F9598962F97B" bold="true" box="[151,287,1673,1695]" pageId="5" pageNumber="478">FIGURE 3a.</emphasis>
Dorsal view of preserved
<taxonomicName id="DEF54D209514FF9DD87DF9598B49F97B" box="[554,820,1673,1695]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DD87DF9598B49F97B" box="[554,820,1673,1695]" italics="true" pageId="5" pageNumber="478">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9514FF9DD96CF9598BF7F97B" box="[827,906,1673,1695]" pageId="5" pageNumber="478" rank="species">
<emphasis id="2B81EAB19514FF9DD96CF9598BF7F97B" bold="true" box="[827,906,1673,1695]" pageId="5" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
adult female paratype PSUZC-RT 2012.5. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9514FF9DDAC0F9168935F91E" pageId="5" pageNumber="478">
<paragraph id="194A36A39514FF9DDAC0F9168935F91E" blockId="5.[151,1436,1673,1786]" pageId="5" pageNumber="478">
<emphasis id="2B81EAB19514FF9DDAC0F916895DF938" bold="true" box="[151,288,1734,1756]" pageId="5" pageNumber="478">FIGURE 3b.</emphasis>
Ventral view of preserved
<taxonomicName id="DEF54D209514FF9DD879F9168B45F938" box="[558,824,1734,1756]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DD879F9168B45F938" box="[558,824,1734,1756]" italics="true" pageId="5" pageNumber="478">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9514FF9DD969F9168BF3F938" box="[830,910,1734,1756]" pageId="5" pageNumber="478" rank="species">
<emphasis id="2B81EAB19514FF9DD969F9168BF3F938" bold="true" box="[830,910,1734,1756]" pageId="5" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
adult female paratype PSUZC-RT 2012.5. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<paragraph id="194A36A39514FF9FDA90F8F88AF2FDD9" blockId="5.[151,1436,1831,2036]" lastBlockId="7.[151,1436,151,717]" lastPageId="7" lastPageNumber="480" pageId="5" pageNumber="478">
<emphasis id="2B81EAB19514FF9DDA90F8F889F9F8A4" bold="true" box="[199,388,1831,1856]" pageId="5" pageNumber="478">Natural history.</emphasis>
<taxonomicName id="DEF54D209514FF9DDBDCF8F78ACFF8A4" box="[395,690,1831,1856]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19514FF9DDBDCF8F78ACFF8A4" box="[395,690,1831,1856]" italics="true" pageId="5" pageNumber="478">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9514FF9DD8EFF8F88B6CF8A4" box="[696,785,1832,1856]" pageId="5" pageNumber="478" rank="species">
<emphasis id="2B81EAB19514FF9DD8EFF8F88B6CF8A4" bold="true" box="[696,785,1832,1856]" pageId="5" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
is primarily an arboreal species, as shown on published
<emphasis id="2B81EAB19514FF9DDFD1F8F888BDF880" italics="true" pageId="5" pageNumber="478">in situ</emphasis>
photographs (
<bibRefCitation id="7D644B529514FF9DDB09F89B8A2DF880" author="Chan-ard" box="[350,592,1867,1892]" pageId="5" pageNumber="478" refString="Chan-ard, T., Grossmann, W., Gumprecht, A. &amp; Schulz, K. - D. (1999) Amphibians and reptiles of Peninsular Malaysia and Thailand. An illustrated checklist. Amphibien und reptilien der Halbinsel Malaysia und Thailands. Eine illustrierte Checkliste. Bushmaster Publications, Wuerselen, 240 pp." type="book" year="1999">
Chan-ard
<emphasis id="2B81EAB19514FF9DDB98F89D8A7DF880" box="[463,512,1867,1892]" italics="true" pageId="5" pageNumber="478">et al</emphasis>
., 1999
</bibRefCitation>
;
<bibRefCitation id="7D644B529514FF9DD80DF89C8BE9F880" author="Grossmann" box="[602,916,1867,1892]" pageId="5" pageNumber="478" refString="Grossmann, W. &amp; Tillack, F. (2000) Bemerkungen zur Herpetofauna des Khao Lak, Phang Nga, thailandische Halbinsel. Teil I: Einfuhrung; Amphibia; Reptilia: Sauria. Sauria, 22 (4), 23 - 38." type="journal article" year="2000">Grossmann &amp; Tillack, 2000</bibRefCitation>
;
<bibRefCitation id="7D644B529514FF9DD9C8F89B8C3EF880" author="Manthey" box="[927,1091,1867,1892]" pageId="5" pageNumber="478" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey 2008</bibRefCitation>
) and according to our personal field observations. The Phuket
<typeStatus id="C64E88019514FF9DD856F8A08A4CF86C" box="[513,561,1904,1928]" pageId="5" pageNumber="478">type</typeStatus>
specimens were collected at night while they were asleep on large trees in mature secondary forest. However this species also regularly descends on the ground.
<bibRefCitation id="7D644B529514FF9DDE34F8438D2EF848" box="[1123,1363,1939,1964]" pageId="5" pageNumber="478" refString="Pauwels, O. S. G., Laohawat, O. - A., David, P., Bour, R., Dangsee, P., Puangjit, C. &amp; Chimsunchart, C. (2000) Herpetological investigations in Phang-Nga Province, southern peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilia, 4 (2), 123 - 154." type="journal article">
Pauwels
<emphasis id="2B81EAB19514FF9DDE9FF8458C84F848" box="[1224,1273,1939,1964]" italics="true" pageId="5" pageNumber="478">et al</emphasis>
. (2000)
</bibRefCitation>
found by day an individual under a stone in the dry bed of a stream, a few meters from another individual found active on the ground, in mature secondary forest. This diurnal species can be locally abundant; many individuals were observed in mature secondary forest at the
<typeStatus id="C64E88019517FF9ED829F96B8AD3F937" box="[638,686,1723,1747]" pageId="6" pageNumber="479">type</typeStatus>
locality, in primary forest in Khao Phra Thaeo Non-hunting Area, and in primary and mature secondary forest in Phang-Nga Wildlife Breeding Station. At the
<typeStatus id="C64E88019517FF9EDE96F9308C8CF91C" box="[1217,1265,1760,1784]" pageId="6" pageNumber="479">type</typeStatus>
locality it was observed by us (AN, KK, MS, OSGP, PS) in syntopy with 23 species of squamates, namely
<taxonomicName id="DEF54D209517FF9EDECFF8D3893DF8A4" authority="Das &amp; Leong" authorityName="Das &amp; Leong" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="phuketensis">
<emphasis id="2B81EAB19517FF9EDECFF8D38DE1F8FF" box="[1176,1436,1794,1819]" italics="true" pageId="6" pageNumber="479">Cnemaspis phuketensis</emphasis>
Das &amp; Leong
</taxonomicName>
and
<taxonomicName id="DEF54D209517FF9EDBDEF8F78DE5F8DB" authority="Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya, 2010" authorityName="Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya" authorityYear="2010" box="[393,1432,1830,1856]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="vandeventeri">
<emphasis id="2B81EAB19517FF9EDBDEF8F78A39F8DB" box="[393,580,1830,1855]" italics="true" pageId="6" pageNumber="479">C. vandeventeri</emphasis>
<bibRefCitation id="7D644B529517FF9ED803F8F68DE5F8DB" author="Grismer" box="[596,1432,1830,1856]" pageId="6" pageNumber="479" refString="Grismer, L. L., Sumontha, M., Cota, M., Grismer, J. L., Wood, P. L. Jr., Pauwels, O. S. G. &amp; Kunya, K. (2010) A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa, 2576, 1 - 55." type="journal article" year="2010">Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDAC0F89A8A47F887" authority="Smith" authorityName="Smith" box="[151,570,1866,1891]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="2B81EAB19517FF9EDAC0F89A89A7F887" box="[151,474,1866,1891]" italics="true" pageId="6" pageNumber="479">Cyrtodactylus brevipalmatus</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED811F89B8B37F887" authority="Theobald" authorityName="Theobald" box="[582,842,1866,1891]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="oldhami">
<emphasis id="2B81EAB19517FF9ED811F89B8ABEF887" box="[582,707,1866,1891]" italics="true" pageId="6" pageNumber="479">C. oldhami</emphasis>
(Theobald)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED902F89B8ACEF863" authority="Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer, 2012" authorityName="Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="phuketensis">
<emphasis id="2B81EAB19517FF9ED902F89B8B84F887" box="[853,1017,1866,1891]" italics="true" pageId="6" pageNumber="479">C. phuketensis</emphasis>
<bibRefCitation id="7D644B529517FF9EDE56F89A8A73F863" author="Sumontha" pageId="6" pageNumber="479" refString="Sumontha, M., Pauwels, O. S. G., Kunya, K., Nitikul, A., Samphanthamit, P. &amp; Grismer, L. L. (2012) A new forest-dwelling gecko from Phuket Island, Southern Thailand, related to Cyrtodactylus macrotuberculatus (Squamata: Gekkonidae). Zootaxa, 3522, 61 - 72." type="journal article" year="2012">Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer, 2012</bibRefCitation>
(Gekkonidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED8E9F8BE8C66F863" authority="Kuhl" authorityName="Kuhl" box="[702,1051,1902,1927]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="2B81EAB19517FF9ED8E9F8BE8BBBF863" box="[702,966,1902,1927]" italics="true" pageId="6" pageNumber="479">Bronchocela cristatella</emphasis>
(Kuhl)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDE70F8BE8DE5F863" authority="Gray" authorityName="Gray" box="[1063,1432,1902,1927]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="emma">
<emphasis id="2B81EAB19517FF9EDE70F8BE8CB5F863" box="[1063,1224,1902,1927]" italics="true" pageId="6" pageNumber="479">Calotes emma</emphasis>
Gray (Agamidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDAC0F842899AF84F" authority="Fischer" authorityName="Fischer" box="[151,487,1938,1963]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="fasciolata">
<emphasis id="2B81EAB19517FF9EDAC0F8428906F84F" box="[151,379,1938,1963]" italics="true" pageId="6" pageNumber="479">Ahaetulla fasciolata</emphasis>
(Fischer)
</taxonomicName>
and
<taxonomicName id="DEF54D209517FF9ED848F8438A9AF84F" authority="Boie" authorityName="Boie" box="[543,743,1938,1963]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="prasina">
<emphasis id="2B81EAB19517FF9ED848F8438AE9F84F" box="[543,660,1939,1963]" italics="true" pageId="6" pageNumber="479">A. prasina</emphasis>
(Boie)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED8A4F8448C99F84F" authority="Dumeril, Bibron &amp; Dumeril" authorityName="Dumeril, Bibron &amp; Dumeril" box="[755,1252,1938,1963]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="cyanea">
<emphasis id="2B81EAB19517FF9ED8A4F8448BF0F84F" box="[755,909,1940,1963]" italics="true" pageId="6" pageNumber="479">Boiga cyanea</emphasis>
(Duméril, Bibron &amp; Duméril)
</taxonomicName>
and
<taxonomicName id="DEF54D209517FF9EDF4AF844889CF82B" authority="Boie" authorityName="Boie" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="cynodon">
<emphasis id="2B81EAB19517FF9EDF4AF8448DE1F84F" box="[1309,1436,1938,1963]" italics="true" pageId="6" pageNumber="479">B. cynodon</emphasis>
(Boie)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDABAF8668AFDF82B" authority="Gray" authorityName="Gray" box="[237,640,1974,1999]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="caudolineatus">
<emphasis id="2B81EAB19517FF9EDABAF8668A57F82B" box="[237,554,1974,1999]" italics="true" pageId="6" pageNumber="479">Dendrelaphis caudolineatus</emphasis>
(Gray)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED8DBF8688BEEF82B" authority="Wall" authorityName="Wall" box="[652,915,1974,1999]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="cyanochloris">
<emphasis id="2B81EAB19517FF9ED8DBF8688B3CF82B" box="[652,833,1974,1999]" italics="true" pageId="6" pageNumber="479">D. cyanochloris</emphasis>
(Wall)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9ED9C8F8688C01F82B" authority="Boie" authorityName="Boie" box="[927,1148,1974,1999]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="formosus">
<emphasis id="2B81EAB19517FF9ED9C8F8688C57F82B" box="[927,1066,1975,1999]" italics="true" pageId="6" pageNumber="479">D. formosus</emphasis>
(Boie)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDEDEF86888CFF817" authority="Van Rooijen &amp; Vo" authorityName="Van Rooijen &amp; Vo" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="haasi">
<emphasis id="2B81EAB19517FF9EDEDEF8688C94F82B" box="[1161,1257,1974,1999]" italics="true" pageId="6" pageNumber="479">D. haasi</emphasis>
Van Rooijen &amp; Vo
</taxonomicName>
g el a n d
<taxonomicName id="DEF54D209517FF9EDB40F80C8993F810" authority="Cohn" authorityName="Cohn" box="[279,494,2011,2036]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="striatus">
<emphasis id="2B81EAB19517FF9EDB40F80C89ECF817" box="[279,401,2012,2035]" italics="true" pageId="6" pageNumber="479">D. striatus</emphasis>
(Cohn)
</taxonomicName>
,
<taxonomicName id="DEF54D209517FF9EDBA9F80C8B5CF810" authority="Gunther" authorityName="Gunther" box="[510,801,2011,2036]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="carinata">
<emphasis id="2B81EAB19517FF9EDBA9F80C8ADAF817" box="[510,679,2012,2035]" italics="true" pageId="6" pageNumber="479">Ptyas carinata</emphasis>
(Günther)
</taxonomicName>
and
<emphasis id="2B81EAB19517FF9ED908F80C8BCAF817" box="[863,951,2011,2035]" italics="true" pageId="6" pageNumber="479">P. f u s c a</emphasis>
(Günther) (
<taxonomicName id="DEF54D209517FF9EDE10F80B8CB7F810" box="[1095,1226,2011,2036]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Colubridae</taxonomicName>
),
<taxonomicName id="DEF54D209517FF9FDEB6F80B889CFF54" authority="Boie" authorityName="Boie" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" lastPageId="7" lastPageNumber="480" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="boa">
<emphasis id="2B81EAB19517FF9EDEB6F80B8DE1F810" box="[1249,1436,2011,2036]" italics="true" pageId="6" pageNumber="479">Aplopeltura boa</emphasis>
(Boie)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDAA7FF498AE4FF54" authority="Boie" authorityName="Boie" box="[240,665,151,176]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="2B81EAB19516FF9FDAA7FF4989CFFF54" box="[240,434,153,176]" italics="true" pageId="7" pageNumber="480">Pareas carinatus</emphasis>
(Boie) (Pareatidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD8F0FF478C11FF54" authority="Stull" authorityName="Stull" box="[679,1132,151,176]" class="Reptilia" family="Pythonidae" genus="Python" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="brongersmai">
<emphasis id="2B81EAB19516FF9FD8F0FF478BF2FF54" box="[679,911,151,176]" italics="true" pageId="7" pageNumber="480">Python brongersmai</emphasis>
Stull (Pythonidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDE2CFF498B7BFF30" authority="Sumontha, Kunya, Pauwels, Nitikul &amp; Punnadee, 2011" authorityName="Sumontha, Kunya, Pauwels, Nitikul &amp; Punnadee" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="phuketensis">
<emphasis id="2B81EAB19516FF9FDE2CFF498DE1FF54" box="[1147,1436,151,176]" italics="true" pageId="7" pageNumber="480">Trimeresurus phuketensis</emphasis>
<bibRefCitation id="7D644B529516FF9FDAC0FF6C8B7BFF30" author="Sumontha" box="[151,774,188,213]" pageId="7" pageNumber="480" refString="Sumontha, M., Kunya, K., Pauwels, O. S. G., Nitikul, A. &amp; Punnadee, S. (2011) Trimeresurus (Popeia) phuketensis, a new pitviper (Squamata: Viperidae) from Phuket Island, southwestern Thailand. Russian Journal of Herpetology, 18 (3), 185 - 194." type="journal article" year="2011">Sumontha, Kunya, Pauwels, Nitikul &amp; Punnadee, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD945FF6C8C85FF31" authority="Wagler" authorityName="Wagler" box="[786,1272,188,213]" class="Reptilia" family="Viperidae" genus="Tropidolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="2B81EAB19516FF9FD945FF6C8C6BFF31" box="[786,1046,188,213]" italics="true" pageId="7" pageNumber="480">Tropidolaemus wagleri</emphasis>
Wagler (Viperidae)
</taxonomicName>
, and with four species of anuran amphibians,
<taxonomicName id="DEF54D209516FF9FD85AFF318B3DFF1C" authority="Smith" authorityName="Smith" box="[525,832,223,248]" class="Amphibia" family="Dicroglossidae" genus="Ingerana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="tasanae">
<emphasis id="2B81EAB19516FF9FD85AFF318AA4FF1C" box="[525,729,225,248]" italics="true" pageId="7" pageNumber="480">Ingerana tasanae</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD904FF318DE5FF1C" authority="Boulenger" authorityName="Boulenger" box="[851,1432,223,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="blythii">
<emphasis id="2B81EAB19516FF9FD904FF318C4BFF1C" box="[851,1078,223,248]" italics="true" pageId="7" pageNumber="480">Limnonectes blythii</emphasis>
(Boulenger) (Dicroglossidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDAC0FED48B6AFEF9" authority="Inger, Stuart &amp; Iskandar" authorityName="Inger, Stuart &amp; Iskandar" box="[151,791,260,285]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="2B81EAB19516FF9FDAC0FED48914FEF9" box="[151,361,260,285]" italics="true" pageId="7" pageNumber="480">Hylarana eschatia</emphasis>
(Inger, Stuart &amp; Iskandar) (Ranidae)
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FD904FED48D3FFEF9" authority="Peters" authorityName="Peters" box="[851,1346,260,285]" class="Amphibia" family="Rhacophoridae" genus="Nyctixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="pictus">
<emphasis id="2B81EAB19516FF9FD904FED48C6AFEF8" box="[851,1047,260,285]" italics="true" pageId="7" pageNumber="480">Nyctixalus pictus</emphasis>
(Peters) (Rhacophoridae)
</taxonomicName>
. In Suk Samran District in Ranong Province it has been observed in secondary forest at the proximity of
<taxonomicName id="DEF54D209516FF9FDF59FEF78974FE81" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="2B81EAB19516FF9FDF59FEF78974FE81" italics="true" pageId="7" pageNumber="480">Bronchocela cristatella</emphasis>
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDB77FE9C8A1EFE81" box="[288,611,332,357]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="emma">
<emphasis id="2B81EAB19516FF9FDB77FE9C89B4FE80" box="[288,457,332,357]" italics="true" pageId="7" pageNumber="480">Calotes emma</emphasis>
(Agamidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD82FFE9C8BB9FE81" box="[632,964,332,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="2B81EAB19516FF9FD82FFE9C8BB9FE81" box="[632,964,332,357]" italics="true" pageId="7" pageNumber="480">Cyrtodactylus brevipalmatus</emphasis>
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FDE58FE9D8CBFFE80" box="[1039,1218,333,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="ranongensis">
<emphasis id="2B81EAB19516FF9FDE58FE9D8CBFFE80" box="[1039,1218,333,356]" italics="true" pageId="7" pageNumber="480">C. ranongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDE8DFE9C895AFE6C" authority="Wiegmann" authorityName="Wiegmann" class="Reptilia" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="mutilata">
<emphasis id="2B81EAB19516FF9FDE8DFE9C8DE1FE81" box="[1242,1436,332,357]" italics="true" pageId="7" pageNumber="480">Gehyra mutilata</emphasis>
(Wiegmann)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDB63FEBF8A30FE6C" authority="Linnaeus" authorityName="Linnaeus" box="[308,589,367,392]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="gecko">
<emphasis id="2B81EAB19516FF9FDB63FEBF89BBFE6C" box="[308,454,367,392]" italics="true" pageId="7" pageNumber="480">Gekko gecko</emphasis>
(Linnaeus)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD80CFEBF8BA4FE6C" authority="Schlegel" authorityName="Schlegel" box="[603,985,367,392]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="2B81EAB19516FF9FD80CFEBF8B24FE6C" box="[603,857,367,392]" italics="true" pageId="7" pageNumber="480">Hemidactylus frenatus</emphasis>
(Schlegel)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD9B0FEA18D38FE6C" authority="Dumeril &amp; Bibron" authorityName="Dumeril &amp; Bibron" box="[999,1349,367,392]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="garnotii">
<emphasis id="2B81EAB19516FF9FD9B0FEA18C1BFE6C" box="[999,1126,369,392]" italics="true" pageId="7" pageNumber="480">H. garnotii</emphasis>
Duméril &amp; Bibron
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FDFD6FEA189EDFE49" authority="Schneider" authorityName="Schneider" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="platyurus">
<emphasis id="2B81EAB19516FF9FDFD6FEA1897CFE49" italics="true" pageId="7" pageNumber="480">H. platyurus</emphasis>
(Schneider)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDBC9FE448BC4FE49" authority="Annandale" authorityName="Annandale" box="[414,953,404,429]" class="Reptilia" family="Gekkonidae" genus="Ptychozoon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="lionotum">
<emphasis id="2B81EAB19516FF9FDBC9FE448AF2FE49" box="[414,655,404,429]" italics="true" pageId="7" pageNumber="480">Ptychozoon lionotum</emphasis>
Annandale (Gekkonidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD990FE448D3AFE49" authority="Linnaeus" authorityName="Linnaeus" box="[967,1351,404,429]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="mycterizans">
<emphasis id="2B81EAB19516FF9FD990FE448CB9FE48" box="[967,1220,404,429]" italics="true" pageId="7" pageNumber="480">Ahaetulla mycterizans</emphasis>
(Linnaeus)
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FDFD3FE448891FE34" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="prasina">
<emphasis id="2B81EAB19516FF9FDFD3FE448891FE34" italics="true" pageId="7" pageNumber="480">A. prasina</emphasis>
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDB54FE698A48FE34" authority="Boie" authorityName="Boie" box="[259,565,439,464]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="dendrophila">
<emphasis id="2B81EAB19516FF9FDB54FE6989A0FE34" box="[259,477,439,464]" italics="true" pageId="7" pageNumber="480">Boiga dendrophila</emphasis>
(Boie)
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FD82BFE698C7CFE34" authority="Boie" authorityName="Boie" box="[636,1025,439,464]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="drapiezii">
<emphasis id="2B81EAB19516FF9FD82BFE698B75FE34" box="[636,776,439,464]" italics="true" pageId="7" pageNumber="480">B. drapiezii</emphasis>
(Boie) (Colubridae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDE43FE678DEAFE34" authority="Blyth" authorityName="Blyth" box="[1044,1431,439,464]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="nigrocinctus">
<emphasis id="2B81EAB19516FF9FDE43FE678D49FE34" box="[1044,1332,439,464]" italics="true" pageId="7" pageNumber="480">Rhabdophis nigrocinctus</emphasis>
(Blyth)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FDAC0FE0C8AB7FE11" authority="Boie" authorityName="Boie" box="[151,714,476,501]" class="Reptilia" family="Colubridae" genus="Xenochrophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="trianguligerus">
<emphasis id="2B81EAB19516FF9FDAC0FE0C89A3FE11" box="[151,478,476,501]" italics="true" pageId="7" pageNumber="480">Xenochrophis trianguligerus</emphasis>
(Boie) (Natricidae)
</taxonomicName>
,
<taxonomicName id="DEF54D209516FF9FD88CFE0D8C49FE11" box="[731,1076,476,501]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="2B81EAB19516FF9FD88CFE0D8BDDFE10" box="[731,928,477,500]" italics="true" pageId="7" pageNumber="480">Pareas carinatus</emphasis>
(Pareatidae)
</taxonomicName>
and
<taxonomicName id="DEF54D209516FF9FDE21FE0C8A4EFDFC" authority="(Schneider) (Sumontha et al., 2015)" authorityName="(Schneider) (Sumontha et al." authorityYear="2015" class="Reptilia" family="Pythonidae" genus="Malayopython" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="2B81EAB19516FF9FDE21FE0C8DE1FE11" box="[1142,1436,476,501]" italics="true" pageId="7" pageNumber="480">Malayopython reticulatus</emphasis>
(Schneider) (
<bibRefCitation id="7D644B529516FF9FDB7EFE2F8A56FDFC" author="Sumontha" box="[297,555,511,536]" pageId="7" pageNumber="480" refString="Sumontha, M., Pauwels, O. S. G., Panitvong, N., Kunya, K. &amp; Grismer, L. L. (2015) A new lowland forest Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Ranong Province, peninsular Thailand. Zootaxa, 3911 (1), 106 - 118. http: // dx. doi. org / 10.11646 / zootaxa. 3911.1.6" type="journal article" year="2015">
Sumontha
<emphasis id="2B81EAB19516FF9FDBF2FDD189AAFDFC" box="[421,471,511,536]" italics="true" pageId="7" pageNumber="480">et al</emphasis>
., 2015
</bibRefCitation>
)
</taxonomicName>
. The species seems to be strictly sylvicolous, and was never found in highly degraded secondary forest nor in plantations.
</paragraph>
<caption id="4D8A662B9517FF9EDAC0F9F18954F9B2" httpUri="https://zenodo.org/record/399935/files/figure.png" pageId="6" pageNumber="479" targetBox="[151,1436,202,1528]" targetPageId="6">
<paragraph id="194A36A39517FF9EDAC0F9F18954F9B2" blockId="6.[151,1436,1569,1683]" pageId="6" pageNumber="479">
<emphasis id="2B81EAB19517FF9EDAC0F9F18962F9D3" bold="true" box="[151,287,1569,1591]" pageId="6" pageNumber="479">FIGURE 4a.</emphasis>
Dorsal view of preserved
<taxonomicName id="DEF54D209517FF9ED87CF9F18B48F9D3" box="[555,821,1569,1591]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19517FF9ED87CF9F18B48F9D3" box="[555,821,1569,1591]" italics="true" pageId="6" pageNumber="479">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9517FF9ED96CF9F18BF6F9D3" box="[827,907,1569,1591]" pageId="6" pageNumber="479" rank="species">
<emphasis id="2B81EAB19517FF9ED96CF9F18BF6F9D3" bold="true" box="[827,907,1569,1591]" pageId="6" pageNumber="479">sp. nov.</emphasis>
</taxonomicNameLabel>
adult female paratype THNHM 22543. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9517FF9EDAC0F98E8954F977" pageId="6" pageNumber="479">
<paragraph id="194A36A39517FF9EDAC0F98E8954F977" blockId="6.[151,1436,1569,1683]" pageId="6" pageNumber="479">
<emphasis id="2B81EAB19517FF9EDAC0F98E895DF990" bold="true" box="[151,288,1630,1652]" pageId="6" pageNumber="479">FIGURE 4b.</emphasis>
Ventral view of preserved
<taxonomicName id="DEF54D209517FF9ED878F98E8B44F990" box="[559,825,1630,1652]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19517FF9ED878F98E8B44F990" box="[559,825,1630,1652]" italics="true" pageId="6" pageNumber="479">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9517FF9ED968F98E8BF2F990" box="[831,911,1630,1652]" pageId="6" pageNumber="479" rank="species">
<emphasis id="2B81EAB19517FF9ED968F98E8BF2F990" bold="true" box="[831,911,1630,1652]" pageId="6" pageNumber="479">sp. nov.</emphasis>
</taxonomicNameLabel>
adult female paratype THNHM 22543. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<paragraph id="194A36A39516FF9FDA90FD97896BFD29" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="480">
<figureCitation id="81CE2A269516FF9FDA90FD978955FD84" box="[199,296,583,608]" captionStart="FIGURE 9" captionStartId="13.[151,250,1965,1987]" captionTargetBox="[201,1372,193,1937]" captionTargetId="figure@13.[201,1372,193,1938]" captionTargetPageId="13" captionText="FIGURE 9. Live subadult male Acanthosaura phuketensis sp. nov., eating a grasshopper on a tree in situ at Ban Bangrong, Phuket Island, southwestern Thailand. Photo by P. Samphanthamit." httpUri="https://zenodo.org/record/399940/files/figure.png" pageId="7" pageNumber="480">Figure 9</figureCitation>
shows a subadult male
<taxonomicName id="DEF54D209516FF9FD815FD978B11FD84" box="[578,876,583,608]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19516FF9FD815FD978B11FD84" box="[578,876,583,608]" italics="true" pageId="7" pageNumber="480">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9516FF9FD921FD988BAEFD84" box="[886,979,584,608]" pageId="7" pageNumber="480" rank="species">
<emphasis id="2B81EAB19516FF9FD921FD988BAEFD84" bold="true" box="[886,979,584,608]" pageId="7" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
eating a large leaf-colored grasshopper (Orthoptera). In captivity,
<taxonomicName id="DEF54D209516FF9FDB84FDBC8A01FD61" box="[467,636,620,645]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19516FF9FDB84FDBC8A01FD61" box="[467,636,620,645]" italics="true" pageId="7" pageNumber="480">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9516FF9FD8DDFDBC8A96FD60" box="[650,747,620,644]" pageId="7" pageNumber="480" rank="species">
<emphasis id="2B81EAB19516FF9FD8DDFDBC8A96FD60" bold="true" box="[650,747,620,644]" pageId="7" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
accepts meal worms, crickets, woodlice and earthworms (
<bibRefCitation id="7D644B529516FF9FDAC9FD5F89F5FD4C" author="Pauwels" box="[158,392,655,680]" pageId="7" pageNumber="480" refString="Pauwels, O. S. G., Laohawat, O. - A., David, P., Bour, R., Dangsee, P., Puangjit, C. &amp; Chimsunchart, C. (2000) Herpetological investigations in Phang-Nga Province, southern peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilia, 4 (2), 123 - 154." type="journal article" year="2000">
Pauwels
<emphasis id="2B81EAB19516FF9FDB51FD418945FD4C" box="[262,312,655,680]" italics="true" pageId="7" pageNumber="480">et al</emphasis>
., 2000
</bibRefCitation>
and unpubl. obs.). The abdominal cavity of the
<typeStatus id="C64E88019516FF9FD9E4FD408C6EFD4C" box="[947,1043,656,680]" pageId="7" pageNumber="480" type="paratype">paratype</typeStatus>
IRSNB 2681 contained numerous nematodes.
</paragraph>
</subSubSection>
<caption id="4D8A662B9516FF9FDAC0F88D8973F876" pageId="7" pageNumber="480">
<paragraph id="194A36A39516FF9FDAC0F88D8973F876" blockId="7.[151,1436,1885,1999]" pageId="7" pageNumber="480">
<emphasis id="2B81EAB19516FF9FDAC0F88D8962F897" bold="true" box="[151,287,1885,1907]" pageId="7" pageNumber="480">FIGURE 5a.</emphasis>
Dorsal view of preserved
<taxonomicName id="DEF54D209516FF9FD87CF88D8B48F897" box="[555,821,1885,1907]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19516FF9FD87CF88D8B48F897" box="[555,821,1885,1907]" italics="true" pageId="7" pageNumber="480">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9516FF9FD96BF88D8BF1F897" box="[828,908,1885,1907]" pageId="7" pageNumber="480" rank="species">
<emphasis id="2B81EAB19516FF9FD96BF88D8BF1F897" bold="true" box="[828,908,1885,1907]" pageId="7" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
subadult male paratype QSMI 1166. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9516FF9FDAC0F84A8973F82B" httpUri="https://zenodo.org/record/399936/files/figure.png" pageId="7" pageNumber="480" targetBox="[151,1436,775,1843]" targetPageId="7">
<paragraph id="194A36A39516FF9FDAC0F84A8973F82B" blockId="7.[151,1436,1885,1999]" pageId="7" pageNumber="480">
<emphasis id="2B81EAB19516FF9FDAC0F84A895DF854" bold="true" box="[151,288,1946,1968]" pageId="7" pageNumber="480">FIGURE 5b.</emphasis>
Ventral view of preserved
<taxonomicName id="DEF54D209516FF9FD878F84A8B47F854" box="[559,826,1946,1968]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19516FF9FD878F84A8B47F854" box="[559,826,1946,1968]" italics="true" pageId="7" pageNumber="480">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9516FF9FD968F84A8BEDF854" box="[831,912,1946,1968]" pageId="7" pageNumber="480" rank="species">
<emphasis id="2B81EAB19516FF9FD968F84A8BEDF854" bold="true" box="[831,912,1946,1968]" pageId="7" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
subadult male paratype QSMI 1166. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9519FF90DAC0FA9F8915FA67" pageId="8" pageNumber="481">
<paragraph id="194A36A39519FF90DAC0FA9F8915FA67" blockId="8.[151,1436,1359,1473]" pageId="8" pageNumber="481">
<emphasis id="2B81EAB19519FF90DAC0FA9F895DFA81" bold="true" box="[151,288,1359,1381]" pageId="8" pageNumber="481">FIGURE 6a.</emphasis>
Dorsal view of preserved
<taxonomicName id="DEF54D209519FF90D866FA9F8B41FA81" box="[561,828,1359,1381]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19519FF90D866FA9F8B41FA81" box="[561,828,1359,1381]" italics="true" pageId="8" pageNumber="481">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9519FF90D914FA9F8BE8FA81" box="[835,917,1359,1381]" pageId="8" pageNumber="481" rank="species">
<emphasis id="2B81EAB19519FF90D914FA9F8BE8FA81" bold="true" box="[835,917,1359,1381]" pageId="8" pageNumber="481">sp. nov.</emphasis>
</taxonomicNameLabel>
subadult male paratype PSUZC-RT 2012.6. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<caption id="4D8A662B9519FF90DAC0FA5C8915FA25" httpUri="https://zenodo.org/record/399937/files/figure.png" pageId="8" pageNumber="481" targetBox="[151,1436,198,1320]" targetPageId="8">
<paragraph id="194A36A39519FF90DAC0FA5C8915FA25" blockId="8.[151,1436,1359,1473]" pageId="8" pageNumber="481">
<emphasis id="2B81EAB19519FF90DAC0FA5C895CFA46" bold="true" box="[151,289,1420,1442]" pageId="8" pageNumber="481">FIGURE 6b.</emphasis>
Ventral view of preserved
<taxonomicName id="DEF54D209519FF90D862FA5C8B3DFA46" box="[565,832,1420,1442]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19519FF90D862FA5C8B3DFA46" box="[565,832,1420,1442]" italics="true" pageId="8" pageNumber="481">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9519FF90D911FA5C8BE5FA46" box="[838,920,1420,1442]" pageId="8" pageNumber="481" rank="species">
<emphasis id="2B81EAB19519FF90D911FA5C8BE5FA46" bold="true" box="[838,920,1420,1442]" pageId="8" pageNumber="481">sp. nov.</emphasis>
</taxonomicNameLabel>
subadult male paratype PSUZC-RT 2012.6. Photo by O. S. G. Pauwels.
</paragraph>
</caption>
<subSubSection id="51EF65289519FF90DA90FA38888EF989" pageId="8" pageNumber="481" type="etymology">
<paragraph id="194A36A39519FF90DA90FA38888EF989" blockId="8.[151,1436,1512,2005]" pageId="8" pageNumber="481">
<emphasis id="2B81EAB19519FF90DA90FA388933F9E4" bold="true" box="[199,334,1512,1536]" pageId="8" pageNumber="481">Etymology.</emphasis>
The specific epithet is an adjective in reference to Phuket Island and Phuket Range, on which the
<typeStatus id="C64E88019519FF90DAC0F9DD88BBF9C1" box="[151,198,1549,1573]" pageId="8" pageNumber="481">type</typeStatus>
locality lies. We suggest the following common names:
<emphasis id="2B81EAB19519FF90D96DF9DC8C1FF9C1" box="[826,1122,1548,1573]" italics="true" pageId="8" pageNumber="481">Kingkakhaownaam Phuket</emphasis>
(
<collectingCountry id="61E276339519FF90DE25F9DC8CD4F9C1" box="[1138,1193,1548,1573]" name="Thailand" pageId="8" pageNumber="481">Thai</collectingCountry>
), Phuket Horned Tree Agamid (English),
<emphasis id="2B81EAB19519FF90DB2EF9E08AEEF9AD" box="[377,659,1584,1609]" italics="true" pageId="8" pageNumber="481">Acanthosaure de Phuket</emphasis>
(French),
<emphasis id="2B81EAB19519FF90D942F9E08C69F9AD" box="[789,1044,1584,1609]" italics="true" pageId="8" pageNumber="481">Phuket-Nackenstachler</emphasis>
(
<collectingCountry id="61E276339519FF90DE7DF9E08CF5F9AC" box="[1066,1160,1584,1608]" name="Germany" pageId="8" pageNumber="481">German</collectingCountry>
),
<emphasis id="2B81EAB19519FF90DEF4F9E08DE1F9AD" box="[1187,1436,1584,1609]" italics="true" pageId="8" pageNumber="481">Phuketstekelnekagame</emphasis>
(Dutch).
</paragraph>
</subSubSection>
<subSubSection id="51EF65289519FF93DA90F9A78DEAF801" lastPageId="11" lastPageNumber="484" pageId="8" pageNumber="481" type="materials_examined">
<paragraph id="194A36A39519FF93DA90F9A78AC6FE83" blockId="8.[151,1436,1512,2005]" lastBlockId="11.[151,1437,151,359]" lastPageId="11" lastPageNumber="484" pageId="8" pageNumber="481">
<emphasis id="2B81EAB19519FF90DA90F9A78A40F974" bold="true" box="[199,573,1655,1680]" pageId="8" pageNumber="481">Comparisons to other species.</emphasis>
<tableCitation id="547703189519FF90D81DF9A88ADFF974" box="[586,674,1656,1681]" captionStart="TABLE 2" captionStartId="18.[119,197,528,548]" captionText="TABLE 2. Comparison of morphometrical (in mm) and meristic data for all currently recognized species of Acanthosaura and A. phuketensis sp. nov. For the ? see the comparison section." pageId="8" pageNumber="481">Table 2</tableCitation>
shows a comparison of morphometric and meristic data for all currently recognized species of
<taxonomicName id="DEF54D209519FF90D854F94C8ADDF951" box="[515,672,1692,1717]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="genus">
<emphasis id="2B81EAB19519FF90D854F94C8ADDF951" box="[515,672,1692,1717]" italics="true" pageId="8" pageNumber="481">Acanthosaura</emphasis>
</taxonomicName>
and
<taxonomicName id="DEF54D209519FF90D88CF94D8B02F951" box="[731,895,1692,1717]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19519FF90D88CF94D8B02F951" box="[731,895,1692,1717]" italics="true" pageId="8" pageNumber="481">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9519FF90D9DFF94C8B99F950" box="[904,996,1692,1716]" pageId="8" pageNumber="481" rank="species">
<emphasis id="2B81EAB19519FF90D9DFF94C8B99F950" bold="true" box="[904,996,1692,1716]" pageId="8" pageNumber="481">sp. nov.</emphasis>
</taxonomicNameLabel>
It is mostly based on the interspecific comparison table provided by Wood
<emphasis id="2B81EAB19519FF90D86BF9118A0BF93D" box="[572,630,1728,1753]" italics="true" pageId="8" pageNumber="481">et al.</emphasis>
(2010, Table 5) and additional data from
<bibRefCitation id="7D644B529519FF90DE02F9108A3DF919" author="Hallermann" pageId="8" pageNumber="481" refString="Hallermann, J. (2000) The taxonomic status of Acanthosaura fruhstorferi Werner, 1904 and Calotes brevipes Werner, 1904 (Squamata, Agamidae). Mitteilungen des Museum fur Naturkunde Berlin, Zoologische Reihe, 76 (1), 143 - 150. http: // dx. doi. org / 10.1002 / mmnz. 20000760113" type="journal article" year="2000">
Hallermann (2000; only data for
<typeStatus id="C64E88019519FF90DA96F9348955F919" box="[193,296,1764,1789]" pageId="8" pageNumber="481" type="lectotype">lectotype</typeStatus>
<taxonomicName id="DEF54D209519FF90DB67F9358A3DF919" authority="Werner" authorityName="Werner" box="[304,576,1764,1789]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="fruhstorferi">
<emphasis id="2B81EAB19519FF90DB67F93589AEF919" box="[304,467,1764,1789]" italics="true" pageId="8" pageNumber="481">A. fruhstorferi</emphasis>
(Werner)
</taxonomicName>
</bibRefCitation>
included),
<bibRefCitation id="7D644B529519FF90D893F9348BB8F919" box="[708,965,1764,1789]" pageId="8" pageNumber="481" refString="Ananjeva, N. B., Orlov, N. L., Nguyen, T. T. &amp; Ryabov, S. A. (2011) A new species of Acanthosaura (Agamidae, Sauria) from northwest Vietnam. Russian Journal of Herpetology, 18 (3), 195 - 202." type="journal article">
Ananjeva
<emphasis id="2B81EAB19519FF90D96FF9358B0EF919" box="[824,883,1764,1789]" italics="true" pageId="8" pageNumber="481">et al.</emphasis>
(2011)
</bibRefCitation>
and
<bibRefCitation id="7D644B529519FF90DE57F9348CD6F919" author="Bobrov" box="[1024,1195,1764,1789]" pageId="8" pageNumber="481" refString="Bobrov, V. V. (2013) Acanthosaura cardamomensis (Reptilia, Sauria, Agamidae) as a lizard species new for the Vietnam fauna. Modern Herpetology, 13 (1 / 2), 64 - 66." type="book chapter" year="2013">Bobrov (2013)</bibRefCitation>
. However, it is to be noted that Wood
<emphasis id="2B81EAB19519FF90DB0CF8D989E9F8C5" box="[347,404,1800,1825]" italics="true" pageId="8" pageNumber="481">et al.</emphasis>
(2010) wrongly defined the character ORBIT used in their comparison table, as the eye diameter, measured from the posterior to the anterior edge of the eye, while they actually meant and measured the orbit diameter, not the eye diameter. About the same table presented by Wood
<emphasis id="2B81EAB19519FF90DE50F8818C42F88D" box="[1031,1087,1872,1897]" italics="true" pageId="8" pageNumber="481">et al.</emphasis>
(2010), it should also be noted that there was a
<emphasis id="2B81EAB19519FF90DB19F8A48995F869" box="[334,488,1908,1933]" italics="true" pageId="8" pageNumber="481">lapsus calami</emphasis>
regarding the ratio TaL/SVL for
<taxonomicName id="DEF54D209519FF90D908F8A58B9FF868" box="[863,994,1909,1932]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="coronata">
<emphasis id="2B81EAB19519FF90D908F8A58B9FF868" box="[863,994,1909,1932]" italics="true" pageId="8" pageNumber="481">A. coronata</emphasis>
</taxonomicName>
, erroneously indicated as varying from 0.061.01, while the corrected variation has been used in our
<tableCitation id="547703189519FF90D92FF8488BADF854" box="[888,976,1944,1969]" captionStart="TABLE 2" captionStartId="18.[119,197,528,548]" captionText="TABLE 2. Comparison of morphometrical (in mm) and meristic data for all currently recognized species of Acanthosaura and A. phuketensis sp. nov. For the ? see the comparison section." pageId="8" pageNumber="481">Table 2</tableCitation>
. A close re-examination of the material regarded by Wood
<emphasis id="2B81EAB19519FF90DB3BF86D89E0F831" box="[364,413,1980,2005]" italics="true" pageId="8" pageNumber="481">et al</emphasis>
. (2010) to represent
<taxonomicName id="DEF54D209519FF90D8D0F86D8B6EF830" box="[647,787,1981,2004]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19519FF90D8D0F86D8B6EF830" box="[647,787,1981,2004]" italics="true" pageId="8" pageNumber="481">A. crucigera</emphasis>
</taxonomicName>
led to separate populations from western
<collectingCountry id="61E276339519FF90DEBFF86C8D31F831" box="[1256,1356,1980,2005]" name="Thailand" pageId="8" pageNumber="481">Thailand</collectingCountry>
and at and near its
<typeStatus id="C64E88019518FF91DB6BFB6A8911FB36" box="[316,364,1210,1234]" pageId="9" pageNumber="482">type</typeStatus>
locality in
<collectingCountry id="61E276339518FF91D850FB6A8A09FB36" box="[519,628,1210,1234]" name="Myanmar" pageId="9" pageNumber="482">Myanmar</collectingCountry>
from those from the Kra Isthmus area based on coloration and morphometrical characteristics, as well as on molecular data (Wood
<emphasis id="2B81EAB19518FF91D9C0FB0F8BB5FB12" box="[919,968,1245,1270]" italics="true" pageId="9" pageNumber="482">et al</emphasis>
., in prep.). Only the northern populations are actually referrable to
<taxonomicName id="DEF54D209518FF91DBE4FAD28A3DFAFE" box="[435,576,1282,1306]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19518FF91DBE4FAD28A3DFAFE" box="[435,576,1282,1306]" italics="true" pageId="9" pageNumber="482">A. crucigera</emphasis>
</taxonomicName>
, showing, similarly to the
<typeStatus id="C64E88019518FF91D939FAD28BE3FAFE" box="[878,926,1282,1306]" pageId="9" pageNumber="482">type</typeStatus>
material, a relatively low nuchal crest, a low dorsal crest, and no contact between the black eye patch and the nuchal crest in both sexes, among other characteristics. The morphological variation for
<taxonomicName id="DEF54D209518FF91D893FA9A8B2EFA86" box="[708,851,1354,1378]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19518FF91D893FA9A8B2EFA86" box="[708,851,1354,1378]" italics="true" pageId="9" pageNumber="482">A. crucigera</emphasis>
</taxonomicName>
that was presented by Wood
<emphasis id="2B81EAB19518FF91DEF9FA9B8C94FA86" box="[1198,1257,1353,1378]" italics="true" pageId="9" pageNumber="482">et al.</emphasis>
(2010) in their interspecific comparison table has consequently been reviewed to include only what we regard here as true
<taxonomicName id="DEF54D209518FF91DFD3FABE8979FA4E" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB19518FF91DFD3FABE8979FA4E" italics="true" pageId="9" pageNumber="482">A. crucigera</emphasis>
</taxonomicName>
(see Appendix: Material examined).
<taxonomicName id="DEF54D209518FF91D8E3FA418BA2FA4E" box="[692,991,1425,1450]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91D8E3FA418BA2FA4E" box="[692,991,1425,1450]" italics="true" pageId="9" pageNumber="482">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91D9BFFA428C39FA4E" box="[1000,1092,1426,1450]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91D9BFFA428C39FA4E" bold="true" box="[1000,1092,1426,1450]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from
<taxonomicName id="DEF54D209518FF91DFD3FA428895FA2A" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="armata">
<emphasis id="2B81EAB19518FF91DFD3FA428895FA2A" italics="true" pageId="9" pageNumber="482">A. armata</emphasis>
</taxonomicName>
by its higher maximal length of the postorbital spines (11.8 vs.
<quantity id="DE0D9B469518FF91D9EFFA668C6EFA2A" box="[952,1043,1462,1486]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="9" pageNumber="482" unit="mm" value="9.9">9.9 mm</quantity>
), its much higher number of scales in the diastema between the nuchal and the dorsal crests (1217 vs. 18), a lower number of infralabials (1012 vs. 1215) and the presence of a black eye patch (vs. absence). It can be distinguished from
<taxonomicName id="DEF54D209518FF91DE2EFA2E8D58F9F2" box="[1145,1317,1533,1558]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="bintangensis">
<emphasis id="2B81EAB19518FF91DE2EFA2E8D58F9F2" box="[1145,1317,1533,1558]" italics="true" pageId="9" pageNumber="482">A. bintangensis</emphasis>
</taxonomicName>
by having a larger tympanum (TD/HD 0.220.33 vs. 0.16), much longer postorbital spines (PS 4.611.8 vs.
<quantity id="DE0D9B469518FF91DE88F9F28D15F9DD" box="[1247,1384,1570,1594]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="4.2" metricValueMin="1.9" pageId="9" pageNumber="482" unit="mm" value="3.05" valueMax="4.2" valueMin="1.9">1.94.2 mm</quantity>
, and PS/HL 0.230.38 vs. 0.070.19), longer spines in the nuchal crest (NSL/HL 0.210.39 vs. 0.170.21), longer spines in the dorsal crest (DS 2.38.3 vs.
<quantity id="DE0D9B469518FF91D822F9BA8A82F965" box="[629,767,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="9" pageNumber="482" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
, and DS/HL 0.110.26 vs. 0.080.09), more ventral scales (5767 vs. 5155), less subdigital lamellae on the fourth finger (1517 vs. 23), less subdigital lamellae on the fourth toe (2124 vs. 2628), much longer occipital spines (OS 2.69.5 vs.
<quantity id="DE0D9B469518FF91D9BFF9618C0CF92D" box="[1000,1137,1713,1738]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="9" pageNumber="482" unit="mm" value="1.9" valueMax="2.6" valueMin="1.2">1.22.6 mm</quantity>
, and OS/HL 0.130.30 vs. 0.100.11), less scales surrounding the occipital spine (45 vs. 67), more scales between the fifth canthals (1213 vs. 1011), presence of a light knee patch (vs. absence), less developed gular pouch (GP 02 vs. 34) and absence of an enlarged row of keeled scales below orbit (vs. presence). From
<taxonomicName id="DEF54D209518FF91D9C0F8CE8C41F8D2" box="[919,1084,1821,1846]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB19518FF91D9C0F8CE8C41F8D2" box="[919,1084,1821,1846]" italics="true" pageId="9" pageNumber="482">A. brachypoda</emphasis>
</taxonomicName>
, A.
<taxonomicName id="DEF54D209518FF91DE3CF8CD8C93F8D2" box="[1131,1262,1821,1846]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91DE3CF8CD8C93F8D2" box="[1131,1262,1821,1846]" italics="true" pageId="9" pageNumber="482">phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91DEA3F8CE8D33F8D2" box="[1268,1358,1822,1846]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91DEA3F8CE8D33F8D2" bold="true" box="[1268,1358,1822,1846]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated by a larger tympanum (TD/HD 0.220.33 vs. 0.21), much longer postorbital spines (PS/HL
<quantity id="DE0D9B469518FF91DF0AF8928894F89A" metricMagnitude="-3" metricUnit="m" metricValue="6.985" metricValueMax="7.112" metricValueMin="6.858" pageId="9" pageNumber="482" unit="in" value="0.275" valueMax="0.28" valueMin="0.27">0.27 0.28 in</quantity>
the two adult female
<typeStatus id="C64E88019518FF91DBA7F8B78A21F89B" box="[496,604,1895,1919]" pageId="9" pageNumber="482" type="paratype">paratypes</typeStatus>
of A.
<taxonomicName id="DEF54D209518FF91D8F8F8B58B4FF89A" box="[687,818,1893,1918]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91D8F8F8B58B4FF89A" box="[687,818,1893,1918]" italics="true" pageId="9" pageNumber="482">phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91D96AF8B68BE6F89A" box="[829,923,1894,1918]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91D96AF8B68BE6F89A" bold="true" box="[829,923,1894,1918]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
vs.
<quantity id="DE0D9B469518FF91D987F8B68C5FF89A" box="[976,1058,1893,1918]" metricMagnitude="-3" metricUnit="m" metricValue="2.7939999999999996" pageId="9" pageNumber="482" unit="in" value="0.11">0.11 in</quantity>
the adult female
<typeStatus id="C64E88019518FF91DEA3F8B58D2BF89A" box="[1268,1366,1893,1918]" pageId="9" pageNumber="482" type="holotype">holotype</typeStatus>
of
<taxonomicName id="DEF54D209518FF91DFD3F8B68960F846" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB19518FF91DFD3F8B68960F846" italics="true" pageId="9" pageNumber="482">A. brachypoda</emphasis>
</taxonomicName>
), longer spines in the nuchal crest (NSL/HL 0.23 and
<quantity id="DE0D9B469518FF91D9CDF85A8B96F846" box="[922,1003,1929,1954]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="482" unit="in" value="0.3">0.30 in</quantity>
the two adult female
<typeStatus id="C64E88019518FF91DEBAF85A8D24F846" box="[1261,1369,1930,1954]" pageId="9" pageNumber="482" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="DEF54D209518FF91DFD3F85A8967F822" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91DFD3F85A8967F822" italics="true" pageId="9" pageNumber="482">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91DB77F87E8904F822" box="[288,377,1966,1990]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91DB77F87E8904F822" bold="true" box="[288,377,1966,1990]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
vs.
<quantity id="DE0D9B469518FF91DBF1F87D8989F822" box="[422,500,1965,1990]" metricMagnitude="-3" metricUnit="m" metricValue="4.064" pageId="9" pageNumber="482" unit="in" value="0.16">0.16 in</quantity>
the adult female
<typeStatus id="C64E88019518FF91D8E2F87D8B6AF822" box="[693,791,1965,1990]" pageId="9" pageNumber="482" type="holotype">holotype</typeStatus>
of
<taxonomicName id="DEF54D209518FF91D96BF87E8B9DF822" box="[828,992,1965,1990]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB19518FF91D96BF87E8B9DF822" box="[828,992,1965,1990]" italics="true" pageId="9" pageNumber="482">A. brachypoda</emphasis>
</taxonomicName>
), much longer spines in the dorsal crest (DS/HL 0.11 and
<quantity id="DE0D9B469518FF91DB32F80289C9F80E" box="[357,436,2001,2026]" metricMagnitude="-3" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="482" unit="in" value="0.14">0.14 in</quantity>
the two adult female
<typeStatus id="C64E88019518FF91D8E5F8028B60F80E" box="[690,797,2002,2026]" pageId="9" pageNumber="482" type="paratype">paratypes</typeStatus>
of A.
<taxonomicName id="DEF54D209518FF91D93DF8018B90F80E" box="[874,1005,2001,2026]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91D93DF8018B90F80E" box="[874,1005,2001,2026]" italics="true" pageId="9" pageNumber="482">phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91D9A1F8028C2FF80E" box="[1014,1106,2002,2026]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91D9A1F8028C2FF80E" bold="true" box="[1014,1106,2002,2026]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
vs. 0.06), a distinctly higher number of scales in the diastema between the nuchal and the dorsal crests (1217 vs. 7), less subdigital lamellae on the fourth finger (1517 vs. 18), a much longer occipital spine (OS/HL 0.19 and
<quantity id="DE0D9B46951BFF92DE06FF6C8CDFFF31" box="[1105,1186,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" pageId="10" pageNumber="483" unit="in" value="0.24">0.24 in</quantity>
the two adult female
<typeStatus id="C64E8801951BFF92DAC0FF30897FFF1C" box="[151,258,224,248]" pageId="10" pageNumber="483" type="paratype">paratypes</typeStatus>
of A.
<taxonomicName id="DEF54D20951BFF92DB07FF0F89AEFF1C" box="[336,467,223,248]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92DB07FF0F89AEFF1C" box="[336,467,223,248]" italics="true" pageId="10" pageNumber="483">phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92DB8BFF308A45FF1C" box="[476,568,224,248]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92DB8BFF308A45FF1C" bold="true" box="[476,568,224,248]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
vs. 0.03), and less scales between the nasals (NS 78 vs. 9). The character
<collectionCode id="7FE4AE66951BFF92DAC0FED488A5FEF8" box="[151,216,260,284]" pageId="10" pageNumber="483">DIAS</collectionCode>
is given both as 4.5 and
<quantity id="DE0D9B46951BFF92DBA8FED48A25FEF8" box="[511,600,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="483" unit="mm" value="1.9">1.9 mm</quantity>
in the original description of
<taxonomicName id="DEF54D20951BFF92D9E4FED48C27FEF9" box="[947,1114,260,285]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB1951BFF92D9E4FED48C27FEF9" box="[947,1114,260,285]" italics="true" pageId="10" pageNumber="483">A. brachypoda</emphasis>
</taxonomicName>
, which is based on a single specimen, so this character cannot be used here for comparison. The methodology provided for measuring
<collectionCode id="7FE4AE66951BFF92DF13FEF88DE0FEA4" box="[1348,1437,296,320]" pageId="10" pageNumber="483">FOREL</collectionCode>
and
<collectionCode id="7FE4AE66951BFF92DA9FFE9D8962FE80" box="[200,287,333,356]" pageId="10" pageNumber="483">HINDL</collectionCode>
in the original description of
<taxonomicName id="DEF54D20951BFF92D82DFE9C8B5CFE81" box="[634,801,332,357]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB1951BFF92D82DFE9C8B5CFE81" box="[634,801,332,357]" italics="true" pageId="10" pageNumber="483">A. brachypoda</emphasis>
</taxonomicName>
is insufficiently described to allow verifying that these measurements were taken according to our methodology, so we could not use these characters here; CS,
<collectionCode id="7FE4AE66951BFF92DF7DFEA08D1EFE6C" box="[1322,1379,368,392]" httpUri="http://grbio.org/cool/2iqx-b0mr" name="North Carolina State University" pageId="10" pageNumber="483">NCS</collectionCode>
,
<collectionCode id="7FE4AE66951BFF92DF39FEA08DE5FE63" box="[1390,1432,368,391]" pageId="10" pageNumber="483">NR</collectionCode>
,
<collectionCode id="7FE4AE66951BFF92DAC0FE44888CFE48" box="[151,241,404,428]" pageId="10" pageNumber="483">NSCSL</collectionCode>
,
<collectionCode id="7FE4AE66951BFF92DAA9FE448928FE48" box="[254,341,404,428]" pageId="10" pageNumber="483">NSSLC</collectionCode>
and
<collectionCode id="7FE4AE66951BFF92DBD8FE44899BFE48" box="[399,486,404,428]" pageId="10" pageNumber="483">NSSOS</collectionCode>
were not provided in the original description of
<taxonomicName id="DEF54D20951BFF92DE42FE448CC6FE49" box="[1045,1211,404,429]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="brachypoda">
<emphasis id="2B81EAB1951BFF92DE42FE448CC6FE49" box="[1045,1211,404,429]" italics="true" pageId="10" pageNumber="483">A. brachypoda</emphasis>
</taxonomicName>
. A.
<taxonomicName id="DEF54D20951BFF92DEB8FE448D0FFE49" box="[1263,1394,404,429]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92DEB8FE448D0FFE49" box="[1263,1394,404,429]" italics="true" pageId="10" pageNumber="483">phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92DF2DFE4488B5FE34" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92DF2DFE4488B5FE34" bold="true" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="DEF54D20951BFF92DBA1FE688B72FE34" authority="Gunther, 1861" authorityName="Gunther" authorityYear="1861" box="[502,783,439,464]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="capra">
<emphasis id="2B81EAB1951BFF92DBA1FE688A2BFE34" box="[502,598,440,464]" italics="true" pageId="10" pageNumber="483">A. capra</emphasis>
(Günther, 1861)
</taxonomicName>
based on a generally larger tympanum (TD/HD 0.220.33 vs. 0.210.23), shorter spines in the nuchal crest (NSL/HL 0.210.39 vs. 0.420.43), slightly higher maximal length of dorsal crest spines (8.3 vs.
<quantity id="DE0D9B46951BFF92D86EFE2F8AE9FDFC" box="[569,660,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="483" unit="mm" value="6.8">6.8 mm</quantity>
), much higher number of scales in the diastema between nuchal and dorsal crests (1217 vs. 47), presence of occipital spines (vs. absence), lower number of scales between the nasals (78 vs. 9), higher number of scales between the fifth canthals (1213 vs. 9), generally more scales between the seventh supralabial and the sixth canthal (1114 vs. 911) and a notably less developed gular pouch (GP 02 vs. 3 4). From
<taxonomicName id="DEF54D20951BFF92DB56FD4089ADFD4C" box="[257,464,655,680]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="2B81EAB1951BFF92DB56FD4089ADFD4C" box="[257,464,655,680]" italics="true" pageId="10" pageNumber="483">A. cardamomensis</emphasis>
</taxonomicName>
, it can be separated based on a much lower maximal length of dorsal crest spines (8.3 vs.
<quantity id="DE0D9B46951BFF92DAEAFD64895EFD28" box="[189,291,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="10" pageNumber="483" unit="mm" value="14.2">14.2 mm</quantity>
, with a sudden decrease in size before dorsum mid-length in male
<taxonomicName id="DEF54D20951BFF92DE5BFD648CD3FD29" box="[1036,1198,692,717]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92DE5BFD648CD3FD29" box="[1036,1198,692,717]" italics="true" pageId="10" pageNumber="483">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92DEE3FD648D73FD28" box="[1204,1294,692,716]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92DEE3FD648D73FD28" bold="true" box="[1204,1294,692,716]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
, posterior to dorsum mid-length in male
<taxonomicName id="DEF54D20951BFF92DB98FD088AE0FD14" box="[463,669,727,752]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="2B81EAB1951BFF92DB98FD088AE0FD14" box="[463,669,727,752]" italics="true" pageId="10" pageNumber="483">A. cardamomensis</emphasis>
</taxonomicName>
), a generally higher number of scales in the diastema separating the nuchal and the dorsal crests (1217 vs 615), a lower maximal length of occipital spines (9.5 vs.
<quantity id="DE0D9B46951BFF92DE86FD2C8D4BFCF0" box="[1233,1334,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="10" pageNumber="483" unit="mm" value="13.3">13.3 mm</quantity>
, and OS/ HL 0.130.30 vs. 0.240.56), a generally less-developed gular pouch (GP 02 vs. 14); another character to separate them is based on coloration: in male
<taxonomicName id="DEF54D20951BFF92D8FDFC948B33FCB9" box="[682,846,836,861]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92D8FDFC948B33FCB9" box="[682,846,836,861]" italics="true" pageId="10" pageNumber="483">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92D900FC948BCEFCB8" box="[855,947,836,860]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92D900FC948BCEFCB8" bold="true" box="[855,947,836,860]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
the black eye patch extends posteriorly to reach the nuchal crest, while it never does in
<taxonomicName id="DEF54D20951BFF92D8D8FCB88B20FC64" box="[655,861,871,896]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="2B81EAB1951BFF92D8D8FCB88B20FC64" box="[655,861,871,896]" italics="true" pageId="10" pageNumber="483">A. cardamomensis</emphasis>
</taxonomicName>
(see species description and photographs in Wood
<emphasis id="2B81EAB1951BFF92DAC0FC5D88ADFC41" box="[151,208,908,933]" italics="true" pageId="10" pageNumber="483">et al.</emphasis>
2010 and all photographs of
<taxonomicName id="DEF54D20951BFF92D84AFC5C8AB2FC40" box="[541,719,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB1951BFF92D84AFC5C8A4BFC47" box="[541,566,908,931]" italics="true" pageId="10" pageNumber="483">A.</emphasis>
cf.
<emphasis id="2B81EAB1951BFF92D835FC5D8AB2FC40" box="[610,719,909,932]" italics="true" pageId="10" pageNumber="483">crucigera</emphasis>
</taxonomicName>
from Cardamom Mountains [=
<taxonomicName id="DEF54D20951BFF92DE6CFC5C8D77FC41" box="[1083,1290,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="2B81EAB1951BFF92DE6CFC5C8D77FC41" box="[1083,1290,908,933]" italics="true" pageId="10" pageNumber="483">A. cardamomensis</emphasis>
</taxonomicName>
] in
<bibRefCitation id="7D644B52951BFF92DF6FFC5C88A9FC2C" author="Manthey" pageId="10" pageNumber="483" refString="Manthey, U. (2008) Agamid Lizards of Southern Asia - Agamen des sudlichen Asien - Draconinae 1. Terralog Vol. 7 a. Edition Chimaira, Frankfurt am Main, 160 pp." type="book" year="2008">Manthey 2008</bibRefCitation>
).
<taxonomicName id="DEF54D20951BFF92DABFFC7F8A6CFC2C" box="[232,529,943,968]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92DABFFC7F8A6CFC2C" box="[232,529,943,968]" italics="true" pageId="10" pageNumber="483">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92D84DFC608A08FC2C" box="[538,629,944,968]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92D84DFC608A08FC2C" bold="true" box="[538,629,944,968]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguishable from
<taxonomicName id="DEF54D20951BFF92D9DBFC608CB2FC2C" authority="Gunther, 1861" authorityName="Gunther" authorityYear="1861" box="[908,1231,943,968]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="coronata">
<emphasis id="2B81EAB1951BFF92D9DBFC608C6DFC2C" box="[908,1040,944,968]" italics="true" pageId="10" pageNumber="483">A. coronata</emphasis>
(Günther, 1861)
</taxonomicName>
based on a larger tympanum (0.220.33 vs.
<quantity id="DE0D9B46951BFF92DB91FC048A0DFC08" box="[454,624,980,1004]" metricMagnitude="-4" metricUnit="m" metricValue="1.5500000000000003" metricValueMax="1.7000000000000002" metricValueMin="1.4000000000000001" pageId="10" pageNumber="483" unit="mm" value="0.155" valueMax="0.17" valueMin="0.14">0.140.17 mm</quantity>
), presence of postorbital spines, occipital spines, of nuchal and dorsal crests (vs. absence), higher number of subdigital lamellae on the fourth finger (1517 vs. 1314), higher number of subdigital lamellae on the fourth toe (2124 vs. 1719), longer tail (TaL/SVL 1.41.7 vs. 0.61.0), higher number of scales between the fifth canthals (1213 vs. 811), higher number of scales from the fifth canthal to the fifth supralabial (810 vs. 56), lower number of scales between the nasals and the rostrals (12 vs. 34), generally higher number of scales between the seventh supralabial and the sixth canthal (1114 vs. 611), presence of a black, diamond shaped, nuchal collar (vs. absence) and presence of a black eye patch (vs. absence). From
<taxonomicName id="DEF54D20951BFF92DFD3FB7C8979FB0C" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB1951BFF92DFD3FB7C8979FB0C" italics="true" pageId="10" pageNumber="483">A. crucigera</emphasis>
</taxonomicName>
, it is distinguished by a larger tympanum (TD/HD 0.220.33 vs. 0.140.21), a higher maximal length of postorbital spines (11.8 vs.
<quantity id="DE0D9B46951BFF92DB81FB248A4EFAE8" box="[470,563,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="10" pageNumber="483" unit="mm" value="7.8">7.8 mm</quantity>
), a distinctly higher maximal length of occipital spines (9.5 vs.
<quantity id="DE0D9B46951BFF92DF4AFB248D07FAE8" box="[1309,1402,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="483" unit="mm" value="4.9">4.9 mm</quantity>
), a higher maximal length of nuchal crest spines (12.2 vs.
<quantity id="DE0D9B46951BFF92D8ADFAC88B29FAD4" box="[762,852,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="10" pageNumber="483" unit="mm" value="8.9">8.9 mm</quantity>
), a higher maximal length of dorsal crest spines (8.3 vs.
<quantity id="DE0D9B46951BFF92DA97FAEC8960FAB0" box="[192,285,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="483" unit="mm" value="5.5">5.5 mm</quantity>
); although several morphometrical characters of both species are overlapping (see
<tableCitation id="54770318951BFF92DE80FAEC8D52FAB0" box="[1239,1327,1340,1365]" captionStart="TABLE 2" captionStartId="18.[119,197,528,548]" captionText="TABLE 2. Comparison of morphometrical (in mm) and meristic data for all currently recognized species of Acanthosaura and A. phuketensis sp. nov. For the ? see the comparison section." pageId="10" pageNumber="483">Table 2</tableCitation>
), an easy character to separate them is based on coloration: in male
<taxonomicName id="DEF54D20951BFF92D914FAB08B94FA9C" box="[835,1001,1375,1400]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92D914FAB08B94FA9C" box="[835,1001,1375,1400]" italics="true" pageId="10" pageNumber="483">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92D9A4FAB08C2DFA9C" box="[1011,1104,1376,1400]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92D9A4FAB08C2DFA9C" bold="true" box="[1011,1104,1376,1400]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
the black eye patch extends posteriorly to reach the nuchal crest, while it never does in
<taxonomicName id="DEF54D20951BFF92D90BFA548B90FA78" box="[860,1005,1412,1436]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="crucigera">
<emphasis id="2B81EAB1951BFF92D90BFA548B90FA78" box="[860,1005,1412,1436]" italics="true" pageId="10" pageNumber="483">A. crucigera</emphasis>
</taxonomicName>
(see species original description by
<bibRefCitation id="7D644B52951BFF92DAC0FA778932FA24" author="Boulenger" box="[151,335,1447,1472]" pageId="10" pageNumber="483" refString="Boulenger, G. A. (1885) Catalogue of the lizards in the British Museum (Natural History). Second edition. Volume I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. British Museum (Natural History), London, Xii + 436 pp., XXXII Pls." type="book" year="1885">Boulenger 1885</bibRefCitation>
). From
<taxonomicName id="DEF54D20951BFF92DBF2FA788A80FA24" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[421,765,1447,1472]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="lepidogaster">
<emphasis id="2B81EAB1951BFF92DBF2FA788A2DFA24" box="[421,592,1447,1472]" italics="true" pageId="10" pageNumber="483">A. lepidogaster</emphasis>
(Cuvier, 1829)
</taxonomicName>
,
<taxonomicName id="DEF54D20951BFF92D95EFA788BD6FA24" box="[777,939,1447,1472]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92D95EFA788BD6FA24" box="[777,939,1447,1472]" italics="true" pageId="10" pageNumber="483">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92D9E5FA788C71FA24" box="[946,1036,1448,1472]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92D9E5FA788C71FA24" bold="true" box="[946,1036,1448,1472]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated based on its much longer postorbital spines (4.611.8 vs.
<quantity id="DE0D9B46951BFF92D81CFA1C8AA9FA00" box="[587,724,1484,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="10" pageNumber="483" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
, PS/HL 0.230.38 vs. 0.060.11), much higher maximal length of nuchal crest spines (4.112.2 vs.
<quantity id="DE0D9B46951BFF92D87DFA208ACEF9EC" box="[554,691,1520,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="10" pageNumber="483" unit="mm" value="3.15" valueMax="3.4" valueMin="2.9">2.93.4 mm</quantity>
, NSL/HL 0.210.39 vs. 0.120.15) and of occipital spines (9.5 vs.
<quantity id="DE0D9B46951BFF92DAC0F9C4888CF9C8" box="[151,241,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="483" unit="mm" value="3.4">3.4 mm</quantity>
), its much higher maximal length of dorsal crest spines (8.3 vs.
<quantity id="DE0D9B46951BFF92D996F9C48C67F9C8" box="[961,1050,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="483" unit="mm" value="2.7">2.7 mm</quantity>
, DS/HL 0.110.26 vs. 0.070.12), its generally lower number of subdigital lamellae on the 4th finger (1517 vs. 1719), higher number of scales between the fifth canthals (1213 vs. 710), higher number of scales between the seventh supralabial and the sixth canthal (1114 vs. 10) and lower number of scales in contact with the mental (4 vs. 5).
<taxonomicName id="DEF54D20951BFF92DE3CF9538D70F97F" box="[1131,1293,1666,1691]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951BFF92DE3CF9538D70F97F" box="[1131,1293,1666,1691]" italics="true" pageId="10" pageNumber="483">A. phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951BFF92DF43F9538D13F97F" box="[1300,1390,1667,1691]" pageId="10" pageNumber="483" rank="species">
<emphasis id="2B81EAB1951BFF92DF43F9538D13F97F" bold="true" box="[1300,1390,1667,1691]" pageId="10" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated from
<taxonomicName id="DEF54D20951BFF92DB3AF9778B1BF95B" authority="Orlov, Nguyen &amp; Nguyen, 2006" authorityName="Orlov, Nguyen &amp; Nguyen" authorityYear="2006" box="[365,870,1702,1728]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="nataliae">
<emphasis id="2B81EAB1951BFF92DB3AF9778996F95B" box="[365,491,1702,1727]" italics="true" pageId="10" pageNumber="483">A. nataliae</emphasis>
<bibRefCitation id="7D644B52951BFF92DBA3F9768B1BF95B" author="Orlov" box="[500,870,1702,1728]" pageId="10" pageNumber="483" refString="Orlov, N. L., Nguyen, Q. T. &amp; Nguyen, V. S. (2006) A new Acanthosaura allied to A. capra Gunther, 1861 (Agamidae, Sauria) from Central Vietnam and Southern Laos. Russian Journal of Herpetology, 13 (1), 61 - 76." type="journal article" year="2006">Orlov, Nguyen &amp; Nguyen, 2006</bibRefCitation>
</taxonomicName>
by its much lower maximal length of postorbital spines (11.8 vs.
<quantity id="DE0D9B46951BFF92DB05F91B89C1F907" box="[338,444,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="10" pageNumber="483" unit="mm" value="17.8">17.8 mm</quantity>
), nuchal crest spines (12.2 vs.
<quantity id="DE0D9B46951BFF92D977F91B8BF5F906" box="[800,904,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="10" pageNumber="483" unit="mm" value="23.8">23.8 mm</quantity>
, NSL/HL 0.210.39 vs. 0.58) and dorsal crest spines (8.3 vs.
<quantity id="DE0D9B46951BFF92DB17F93F89DAF8E3" box="[320,423,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="10" pageNumber="483" unit="mm" value="17.7">17.7 mm</quantity>
, DS/HL 0.110.26 vs. 0.44), a longer diastema between nuchal and dorsal crests (3.67.6 vs.
<quantity id="DE0D9B46951BFF92DA96F8C28960F8CE" box="[193,285,1810,1835]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="483" unit="mm" value="2.5">2.5 mm</quantity>
,
<collectionCode id="7FE4AE66951BFF92DB7BF8C38910F8CF" box="[300,365,1811,1835]" pageId="10" pageNumber="483">DIAS</collectionCode>
/SVL 0.050.08 vs. 0.04) with more scales (1217 vs. 10), presence of occipital spines (vs. absence), lower number of scales between the fifth canthals (1213 vs. 14), lower number of scales between the seventh supralabial and the sixth canthal (1114 vs. 16), presence of a black, diamond shaped, nuchal collar (vs. absence), presence of light knee patch (vs. absence) and much lesser development of gular pouch (GP 02 vs. 4). From
<taxonomicName id="DEF54D20951BFF92DA89F87389D0F85F" box="[222,429,1955,1979]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="titiwangsaensis">
<emphasis id="2B81EAB1951BFF92DA89F87389D0F85F" box="[222,429,1955,1979]" italics="true" pageId="10" pageNumber="483">A. titiwangsaensis</emphasis>
</taxonomicName>
, it can be distinguished by its larger tympanum (TD/HD 0.220.33 vs. 0.170.20), its much longer postorbital spines (4.611.8 vs.
<quantity id="DE0D9B46951BFF92D8E5F8178B3CF83B" box="[690,833,1991,2015]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.4" metricValueMin="3.3" pageId="10" pageNumber="483" unit="mm" value="3.85" valueMax="4.4" valueMin="3.3">3.34.4 mm</quantity>
, PS/HL 0.230.38 vs. 0.140.18), its much longer occipital spines (2.69.5 vs.
<quantity id="DE0D9B46951AFF93DBB3FF488A0DFF4B" box="[484,624,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="11" pageNumber="484" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.82.3 mm</quantity>
, OS/HL 0.130.30 vs. 0.090.10), its much higher maximal length of nuchal crest spines (12.2 vs.
<quantity id="DE0D9B46951AFF93DB89FF6C8A4BFF30" box="[478,566,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="484" unit="mm" value="4.4">4.4 mm</quantity>
, NSL/HL 0.210.39 vs. 0.110.18), its much longer dorsal crest spines (2.3 8.3 vs.
<quantity id="DE0D9B46951AFF93DABCFF30890BFF13" box="[235,374,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="11" pageNumber="484" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.72.1 mm</quantity>
, DS/HL 0.110.26 vs. 0.070.09), its higher number of ventrals (5767 vs. 4757), its lower number of subdigital lamellae on 4th finger (1517 vs. 2021), lower number of scales bordering the rostral scale (58 vs. 9), lower number of scales bordering the mental (4 vs. 5), presence of light knee patch (vs. absence) and lesser development of gular pouch (02 vs. 24).
</paragraph>
<caption id="4D8A662B9518FF91DAC0FB888B9EFB69" httpUri="https://zenodo.org/record/399938/files/figure.png" pageId="9" pageNumber="482" targetBox="[151,1436,176,1090]" targetPageId="9">
<paragraph id="194A36A39518FF91DAC0FB888B9EFB69" blockId="9.[151,1435,1112,1165]" pageId="9" pageNumber="482">
<emphasis id="2B81EAB19518FF91DAC0FB88896FFB8A" bold="true" box="[151,274,1112,1134]" pageId="9" pageNumber="482">FIGURE 7.</emphasis>
Live adult male
<taxonomicName id="DEF54D209518FF91DBEAFB888ABAFB8A" box="[445,711,1112,1134]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB19518FF91DBEAFB888ABAFB8A" box="[445,711,1112,1134]" italics="true" pageId="9" pageNumber="482">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA9518FF91D89AFB888B63FB8A" box="[717,798,1112,1134]" pageId="9" pageNumber="482" rank="species">
<emphasis id="2B81EAB19518FF91D89AFB888B63FB8A" bold="true" box="[717,798,1112,1134]" pageId="9" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="2B81EAB19518FF91D973FB898B1FFB8A" box="[804,866,1113,1134]" italics="true" pageId="9" pageNumber="482">in situ</emphasis>
at the type locality of
<taxonomicName id="DEF54D209518FF91DE12FB888D28FB8A" box="[1093,1365,1112,1134]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="482" phylum="Chordata" rank="species" species="ranongensis">
<emphasis id="2B81EAB19518FF91DE12FB888D28FB8A" box="[1093,1365,1112,1134]" italics="true" pageId="9" pageNumber="482">Cyrtodactylus ranongensis</emphasis>
</taxonomicName>
in Suk Samran District, Ranong Province, southwestern Thailand. Photo by M. Sumontha.
</paragraph>
</caption>
<caption id="4D8A662B951AFF93DAC0FE468D02FE29" ID-Table-UUID="4D8A662B951AFF93DAC0FE468D02FE29" httpUri="http://table.plazi.org/id/4D8A662B951AFF93DAC0FE468D02FE29" pageId="11" pageNumber="484" targetBox="[159,1431,488,1980]" targetIsTable="true" targetPageId="11">
<paragraph id="194A36A3951AFF93DAC0FE468D02FE29" blockId="11.[151,1435,405,461]" pageId="11" pageNumber="484">
<emphasis id="2B81EAB1951AFF93DAC0FE468970FE49" bold="true" box="[151,269,406,429]" pageId="11" pageNumber="484">TABLE 1.</emphasis>
Morphometrical (in mm) and meristic data for the type series of
<taxonomicName id="DEF54D20951AFF93D9B9FE458D74FE49" box="[1006,1289,405,429]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="484" phylum="Chordata" rank="species" species="phuketensis" status="sp. nov.">
<emphasis id="2B81EAB1951AFF93D9B9FE458D74FE49" box="[1006,1289,405,429]" italics="true" pageId="11" pageNumber="484">Acanthosaura phuketensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B257CA951AFF93DF43FE468D10FE49" box="[1300,1389,406,429]" pageId="11" pageNumber="484" rank="species">
<emphasis id="2B81EAB1951AFF93DF43FE468D10FE49" bold="true" box="[1300,1389,406,429]" pageId="11" pageNumber="484">sp. nov.</emphasis>
</taxonomicNameLabel>
For character abbreviations see material and methods. Paired meristic characters are given left/right. NA = not applicable.
</paragraph>
</caption>
<paragraph id="194A36A3951AFF93DB32FE388D58F858" pageId="11" pageNumber="484">
<table id="6BF5C403951A0067DAC8FE388DEAF858" box="[159,1431,488,1980]" colsContinueIn="12.[159,1405,199,1074]" gridcols="6" gridrows="19" pageId="11" pageNumber="484">
<tr id="A7C534E1951A0067DAC8FE388DEAFDBE" box="[159,1431,488,602]" gridrow="0" pageId="11" pageNumber="484" rowspan-0="1">
<th id="E4145D9D951A0067DB32FE388A75FDBE" box="[357,520,488,602]" gridcol="1" gridrow="0" pageId="11" pageNumber="484">Holotype THNHM 22542 Adult male</th>
<th id="E4145D9D951A0067D874FE388B40FDBE" box="[547,829,488,602]" gridcol="2" gridrow="0" pageId="11" pageNumber="484">Paratype Paratype IRSNB 2681 PSUZC-RT Adult male 2012.5 Adult female</th>
<th id="E4145D9D951A0067D924FE388C68FDBE" box="[883,1045,488,602]" gridcol="3" gridrow="0" pageId="11" pageNumber="484">Paratype THNHM 22543 Adult female</th>
<th id="E4145D9D951A0067DE78FE388CB0FDBE" box="[1071,1229,488,602]" gridcol="4" gridrow="0" pageId="11" pageNumber="484">Paratype QSMI 1166 Subadult male</th>
<th id="E4145D9D951A0067DEBCFE388DEAFDBE" box="[1259,1431,488,602]" gridcol="5" gridrow="0" pageId="11" pageNumber="484">Paratype PSUZC-RT 2012.6 Subadult male</th>
</tr>
<tr id="A7C534E1951A0067DAC8FDBD8DEAFD4F" box="[159,1431,621,683]" gridrow="1" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FDBD896DFD4F" box="[159,272,621,683]" gridcol="0" gridrow="1" pageId="11" pageNumber="484">SVL TaL</th>
<td id="E4145D9D951A0067DB32FDBD8A75FD4F" box="[357,520,621,683]" gridcol="1" gridrow="1" pageId="11" pageNumber="484">123.5 205.6</td>
<td id="E4145D9D951A0067D874FDBD8B40FD4F" box="[547,829,621,683]" gridcol="2" gridrow="1" pageId="11" pageNumber="484">96.3 118.4&gt;101.2 189.8</td>
<td id="E4145D9D951A0067D924FDBD8C68FD4F" box="[883,1045,621,683]" gridcol="3" gridrow="1" pageId="11" pageNumber="484">100.8 145.0</td>
<td id="E4145D9D951A0067DE78FDBD8CB0FD4F" box="[1071,1229,621,683]" gridcol="4" gridrow="1" pageId="11" pageNumber="484">69.2 107.0</td>
<td id="E4145D9D951A0067DEBCFDBD8DEAFD4F" box="[1259,1431,621,683]" gridcol="5" gridrow="1" pageId="11" pageNumber="484">74.2 113.2</td>
</tr>
<tr id="A7C534E1951A0067DAC8FD6E8DEAFD18" box="[159,1431,702,764]" gridrow="2" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FD6E896DFD18" box="[159,272,702,764]" gridcol="0" gridrow="2" pageId="11" pageNumber="484">TaL/SVL TBW</th>
<td id="E4145D9D951A0067DB32FD6E8A75FD18" box="[357,520,702,764]" gridcol="1" gridrow="2" pageId="11" pageNumber="484">1.7 14.3</td>
<td id="E4145D9D951A0067D874FD6E8B40FD18" box="[547,829,702,764]" gridcol="2" gridrow="2" pageId="11" pageNumber="484">NA 1.6 12.1 14.5</td>
<td id="E4145D9D951A0067D924FD6E8C68FD18" box="[883,1045,702,764]" gridcol="3" gridrow="2" pageId="11" pageNumber="484">1.4 9.2</td>
<td id="E4145D9D951A0067DE78FD6E8CB0FD18" box="[1071,1229,702,764]" gridcol="4" gridrow="2" pageId="11" pageNumber="484">1.5 5.5</td>
<td id="E4145D9D951A0067DEBCFD6E8DEAFD18" box="[1259,1431,702,764]" gridcol="5" gridrow="2" pageId="11" pageNumber="484">1.5 5.4</td>
</tr>
<tr id="A7C534E1951A0067DAC8FCDF8DEAFCA9" box="[159,1431,783,845]" gridrow="3" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FCDF896DFCA9" box="[159,272,783,845]" gridcol="0" gridrow="3" pageId="11" pageNumber="484">HL HW</th>
<td id="E4145D9D951A0067DB32FCDF8A75FCA9" box="[357,520,783,845]" gridcol="1" gridrow="3" pageId="11" pageNumber="484">31.4 22.8</td>
<td id="E4145D9D951A0067D874FCDF8B40FCA9" box="[547,829,783,845]" gridcol="2" gridrow="3" pageId="11" pageNumber="484">26.9 29.6 18.0 21.2</td>
<td id="E4145D9D951A0067D924FCDF8C68FCA9" box="[883,1045,783,845]" gridcol="3" gridrow="3" pageId="11" pageNumber="484">27.4 19.1</td>
<td id="E4145D9D951A0067DE78FCDF8CB0FCA9" box="[1071,1229,783,845]" gridcol="4" gridrow="3" pageId="11" pageNumber="484">19.7 14.4</td>
<td id="E4145D9D951A0067DEBCFCDF8DEAFCA9" box="[1259,1431,783,845]" gridcol="5" gridrow="3" pageId="11" pageNumber="484">23.0 14.7</td>
</tr>
<tr id="A7C534E1951A0067DAC8FCB08DEAFC92" box="[159,1431,864,886]" gridrow="4" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FCB0896DFC92" box="[159,272,864,886]" gridcol="0" gridrow="4" pageId="11" pageNumber="484">HD</th>
<td id="E4145D9D951A0067DB32FCB08A75FC92" box="[357,520,864,886]" gridcol="1" gridrow="4" pageId="11" pageNumber="484">18.6</td>
<td id="E4145D9D951A0067D874FCB08B40FC92" box="[547,829,864,886]" gridcol="2" gridrow="4" pageId="11" pageNumber="484">14.8 18.6</td>
<td id="E4145D9D951A0067D924FCB08C68FC92" box="[883,1045,864,886]" gridcol="3" gridrow="4" pageId="11" pageNumber="484">16.3</td>
<td id="E4145D9D951A0067DE78FCB08CB0FC92" box="[1071,1229,864,886]" gridcol="4" gridrow="4" pageId="11" pageNumber="484">10.9</td>
<td id="E4145D9D951A0067DEBCFCB08DEAFC92" box="[1259,1431,864,886]" gridcol="5" gridrow="4" pageId="11" pageNumber="484">12.3</td>
</tr>
<tr id="A7C534E1951A0067DAC8FC598DEAFC23" box="[159,1431,905,967]" gridrow="5" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FC59896DFC23" box="[159,272,905,967]" gridcol="0" gridrow="5" pageId="11" pageNumber="484">SL ORBIT</th>
<td id="E4145D9D951A0067DB32FC598A75FC23" box="[357,520,905,967]" gridcol="1" gridrow="5" pageId="11" pageNumber="484">11.0 10.5</td>
<td id="E4145D9D951A0067D874FC598B40FC23" box="[547,829,905,967]" gridcol="2" gridrow="5" pageId="11" pageNumber="484">10.2 10.1 9.7 11.2</td>
<td id="E4145D9D951A0067D924FC598C68FC23" box="[883,1045,905,967]" gridcol="3" gridrow="5" pageId="11" pageNumber="484">9.3 9.0</td>
<td id="E4145D9D951A0067DE78FC598CB0FC23" box="[1071,1229,905,967]" gridcol="4" gridrow="5" pageId="11" pageNumber="484">6.8 6.6</td>
<td id="E4145D9D951A0067DEBCFC598DEAFC23" box="[1259,1431,905,967]" gridcol="5" gridrow="5" pageId="11" pageNumber="484">8.3 7.3</td>
</tr>
<tr id="A7C534E1951A0067DAC8FC0A8DEAFBFC" box="[159,1431,986,1048]" gridrow="6" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FC0A896DFBFC" box="[159,272,986,1048]" gridcol="0" gridrow="6" pageId="11" pageNumber="484">EYE TD</th>
<td id="E4145D9D951A0067DB32FC0A8A75FBFC" box="[357,520,986,1048]" gridcol="1" gridrow="6" pageId="11" pageNumber="484">6.3 4.7</td>
<td id="E4145D9D951A0067D874FC0A8B40FBFC" box="[547,829,986,1048]" gridcol="2" gridrow="6" pageId="11" pageNumber="484">6.5 4.7 3.2 4.3</td>
<td id="E4145D9D951A0067D924FC0A8C68FBFC" box="[883,1045,986,1048]" gridcol="3" gridrow="6" pageId="11" pageNumber="484">4.0 3.6</td>
<td id="E4145D9D951A0067DE78FC0A8CB0FBFC" box="[1071,1229,986,1048]" gridcol="4" gridrow="6" pageId="11" pageNumber="484">3.7 3.5</td>
<td id="E4145D9D951A0067DEBCFC0A8DEAFBFC" box="[1259,1431,986,1048]" gridcol="5" gridrow="6" pageId="11" pageNumber="484">3.7 4.0</td>
</tr>
<tr id="A7C534E1951A0067DAC8FBFB8DEAFB8D" box="[159,1431,1067,1129]" gridrow="7" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FBFB896DFB8D" box="[159,272,1067,1129]" gridcol="0" gridrow="7" pageId="11" pageNumber="484">TD/HD TN</th>
<td id="E4145D9D951A0067DB32FBFB8A75FB8D" box="[357,520,1067,1129]" gridcol="1" gridrow="7" pageId="11" pageNumber="484">0.25 0</td>
<td id="E4145D9D951A0067D874FBFB8B40FB8D" box="[547,829,1067,1129]" gridcol="2" gridrow="7" pageId="11" pageNumber="484">0.22 0.23 0 0</td>
<td id="E4145D9D951A0067D924FBFB8C68FB8D" box="[883,1045,1067,1129]" gridcol="3" gridrow="7" pageId="11" pageNumber="484">0.22 0</td>
<td id="E4145D9D951A0067DE78FBFB8CB0FB8D" box="[1071,1229,1067,1129]" gridcol="4" gridrow="7" pageId="11" pageNumber="484">0.32 0</td>
<td id="E4145D9D951A0067DEBCFBFB8DEAFB8D" box="[1259,1431,1067,1129]" gridcol="5" gridrow="7" pageId="11" pageNumber="484">0.33 0</td>
</tr>
<tr id="A7C534E1951A0067DAC8FBAC8DEAFB5E" box="[159,1431,1148,1210]" gridrow="8" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FBAC896DFB5E" box="[159,272,1148,1210]" gridcol="0" gridrow="8" pageId="11" pageNumber="484">PS PS/HL</th>
<td id="E4145D9D951A0067DB32FBAC8A75FB5E" box="[357,520,1148,1210]" gridcol="1" gridrow="8" pageId="11" pageNumber="484">11.8 0.38</td>
<td id="E4145D9D951A0067D874FBAC8B40FB5E" box="[547,829,1148,1210]" gridcol="2" gridrow="8" pageId="11" pageNumber="484">9.1 8.0 0.34 0.27</td>
<td id="E4145D9D951A0067D924FBAC8C68FB5E" box="[883,1045,1148,1210]" gridcol="3" gridrow="8" pageId="11" pageNumber="484">7.7 0.28</td>
<td id="E4145D9D951A0067DE78FBAC8CB0FB5E" box="[1071,1229,1148,1210]" gridcol="4" gridrow="8" pageId="11" pageNumber="484">4.6 0.23</td>
<td id="E4145D9D951A0067DEBCFBAC8DEAFB5E" box="[1259,1431,1148,1210]" gridcol="5" gridrow="8" pageId="11" pageNumber="484">6.5 0.28</td>
</tr>
<tr id="A7C534E1951A0067DAC8FB1D8DEAFAEF" box="[159,1431,1229,1291]" gridrow="9" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FB1D896DFAEF" box="[159,272,1229,1291]" gridcol="0" gridrow="9" pageId="11" pageNumber="484">NSL NSL/HL</th>
<td id="E4145D9D951A0067DB32FB1D8A75FAEF" box="[357,520,1229,1291]" gridcol="1" gridrow="9" pageId="11" pageNumber="484">12.2 0.39</td>
<td id="E4145D9D951A0067D874FB1D8B40FAEF" box="[547,829,1229,1291]" gridcol="2" gridrow="9" pageId="11" pageNumber="484">9.1 6.8 0.34 0.23</td>
<td id="E4145D9D951A0067D924FB1D8C68FAEF" box="[883,1045,1229,1291]" gridcol="3" gridrow="9" pageId="11" pageNumber="484">8.1 0.30</td>
<td id="E4145D9D951A0067DE78FB1D8CB0FAEF" box="[1071,1229,1229,1291]" gridcol="4" gridrow="9" pageId="11" pageNumber="484">4.1 0.21</td>
<td id="E4145D9D951A0067DEBCFB1D8DEAFAEF" box="[1259,1431,1229,1291]" gridcol="5" gridrow="9" pageId="11" pageNumber="484">6.5 0.28</td>
</tr>
<tr id="A7C534E1951A0067DAC8FACE8DEAFAB8" box="[159,1431,1310,1372]" gridrow="10" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FACE896DFAB8" box="[159,272,1310,1372]" gridcol="0" gridrow="10" pageId="11" pageNumber="484">DS DS/HL</th>
<td id="E4145D9D951A0067DB32FACE8A75FAB8" box="[357,520,1310,1372]" gridcol="1" gridrow="10" pageId="11" pageNumber="484">8.3 0.26</td>
<td id="E4145D9D951A0067D874FACE8B40FAB8" box="[547,829,1310,1372]" gridcol="2" gridrow="10" pageId="11" pageNumber="484">6.5 3.3 0.24 0.11</td>
<td id="E4145D9D951A0067D924FACE8C68FAB8" box="[883,1045,1310,1372]" gridcol="3" gridrow="10" pageId="11" pageNumber="484">3.9 0.14</td>
<td id="E4145D9D951A0067DE78FACE8CB0FAB8" box="[1071,1229,1310,1372]" gridcol="4" gridrow="10" pageId="11" pageNumber="484">2.3 0.12</td>
<td id="E4145D9D951A0067DEBCFACE8DEAFAB8" box="[1259,1431,1310,1372]" gridcol="5" gridrow="10" pageId="11" pageNumber="484">2.5 0.11</td>
</tr>
<tr id="A7C534E1951A0067DAC8FABF8DEAFA49" box="[159,1431,1391,1453]" gridrow="11" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FABF896DFA49" box="[159,272,1391,1453]" gridcol="0" gridrow="11" pageId="11" pageNumber="484">WNC DIAS</th>
<td id="E4145D9D951A0067DB32FABF8A75FA49" box="[357,520,1391,1453]" gridcol="1" gridrow="11" pageId="11" pageNumber="484">2.9 7.6</td>
<td id="E4145D9D951A0067D874FABF8B40FA49" box="[547,829,1391,1453]" gridcol="2" gridrow="11" pageId="11" pageNumber="484">2.4 2.4 4.5 6.5</td>
<td id="E4145D9D951A0067D924FABF8C68FA49" box="[883,1045,1391,1453]" gridcol="3" gridrow="11" pageId="11" pageNumber="484">2.0 7.6</td>
<td id="E4145D9D951A0067DE78FABF8CB0FA49" box="[1071,1229,1391,1453]" gridcol="4" gridrow="11" pageId="11" pageNumber="484">1.4 3.6</td>
<td id="E4145D9D951A0067DEBCFABF8DEAFA49" box="[1259,1431,1391,1453]" gridcol="5" gridrow="11" pageId="11" pageNumber="484">1.7 3.8</td>
</tr>
<tr id="A7C534E1951A0067DAC8FA108DEAFA32" box="[159,1431,1472,1494]" gridrow="12" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FA10896DFA32" box="[159,272,1472,1494]" gridcol="0" gridrow="12" pageId="11" pageNumber="484">DIAS/SVL</th>
<td id="E4145D9D951A0067DB32FA108A75FA32" box="[357,520,1472,1494]" gridcol="1" gridrow="12" pageId="11" pageNumber="484">0.06</td>
<td id="E4145D9D951A0067D874FA108B40FA32" box="[547,829,1472,1494]" gridcol="2" gridrow="12" pageId="11" pageNumber="484">0.05 0.05</td>
<td id="E4145D9D951A0067D924FA108C68FA32" box="[883,1045,1472,1494]" gridcol="3" gridrow="12" pageId="11" pageNumber="484">0.08</td>
<td id="E4145D9D951A0067DE78FA108CB0FA32" box="[1071,1229,1472,1494]" gridcol="4" gridrow="12" pageId="11" pageNumber="484">0.05</td>
<td id="E4145D9D951A0067DEBCFA108DEAFA32" box="[1259,1431,1472,1494]" gridcol="5" gridrow="12" pageId="11" pageNumber="484">0.05</td>
</tr>
<tr id="A7C534E1951A0067DAC8FA398DEAF9C3" box="[159,1431,1513,1575]" gridrow="13" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8FA39896DF9C3" box="[159,272,1513,1575]" gridcol="0" gridrow="13" pageId="11" pageNumber="484">DIASN FOREL</th>
<td id="E4145D9D951A0067DB32FA398A75F9C3" box="[357,520,1513,1575]" gridcol="1" gridrow="13" pageId="11" pageNumber="484">17 38.8</td>
<td id="E4145D9D951A0067D874FA398B40F9C3" box="[547,829,1513,1575]" gridcol="2" gridrow="13" pageId="11" pageNumber="484">13 12 33.5 42.9</td>
<td id="E4145D9D951A0067D924FA398C68F9C3" box="[883,1045,1513,1575]" gridcol="3" gridrow="13" pageId="11" pageNumber="484">13 33.9</td>
<td id="E4145D9D951A0067DE78FA398CB0F9C3" box="[1071,1229,1513,1575]" gridcol="4" gridrow="13" pageId="11" pageNumber="484">15 22.3</td>
<td id="E4145D9D951A0067DEBCFA398DEAF9C3" box="[1259,1431,1513,1575]" gridcol="5" gridrow="13" pageId="11" pageNumber="484">14 27.5</td>
</tr>
<tr id="A7C534E1951A0067DAC8F9EA8DEAF99C" box="[159,1431,1594,1656]" gridrow="14" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8F9EA896DF99C" box="[159,272,1594,1656]" gridcol="0" gridrow="14" pageId="11" pageNumber="484">HINDL SUPRAL</th>
<td id="E4145D9D951A0067DB32F9EA8A75F99C" box="[357,520,1594,1656]" gridcol="1" gridrow="14" pageId="11" pageNumber="484">60.3 11/11</td>
<td id="E4145D9D951A0067D874F9EA8B40F99C" box="[547,829,1594,1656]" gridcol="2" gridrow="14" pageId="11" pageNumber="484">50.4 60.6 11/11 12/11</td>
<td id="E4145D9D951A0067D924F9EA8C68F99C" box="[883,1045,1594,1656]" gridcol="3" gridrow="14" pageId="11" pageNumber="484">49.1 10/11</td>
<td id="E4145D9D951A0067DE78F9EA8CB0F99C" box="[1071,1229,1594,1656]" gridcol="4" gridrow="14" pageId="11" pageNumber="484">38.2 12/11</td>
<td id="E4145D9D951A0067DEBCF9EA8DEAF99C" box="[1259,1431,1594,1656]" gridcol="5" gridrow="14" pageId="11" pageNumber="484">38.6 10/12</td>
</tr>
<tr id="A7C534E1951A0067DAC8F95B8DEAF92D" box="[159,1431,1675,1737]" gridrow="15" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8F95B896DF92D" box="[159,272,1675,1737]" gridcol="0" gridrow="15" pageId="11" pageNumber="484">INFRAL VENT</th>
<td id="E4145D9D951A0067DB32F95B8A75F92D" box="[357,520,1675,1737]" gridcol="1" gridrow="15" pageId="11" pageNumber="484">12/12 66</td>
<td id="E4145D9D951A0067D874F95B8B40F92D" box="[547,829,1675,1737]" gridcol="2" gridrow="15" pageId="11" pageNumber="484">11/11 12/12 57 59</td>
<td id="E4145D9D951A0067D924F95B8C68F92D" box="[883,1045,1675,1737]" gridcol="3" gridrow="15" pageId="11" pageNumber="484">10/10 57</td>
<td id="E4145D9D951A0067DE78F95B8CB0F92D" box="[1071,1229,1675,1737]" gridcol="4" gridrow="15" pageId="11" pageNumber="484">11/12 67</td>
<td id="E4145D9D951A0067DEBCF95B8DEAF92D" box="[1259,1431,1675,1737]" gridcol="5" gridrow="15" pageId="11" pageNumber="484">11/11 58</td>
</tr>
<tr id="A7C534E1951A0067DAC8F90C8DEAF8FE" box="[159,1431,1756,1818]" gridrow="16" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8F90C896DF8FE" box="[159,272,1756,1818]" gridcol="0" gridrow="16" pageId="11" pageNumber="484">FI TO</th>
<td id="E4145D9D951A0067DB32F90C8A75F8FE" box="[357,520,1756,1818]" gridcol="1" gridrow="16" pageId="11" pageNumber="484">17/16 22/23</td>
<td id="E4145D9D951A0067D874F90C8B40F8FE" box="[547,829,1756,1818]" gridcol="2" gridrow="16" pageId="11" pageNumber="484">16/17 16/17 22/23 23/24</td>
<td id="E4145D9D951A0067D924F90C8C68F8FE" box="[883,1045,1756,1818]" gridcol="3" gridrow="16" pageId="11" pageNumber="484">15/15 21/21</td>
<td id="E4145D9D951A0067DE78F90C8CB0F8FE" box="[1071,1229,1756,1818]" gridcol="4" gridrow="16" pageId="11" pageNumber="484">17/17 22/23</td>
<td id="E4145D9D951A0067DEBCF90C8DEAF8FE" box="[1259,1431,1756,1818]" gridcol="5" gridrow="16" pageId="11" pageNumber="484">17/17 23/23</td>
</tr>
<tr id="A7C534E1951A0067DAC8F8FD8DEAF88F" box="[159,1431,1837,1899]" gridrow="17" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8F8FD896DF88F" box="[159,272,1837,1899]" gridcol="0" gridrow="17" pageId="11" pageNumber="484">OS OS/HL</th>
<td id="E4145D9D951A0067DB32F8FD8A75F88F" box="[357,520,1837,1899]" gridcol="1" gridrow="17" pageId="11" pageNumber="484">9.5 0.30</td>
<td id="E4145D9D951A0067D874F8FD8B40F88F" box="[547,829,1837,1899]" gridcol="2" gridrow="17" pageId="11" pageNumber="484">7.3 5.6 0.27 0.19</td>
<td id="E4145D9D951A0067D924F8FD8C68F88F" box="[883,1045,1837,1899]" gridcol="3" gridrow="17" pageId="11" pageNumber="484">6.7 0.24</td>
<td id="E4145D9D951A0067DE78F8FD8CB0F88F" box="[1071,1229,1837,1899]" gridcol="4" gridrow="17" pageId="11" pageNumber="484">2.6 0.13</td>
<td id="E4145D9D951A0067DEBCF8FD8DEAF88F" box="[1259,1431,1837,1899]" gridcol="5" gridrow="17" pageId="11" pageNumber="484">5.1 0.22</td>
</tr>
<tr id="A7C534E1951A0067DAC8F8AE8DEAF858" box="[159,1431,1918,1980]" gridrow="18" pageId="11" pageNumber="484">
<th id="E4145D9D951A0067DAC8F8AE896DF858" box="[159,272,1918,1980]" gridcol="0" gridrow="18" pageId="11" pageNumber="484">NSSOS CS</th>
<td id="E4145D9D951A0067DB32F8AE8A75F858" box="[357,520,1918,1980]" gridcol="1" gridrow="18" pageId="11" pageNumber="484">5/5 14/14</td>
<td id="E4145D9D951A0067D874F8AE8B40F858" box="[547,829,1918,1980]" gridcol="2" gridrow="18" pageId="11" pageNumber="484">4/4 4/5 13/13 11/12</td>
<td id="E4145D9D951A0067D924F8AE8C68F858" box="[883,1045,1918,1980]" gridcol="3" gridrow="18" pageId="11" pageNumber="484">4/4 10/11</td>
<td id="E4145D9D951A0067DE78F8AE8CB0F858" box="[1071,1229,1918,1980]" gridcol="4" gridrow="18" pageId="11" pageNumber="484">4/5 12/13</td>
<td id="E4145D9D951A0067DEBCF8AE8DEAF858" box="[1259,1431,1918,1980]" gridcol="5" gridrow="18" pageId="11" pageNumber="484">5/5 12/12</td>
</tr>
</table>
</paragraph>
<paragraph id="194A36A3951AFF93DE37F81F8DEAF801" blockId="11.[1120,1431,1999,2021]" box="[1120,1431,1999,2021]" pageId="11" pageNumber="484">
<emphasis id="2B81EAB1951AFF93DE37F81F8DEAF801" box="[1120,1431,1999,2021]" italics="true" pageId="11" pageNumber="484">......continued on the next page</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>