102 lines
14 KiB
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102 lines
14 KiB
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<document id="8F43093B51FDDE2E0082ED220017ECB2" ID-CLB-Dataset="34877" ID-DOI="http://dx.doi.org/10.3897/zookeys.276.4786" ID-GBIF-Dataset="c8f51a00-c823-4313-b379-24dd38b96021" ID-PMC="PMC3677342" ID-Pensoft-Pub="1313-2970-276-55" ID-PubMed="23794816" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-276-55" ModsDocOrigin="ZooKeys 276" ModsDocTitle="Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)" checkinTime="1451247576187" checkinUser="pensoft" docAuthor="Krikken, Jan" docDate="2013" docId="912D707A247E0D0BB9D6BE3BD9115EDF" docLanguage="en" docName="ZooKeys 276: 55-66" docOrigin="ZooKeys 276" docSource="http://dx.doi.org/10.3897/zookeys.276.4786" docTitle="Bolboceras orissicum Krikken, 2013, sp. n." docType="treatment" docVersion="6" lastPageNumber="64" masterDocId="FFB2ED31FFDDFFACDF09FF97FFA5FFD4" masterDocTitle="Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)" masterLastPageNumber="66" masterPageNumber="55" pageNumber="62" updateTime="1732833068396" updateUser="ExternalLinkService">
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<mods:mods id="B259FE246F0642384E5969829745CEEE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E16A04BC609D6A7D1C49A408E05B6749">
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<mods:title id="91F1FB49365A9BEA470B7BB572D9156F">Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)</mods:title>
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<mods:name id="DC07671E957AF548332E75D6C3549024" type="personal">
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<mods:roleTerm id="6A5C687A7E52DD9F4E20EB4BC2FB15AF">Author</mods:roleTerm>
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<mods:namePart id="AEC2E4CB5DEEB7550C83994516E3D08F">Krikken, Jan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="84F9F7A7E52C686087A746FA5C186788">text</mods:typeOfResource>
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<mods:title id="8DA28CD86D6D20D7970CF64B4CD168B7">ZooKeys</mods:title>
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<mods:part id="6837458F2C075DCF78682AE824C8FF39">
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<mods:date id="1E320F07B57C56C5C54E326569A294B9">2013</mods:date>
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<mods:detail id="8C547B93B0D01FAE24916EB5102F8AC0" type="volume">
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<mods:number id="BBA35462C99BC8A89F0D9AE21F4DE03D">276</mods:number>
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</mods:detail>
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<mods:extent id="5989514F29AA27F5F156CC1790D23F86" unit="page">
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<mods:start id="218D0C989281D1A35FBF1D93620FA1C5">55</mods:start>
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<mods:end id="F645E33ED8CAA3C5B398BAD074154258">66</mods:end>
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<mods:location id="AB81020C01E48F4850AA3EA6FE15B1E1">
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<mods:url id="0FD016A24A256ED7624EC93E7E8A55EE">http://dx.doi.org/10.3897/zookeys.276.4786</mods:url>
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</mods:location>
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<mods:classification id="97E5FAFFE883CE43F7A5C2044A0B8791">journal article</mods:classification>
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<mods:identifier id="2A7F395F7AAD7E7286CAF61210D8D03D" type="DOI">http://dx.doi.org/10.3897/zookeys.276.4786</mods:identifier>
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<mods:identifier id="EFCD756DA2D7B2D3B718A69FF6A379C4" type="Pensoft-Pub">1313-2970-276-55</mods:identifier>
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</mods:mods>
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<treatment id="912D707A247E0D0BB9D6BE3BD9115EDF" ID-GBIF-Taxon="152041485" LSID="urn:lsid:zoobank.org:act:64EA91B5-7AB9-4EE0-BAF7-33C24FCB6BE3" httpUri="http://treatment.plazi.org/id/912D707A247E0D0BB9D6BE3BD9115EDF" lastPageId="9" lastPageNumber="64" pageId="7" pageNumber="62" scope_family="Geotrupidae" scope_order="Coleoptera" scope_subFamily="Bolboceratinae">
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<subSubSection id="36047889E61BA8F572AE8CAF614F2125" pageId="7" pageNumber="62" type="nomenclature">
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<paragraph id="E0D3B37CFECA1D5BE5767F13206263B8" pageId="7" pageNumber="62">
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<taxonomicName id="358F69E125B01F5BB63AC11F0FB05FE1" ID-CoL="MBCC" LSID="urn:lsid:zoobank.org:act:64EA91B5-7AB9-4EE0-BAF7-33C24FCB6BE3" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolboceras orissicum" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="orissicum">Bolboceras orissicum</taxonomicName>
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<taxonomicNameLabel id="A4449568CAC881A4831C725E5870C0AC" pageId="7" pageNumber="62">sp. n.</taxonomicNameLabel>
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Figures 7-812, 1422-23
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</paragraph>
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</subSubSection>
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<subSubSection id="0AF07F2FA13140B58612B6F436EA3559" pageId="7" pageNumber="62" type="material examined">
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<paragraph id="84FD73A24A56B322540CC028062F9D76" pageId="7" pageNumber="62">Material examined.</paragraph>
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<paragraph id="2F1FFA74A48DBB67927EBEBB084EE875" pageId="7" pageNumber="62">Holotype male (MNHN, ex Boucomont coll.) from India: [Orissa:] Ganjam: Surada [also spelled Sorada], H. Donckier. Male and female paratypes (SMF) from [India: Jharkhand:] "Burju \ Bengal".</paragraph>
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</subSubSection>
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<subSubSection id="E77D269037CB3B5E52B55FBAD1314A92" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62" type="diagnosis">
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<paragraph id="98CD7FD174366AF7E62F43022806E2F5" pageId="7" pageNumber="62">Diagnosis.</paragraph>
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<paragraph id="C5241597F7A7910DB526611B5A1F726C" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">
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Saddle separating discomedian and discolateral concavities on pronotum very short, broad, not lowered in relation to surrounding discal surface, discoparamedian tubercles distinctly separated by anterior part of discomedian concavity between them; discolateral concavity quasi-extending onto shallowly concave anterolateral corner; posterior edge of discolateral concavity well defined,
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<normalizedToken id="C768FCA3A221DCBFD0198AEB1945417C" originalValue="“swollen”">"swollen"</normalizedToken>
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near saddle with discomedian concavity (outline in dorsal view sinuate). Posterior edge of discomedian concavity virtually rounded. Interocular transverse ridge situated between
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<pageBreakToken id="A57A972B7F28CB3F0C4DDD6644446FB5" pageId="8" pageNumber="63" start="start">posterior</pageBreakToken>
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part of eyes, low, not reaching paraocular ridge on either side. Anterolateral corners of clypeus strongly protuberant (intervening ridge concave in axial view). Scutellum usually abundantly punctate. Aedeagus with narrow, acuminate parameres, median apparatus with pair of strongly sclerotized lateral stalks, their tip very broadly recurved-hooked. Colour uniformly light-brown. Body length roughly 12.5-13.5 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="72E68EF56960D1EF16C101A2977C065F" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63" type="description">
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<paragraph id="AE841D8AEEF0E54379325FC21E47CD3E" pageId="8" pageNumber="63">Description</paragraph>
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<paragraph id="7D19D19175CF5049433E571607504171" pageId="8" pageNumber="63">(holotype, male). Body length ca 13 mm. Colour uniformly light-brown, shiny, certain parts sericeous.</paragraph>
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<paragraph id="6B697972A4902F252BFC79CD7A8D5301" pageId="8" pageNumber="63">Labrum with emarginate anteromedian border, transverse ridge on rugulate upper surface fine, distinct. Clypeal surface with supra-anterolateral angles distinctly raised, dentate, intervening ridge concave in axial view, straight in full-face view; lateral perimarginal ridges distinct, strongly evenly curved, genal angle distinct. Clypeus densely punctate, remainder of head surface abundantly, evenly, distinctly punctate, primary punctures interspersed with fine secondary punctation being denser behind interocular ridge. Anterior edge of eye canthus raised to slight anterolateral angle, thence arcuate along edge of distal lobe; surface coarsely rugulate; paraocular ridge distinct, fine issuing from genal angle, virtually straight, extending posteriorly along eye. Transverse interocular elevation between posterior part of eyes, long, low, not reaching paraocular ridges, lateral slope slight, crest fine, unmodified, lateral angle obtuse on either end (axial view). Much of head surface subsericeous.</paragraph>
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<paragraph id="303B0BE01A9D655C4EBE8FC4FDD6900D" pageId="8" pageNumber="63">Pronotum anteromedially steeply declivous, outline of marginate (raised) anterior border slightly convex (dorsal view); discoparamedian and discolateral protrusions on pronotum distinctly protuberant, subrectangular (lateral view), their tip rounded; discomedian concavity very distinct, evenly concave, broad, anterior part situated between discoparamedian tubercles (not continuing over anterior declivity), posterior edge broadly rounded (dorsal view); discolateral concavity deep, posterior edge sinuate; anteriorly, at bottom, delimited by crowdedly, coarsely punctate patch behind eyes; saddle from discoparamedian tubercle to posterior disc very short, not lowered; bottom of discal concavities and anterior declivity subsericeous; basomedian surface glossy, with abundantly, finely punctate midline impression; anterolateral angle of marginal pronotal ridge ca 100˚ (full-face view). Pronotal surface with double punctation, primary punctation (size variable) laterally generally abundant, denser on anterior declivity; secondary punctation sparse, minute. Pronotal base broadly marginate, lined with numerous punctures. Scutellum with abundant, double punctation.</paragraph>
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<paragraph id="4119043AF26CF125DD0C33B16DF3BB90" pageId="8" pageNumber="63">Intercoxal anterior lobe of metasternum simply truncate in front.</paragraph>
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<paragraph id="4EBECFB672A508E17ACC4799911DDCBA" pageId="8" pageNumber="63">Elytra with discal striae shallowly impressed, finely punctate; punctures separated by 2-3 puncture diameters, slightly crenulating interstriae. Elytral interstriae (on cross-section) very slightly convex, vaguely, sparsely, micropunctate.</paragraph>
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<paragraph id="FE3C286A9336AF95399CBDA587E913D9" pageId="8" pageNumber="63">Protibia with 6 external denticles; apex unmodified, with robust, complanate, slightly tapering spur. Outer side of meso- and metatibiae with bilobate apical and one complete anteapical fossorial elevation (their crest fringed with fine spines).</paragraph>
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<paragraph id="73D17B243B65AD85686B723EA1BAC1DB" pageId="8" pageNumber="63">Aedeagus, Fig. 14; sclerotized lateral stalks broad with recurved, hooked tip, parameral sheaths narrow, acuminate.</paragraph>
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<paragraph id="847AB5165DD219D8AC3D148A5994CF5F" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
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Measurements of body parts. Median length of head (full-face, excluding labrum and mandibles) 2.6 mm, width (including eyes) 4.2 mm. Median length of pronotum
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<pageBreakToken id="A32C16545767E25D17235A3180702061" pageId="9" pageNumber="64" start="start">(</pageBreakToken>
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dorsal) 4.6 mm, maximum width 8.2 mm. Median length of scutellum 1.1 mm, maximum width 1.4 mm. Sutural length of elytra (dorsal) 3.9 mm, maximum width combined 8.3 mm. Width genital capsule 1.35 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="DECBE6313647FE58326CFB82D9FD8173" pageId="9" pageNumber="64" type="variation">
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<paragraph id="85D5265ECDA73968310634E0B0D79B8D" pageId="9" pageNumber="64">Variation.</paragraph>
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<paragraph id="44BAA1EFBBA4C383BF14CA8C2737BB71" pageId="9" pageNumber="64">Variation slight, but beware of possibly deceptive polymorphism (may be obvious in larger series).</paragraph>
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</subSubSection>
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<subSubSection id="D3F1F526C602F30223BACBC0490A79A5" pageId="9" pageNumber="64" type="sexual dimorphism">
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<paragraph id="8AB66B1A32ECE5C4AA1C921C248D8FF9" pageId="9" pageNumber="64">Sexual dimorphism.</paragraph>
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<paragraph id="6B88CE33029CEE62868ABA6B6E4E69AE" pageId="9" pageNumber="64">No obvious sexual dimorphism.</paragraph>
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</subSubSection>
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<subSubSection id="7F55886F2F00FA7CACEF13B775AB3B25" pageId="9" pageNumber="64" type="distribution">
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<paragraph id="9454B08C564D5F124D66C1B76EBEAE29" pageId="9" pageNumber="64">Distribution.</paragraph>
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<paragraph id="B5A3E0F57589B7FDEC080F8B56C46FCA" pageId="9" pageNumber="64">Northeast India, apparently South of the Ganges.</paragraph>
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</subSubSection>
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<subSubSection id="F087F5909809C76B48F4AFD1F7F32CCB" pageId="9" pageNumber="64" type="comment">
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<paragraph id="3C6C0D59A0E082848D7C670AD1C3C9AC" pageId="9" pageNumber="64">Comment.</paragraph>
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<paragraph id="9A956CD2934AB80D73985D58EAB1E338" pageId="9" pageNumber="64">The place called Burju, origin of the paratypes, is mentioned in the list of "post-office pincodes" in the Ranchi District of Jharkhand (India), and at the time of the collection of the SMF specimens there was a German mission school in the area, with staff sending specimens to German entomologists (around 1890-1910).</paragraph>
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</subSubSection>
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<subSubSection id="C2811E48FEFCFE325C809487D06FE4BB" pageId="9" pageNumber="64" type="etymology">
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<paragraph id="F0026CEF757CD4598B7E3F754B8E40BA" pageId="9" pageNumber="64">Etymology.</paragraph>
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<paragraph id="A80997C92D59260AF9F8F7C98B97994D" pageId="9" pageNumber="64">Named after the type region.</paragraph>
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<caption id="8928596C38669BEE79A40EF8D09DE500" pageId="9" pageNumber="64">
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<paragraph id="7CC42195F1E167E419FE77ABD73EDD56" pageId="9" pageNumber="64">
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Figures 16-23. Head, full-face view, and pronotum, dorsal view, of
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<taxonomicName id="DE021D908992792005E4046D8C1A43D4" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolboceras" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="genus">Bolboceras</taxonomicName>
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: 16-19
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<taxonomicName id="6B5C6C74D0D9456C4E6FF8B5CBC0CAA6" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolboceras inaequale" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="inaequale">Bolboceras inaequale</taxonomicName>
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(16-17 major specimen from Kolkata 18-19 minor specimen from Dhanjuri) 20-21duplicatum,holotype 22-23orissicum,holotype. Scale lines 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |