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<mods:title id="97420CDB77AD1D4FA49FB18DD0BC3EEF">Trilobite Assemblage Of Calceola - Bearing Beds In Acanthopyge Limestone (Choteč Formation, Middle Devonian, Eifelian, Prague Basin, The Czech Republic)</mods:title>
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<paragraph id="1938D811FFBCFFFEFEC18417C76AFE86" blockId="4.[366,529,311,336]" box="[366,529,311,336]" pageId="4" pageNumber="83">
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<taxonomicName id="DE87A392FFBCFFFEFEC18417C4A6FE99" ID-CoL="63SY7" authorityName="PUSCH" authorityYear="1833" box="[366,477,311,335]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="2BF30403FFBCFFFEFEC18417C4A6FE99" bold="true" box="[366,477,311,335]" italics="true" pageId="4" pageNumber="83">Scutellum</emphasis>
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? sp.
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<figureCitation id="81BCC494FFBCFFFEFEDA8478C770FEA7" box="[373,523,344,369]" captionStart="Text-fig" captionStartId="6.[129,210,1931,1952]" captionText="Text-fig. 4. Trilobite assemblage at Zadní Kobyla small quarry outcrop (Calceola-bearing locality). ac Longiproetus? sp., a, b poorly preserved pygidium, CGS MM 570, dorsal view, a internal mould, b counterpart of the same specimen, c incomplete free cheek, questionably affiliated to Longiproetus? sp., CGS MM 571, internal mould, dorsal view. d Scabriscutellum sp., fragment of pygidium, internal mould, CGS MM 572, dorsal view. eg Scutellum sp., e fragment of pygidium, internal mould, CGS MM 574, lateral view, fg fragment of pygidium, CGS MM 573, dorsal view, f internal mould, g negative counterpart. h Kettneraspis sp., incomplete free cheek, negative counterpart, CGS MM 576." pageId="4" pageNumber="83">Text-fig. 4eg</figureCitation>
</paragraph>
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<paragraph id="1938D811FFBCFFFEFF0984A9C40AFE16" blockId="4.[128,768,392,449]" pageId="4" pageNumber="83">
M a t e r i a l.
<figureCitation id="81BCC494FFBCFFFEFE9184A8C430FE76" box="[318,331,392,416]" captionStart="Text-fig" captionStartId="1.[108,189,1960,1981]" captionTargetBox="[109,745,1481,1947]" captionTargetId="figure-453@1.[110,745,1481,1946]" captionTargetPageId="1" captionText="Text-fig. 2. General geological map of the Emsian (Suchomasty Limestone) and Eifelian (Acanthopyge Limestone) rocks at Zadní Kobyla ridge with marked Calceola- and trilobite-bearing locality. Modified after Svoboda and Prantl (1949) and Mergl (2014)." figureDoi="http://doi.org/10.5281/zenodo.5383375" httpUri="https://zenodo.org/record/5383375/files/figure.png" pageId="4" pageNumber="83">2</figureCitation>
fragments of pygidia, internal moulds and their counterparts.
</paragraph>
<paragraph id="1938D811FFBCFFFEFF0984F9C7CBFB09" blockId="4.[128,768,473,1248]" pageId="4" pageNumber="83">
R e m a r k s. Among trilobite remains from the locality, apart from ubiquitous phacopids and very fragmentary but common harpetids, remains of scutelluid trilobites are distinctive. They do not enable determination at the species level, and barely at the generic level.Despite this, it is apparent that they do not represent remains of
<emphasis id="2BF30403FFBCFFFEFDB4875CC7D4FD42" box="[539,687,636,660]" italics="true" pageId="4" pageNumber="83">Thysanopeltis</emphasis>
<bibRefCitation id="7D16A5E0FFBCFFFEFD17875CC448FD63" author="Hawle, I. &amp; Corda, A. J. C." pageId="4" pageNumber="83" pagination="117 - 292" refId="ref7970" refString="Hawle, I., Corda, A. J. C. (1847): Prodrom einer Monogra- phie der bohmischen Trilobiten. - Abhandlungen der ko- niglichen bohmischen Gesellschaft der Wissenschaften, Prag, 5 (5): 117 - 292." type="journal article" year="1847">HAWLE et CORDA, 1847</bibRefCitation>
, or
<taxonomicName id="DE87A392FFBCFFFEFEC787BDC5CEFD03" authority="RICHTER et RICHTER, 1956" authorityName="RICHTER et RICHTER" authorityYear="1956" class="Trilobita" family="Styginidae" genus="Scabriscutellum" kingdom="Animalia" order="Corynexochida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="2BF30403FFBCFFFEFEC787BDC76FFD63" box="[360,532,669,693]" italics="true" pageId="4" pageNumber="83">Scabriscutellum</emphasis>
<bibRefCitation id="7D16A5E0FFBCFFFEFD8E87BDC5CEFD03" author="Richter, R. &amp; Richter, M." pageId="4" pageNumber="83" pagination="79 - 124" refId="ref9311" refString="Richter, R., Richter, M. (1956): Grundlagen fur die Beurtei- lung und Einteilung der Scutelluidae (Tril.). - Sencken- bergiana lethaea, 37: 79 - 124." type="journal article" year="1956">RICHTER et RICHTER, 1956</bibRefCitation>
</taxonomicName>
(the only scutelluid genera mentioned in
<taxonomicName id="DE87A392FFBCFFFEFDDE879DC58AFD20" authority="Limestone" authorityName="Limestone (Chlupac" authorityYear="1983" class="Trilobita" family="Lichidae" genus="Acanthopyge" kingdom="Animalia" order="Lichida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">Acanthopyge Limestone</taxonomicName>
by Chlupáč 1983; see also discussion below). Pygidial fragments are characterised by wide, only moderately vaulted paired ribs, middle rib possessing coarse granulation (large granules, pustules), and very narrow inter-rib furrows. The axis is not preserved, with the exception of a narrow fragment figured in
<figureCitation id="81BCC494FFBCFFFEFDEA86A1C7B9FC4F" box="[581,706,897,921]" captionStart="Text-fig" captionStartId="6.[129,210,1931,1952]" captionText="Text-fig. 4. Trilobite assemblage at Zadní Kobyla small quarry outcrop (Calceola-bearing locality). ac Longiproetus? sp., a, b poorly preserved pygidium, CGS MM 570, dorsal view, a internal mould, b counterpart of the same specimen, c incomplete free cheek, questionably affiliated to Longiproetus? sp., CGS MM 571, internal mould, dorsal view. d Scabriscutellum sp., fragment of pygidium, internal mould, CGS MM 572, dorsal view. eg Scutellum sp., e fragment of pygidium, internal mould, CGS MM 574, lateral view, fg fragment of pygidium, CGS MM 573, dorsal view, f internal mould, g negative counterpart. h Kettneraspis sp., incomplete free cheek, negative counterpart, CGS MM 576." pageId="4" pageNumber="83">Text-fig. 4e</figureCitation>
; also, the pygidial outline is barely determinable, possibly semielliptical. Because of insufficient preservation, affiliation of all these samples to the
<taxonomicName id="DE87A392FFBCFFFEFE3F86C3C482FC2D" authorityName="PUSCH" authorityYear="1833" box="[400,505,995,1019]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="2BF30403FFBCFFFEFE3F86C3C482FC2D" box="[400,505,995,1019]" italics="true" pageId="4" pageNumber="83">Scutellum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30C0B978FFBCFFFEFDAB86C3C70AFC2D" box="[516,625,995,1019]" pageId="4" pageNumber="83" sensu="lato">sensu lato</taxonomicNameLabel>
seems to be the most parsimonious solution. Occurrence of
<taxonomicName id="DE87A392FFBCFFFEFD398124C47EFBEA" authority="PUSCH, 1833" authorityName="PUSCH" authorityYear="1833" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="2BF30403FFBCFFFEFD398124C784FBCA" box="[662,767,1028,1052]" italics="true" pageId="4" pageNumber="83">Scutellum</emphasis>
<bibRefCitation id="7D16A5E0FFBCFFFEFF2E8105C47EFBEA" author="Pusch, G. G." box="[129,261,1060,1084]" pageId="4" pageNumber="83" refId="ref9194" refString="Pusch, G. G. (1833): Geognostische Beschreibung von Polen, so wie der ubregen Nordkarpathen-Lander. Part 1. - J. G. Cotta, Stuttgart, Tubingen, 338 pp." type="book" year="1833">PUSCH, 1833</bibRefCitation>
</taxonomicName>
and allied taxa (for example,
<taxonomicName id="DE87A392FFBCFFFEFDE18104C4CBFB8B" authority="BASSE, KOCH et LEMKE, 2016" authorityName="BASSE, KOCH et LEMKE" authorityYear="2016" class="Trilobita" family="Odontopleuridae" genus="Torleyiscutellum" kingdom="Animalia" order="Lichida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="2BF30403FFBCFFFEFDE18104C784FBEA" box="[590,767,1060,1084]" italics="true" pageId="4" pageNumber="83">Torleyiscutellum</emphasis>
<bibRefCitation id="7D16A5E0FFBCFFFEFF2E8165C4CBFB8B" author="Basse, M. &amp; Koch, L. &amp; Lemke, U." box="[129,432,1093,1117]" pageId="4" pageNumber="83" pagination="51 - 93" refId="ref6646" refString="Basse, M., Koch, L., Lemke, U. (2016): Torleyiscutellum herwigorum n. gen., n. sp. (Trilobita) from the Upper Honsel Beds of the northwestern Sauerland (Lower Givetian, Rhenohercynian Zone), with a contribution to scutelluid systematic. - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 281: 51 - 93. https: // doi. org / 10.1127 / njgpa / 2016 / 0587" type="journal article" year="2016">BASSE, KOCH et LEMKE, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="DE87A392FFBCFFFEFE168165C7A8FB8B" authority="DE KONINCK, 1841" authorityName="DE KONINCK" authorityYear="1841" box="[441,723,1093,1117]" class="Trilobita" family="Styginidae" genus="Goldius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="2BF30403FFBCFFFEFE168165C776FB8B" box="[441,525,1093,1117]" italics="true" pageId="4" pageNumber="83">Goldius</emphasis>
<subScript id="8503DA54FFBCFFFEFDBB816BC756FB8D" attach="right" box="[532,557,1099,1115]" fontSize="7" pageId="4" pageNumber="83">DE</subScript>
<bibRefCitation id="7D16A5E0FFBCFFFEFD9D8165C7A8FB8B" author="Koninck, L. - G. de" box="[562,723,1093,1117]" pageId="4" pageNumber="83" pagination="1 - 19" refId="ref8247" refString="Koninck, L. - G. de (1841): Memoire sur les crustaces fos- siles de Belgique. - Nouveaux Memoires de l'Academie royale des Sciences et belles-Lettres de Bruxelles, 14: 1 - 19." type="journal article" year="1841">KONINCK, 1841</bibRefCitation>
</taxonomicName>
; see also
<bibRefCitation id="7D16A5E0FFBCFFFEFF1D8146C402FBA8" author="Weiner, T. &amp; Weinerova, H. &amp; Viktoryn, T." box="[178,377,1126,1150]" pageId="4" pageNumber="83" pagination="121 - 125" refId="ref10384" refString="Weiner, T., Weinerova, H., Viktoryn, T. (2018): A new find of scutelluid trilobite pygidium from the upper Frasnian of the Sumbera area near Brno-Lisen (Moravian Karst, Czech Republic). - Geoscience Research Reports, 51 (2): 121 - 125." type="journal article" year="2018">Weiner et al. 2018</bibRefCitation>
) is characteristic for the Middle and Upper Devonian of Laurussia (current
<collectingCountry id="61909881FFBCFFFEFD9481A7C7FDFB48" box="[571,646,1159,1182]" name="France" pageId="4" pageNumber="83">France</collectingCountry>
,
<collectingCountry id="61909881FFBCFFFEFD3881A6C780FB48" box="[663,763,1158,1182]" name="Germany" pageId="4" pageNumber="83">Germany</collectingCountry>
,
<collectingCountry id="61909881FFBCFFFEFF2F8187C5A7FB69" box="[128,220,1191,1215]" name="Belgium" pageId="4" pageNumber="83">Belgium</collectingCountry>
or
<collectingCountry id="61909881FFBCFFFEFEAF8187C435FB69" box="[256,334,1191,1215]" name="Poland" pageId="4" pageNumber="83">Poland</collectingCountry>
), see, for example,
<bibRefCitation id="7D16A5E0FFBCFFFEFDB28187C7B0FB69" author="Archinal, A." box="[541,715,1191,1215]" pageId="4" pageNumber="83" pagination="291 - 324" refId="ref6333" refString="Archinal, A. (1994): Zur Morphologie, Systematik, Strati- graphie und Okologie der Gattung Scutellum (Trilobita, Devon). - Senckenbergiana lethaea, 74 (1 - 2): 291 - 324." type="journal article" year="1994">Archinal (1994)</bibRefCitation>
,
<bibRefCitation id="7D16A5E0FFBCFFFEFD798187C4D4FB36" author="Van Viersen, A. P. &amp; Prescher, H." pageId="4" pageNumber="83" pagination="55 - 61" refId="ref10068" refString="Van Viersen, A. P., Prescher, H. (2011): Two new species of scutelluid trilobites formerly known as Scutellum costatum from Frasnian bioherms in Belgium. - Bulletin de l'Institut royale des sciences natureles de Belgique, Sciences de la Terre, 81: 55 - 61." type="journal article" year="2011">Van Viersen and Prescher (2011)</bibRefCitation>
and
<bibRefCitation id="7D16A5E0FFBCFFFEFE4D81E8C7D7FB09" author="Basse, M. &amp; Koch, L. &amp; Lemke, U." box="[482,684,1223,1247]" pageId="4" pageNumber="83" pagination="51 - 93" refId="ref6646" refString="Basse, M., Koch, L., Lemke, U. (2016): Torleyiscutellum herwigorum n. gen., n. sp. (Trilobita) from the Upper Honsel Beds of the northwestern Sauerland (Lower Givetian, Rhenohercynian Zone), with a contribution to scutelluid systematic. - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 281: 51 - 93. https: // doi. org / 10.1127 / njgpa / 2016 / 0587" type="journal article" year="2016">Basse et al. (2016)</bibRefCitation>
.
</paragraph>
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