148 lines
14 KiB
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148 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="906E0FB8219F6C5BC72D1738D4C60A34" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha elevata Likhitrakarn, Golovatch & Panha, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="93" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="89" updateTime="1668152297771" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>131</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031115" LSID="urn:lsid:zoobank.org:act:DDACEC9B-1E7D-483E-9B4D-F65E619D2E76" httpUri="http://treatment.plazi.org/id/906E0FB8219F6C5BC72D1738D4C60A34" lastPageId="92" lastPageNumber="93" pageId="88" pageNumber="89">
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<subSubSection pageId="88" pageNumber="89" type="nomenclature">
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<paragraph pageId="88" pageNumber="89">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:DDACEC9B-1E7D-483E-9B4D-F65E619D2E76" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha elevata" order="Polydesmida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="elevata">Orthomorpha elevata</taxonomicName>
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<taxonomicNameLabel pageId="88" pageNumber="89">sp. n.</taxonomicNameLabel>
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Figs 100102
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</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="holotype">
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<paragraph pageId="88" pageNumber="89">Holotype.</paragraph>
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<paragraph pageId="88" pageNumber="89">
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♂ (CUMZ), Malaysia, Perak State, Bondig, Air Banum, ca 500 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="5.6369443">5°38'13"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.71139">101°42'41"E</geoCoordinate>
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, 27.05.2011, leg. S. Panha.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="paratypes">
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<paragraph pageId="88" pageNumber="89">Paratypes.</paragraph>
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<paragraph pageId="88" pageNumber="89">
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2 ♀ (CUMZ), same data, together with holotype. 1 ♀ (CUMZ), Malaysia, Johor State, Sungai Bantang, ca 100 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.3305554">2°19'50"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.1625">103°09'45"E</geoCoordinate>
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, 21.05.2011, leg. R. Chanabun. 1 ♀ (CUMZ), same State, Sungai Bantang, ca 100 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.3305554">2°19'50"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.1625">103°09'45"E</geoCoordinate>
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, 21.05.2011, leg. R. Chanabun. 2 ♀ (CUMZ), Kelantan State, near Gua Pulai, ca 120 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="5.243334">4°74'36"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.94222">101°56'32"E</geoCoordinate>
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, 26.05.2011, leg. R. Chanabun. 2 ♀ (CUMZ), Kelantan State, near Gua Matu Madu, ca. 130 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.8369446">4°50'13"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.94889">101°56'56"E</geoCoordinate>
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, 26.05.2011, leg. R. Chanabun.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="name">
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<paragraph pageId="88" pageNumber="89">Name.</paragraph>
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<paragraph pageId="88" pageNumber="89">To emphasize the mostly strongly elevated paraterga.</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="diagnosis">
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<paragraph pageId="88" pageNumber="89">Diagnosis.</paragraph>
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<paragraph pageId="88" pageNumber="89">Differs by mostly strongly elevated paraterga, coupled with two small sternal cones between ♂ coxae 4 (see also Key below).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="92" lastPageNumber="93" pageId="88" pageNumber="89" type="description">
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<paragraph pageId="88" pageNumber="89">Description.</paragraph>
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<paragraph pageId="88" pageNumber="89">Length 34 (holotype) or 31-38 mm (♀), width of midbody pro- and metazona 2.8 and 4.3 mm (holotype), 2.9-3.5 and 4.3-5.0 mm (♀), respectively.</paragraph>
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<paragraph lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
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Coloration of live material (Fig. 100A) blackish to blackish-brown with contrasting pinkish paraterga legs, and tip of epiproct; coloration of alcohol material after long-term preservation blackish-brown with contrasting light yellowish calluses; two paramedian spots divided by a broad, blackish, axial stripe on prozona; lateral parts of
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<pageBreakToken pageId="89" pageNumber="90" start="start">paraterga</pageBreakToken>
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near calluses, tip of epiproct, legs and venter light yellow-brown, legs slightly infuscate distally; antennae brown, distal part of antennomere 5, distal 2/3 of antennomere 6 and entire antennomere 7 blackish-brown (Fig. 100B-H).
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</paragraph>
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<paragraph lastPageId="91" lastPageNumber="92" pageId="89" pageNumber="90">
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Clypeolabral region densely setose, vertex bare, epicranial suture distinct. Antennae short, slightly clavate (antennomere 6 broadest), extending behind body segment 3 (♂) or 2 (♀) dorsally. Head in width <collum <segments 3 and 4 <2 <5-17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of medium-sized setae, pattern untraceable; paraterga slightly declivous, subtriangular, with a faint lateral incision near midway, evidently discontinuing
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<normalizedToken originalValue="dorsum’s">dorsum's</normalizedToken>
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convexity,
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<pageBreakToken pageId="90" pageNumber="91" start="start">caudal</pageBreakToken>
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corner pointed, about level to rear tergal margin (Fig. 100B & C). Tegument rather poorly shining, metaterga, prozona and most of metazona below paraterga finely shagreened; metaterga dull, leathery, roughly shagreened, their caudal margin evidently ribbed, with 3+3 or 4+4 short wrinkles; surface below paraterga rugulose, on segments 2-4(5) microgranulate. Postcollum metaterga with fully abraded setae, setation pattern untraceable even as insertion points. Axial line visible on collum and both on following pro- and metazona. Paraterga very strongly developed (Fig. 100A-H), especially well so in ♂, set at about 1/4 body height, postcollum ones upturned in lateral view, moderately enlarged on pore-bearing segments, thinner on poreless ones; shoulders broadly rounded and narrowly bordered, fused to callus; paraterga 2 and 19 (♂) or 19 (♀) lying below dorsum, paraterga 3 (♂) or 2-4 and 18 (♀) level to dorsum, remaining postcollum paraterga clearly above dorsum; caudal corner of postcollum paraterga pointed or narrowly rounded, always extending beyond rear tergal margin, best developed on segments 17 and 18 (Fig. 100F & G). Calluses delimited by a sulcus both dorsally and
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<pageBreakToken pageId="91" pageNumber="92" start="start">ventrally</pageBreakToken>
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, with an evident front denticle only on paraterga 2, lateral margin virtually unincised on following paraterga, except for a small sinuosity in front of ozopore on pore-bearing segments. Posterior edge of paraterga strongly concave, especially strongly so on segments 18 and 19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 4-18, narrow, superficial, not reaching bases of paraterga. Stricture between pro- and metazona narrow, rather deep, beaded at bottom down to base of paraterga (Fig. 100B-F & H). Pleurosternal carinae complete crests with a caudal tooth on segments 2-4, thereafter increasingly strongly reduced and remaining visible only as a front bulge and a caudal tooth until segment 17 (♂) or 13 (♀). Epiproct (Fig. 100F-H) conical, flattened dorsoventrally, with two small teeth directed ventrocaudally, emarginate or subtruncate at tip; pre-apical papillae small denticles lying very close to tip. Hypoproct (Fig. 100G) semi-circular, caudal margin with two strong and well-separated setiferous knobs.
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</paragraph>
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<paragraph pageId="91" pageNumber="92">Sterna sparsely setose, shining, without modifications; cross-impressions shallow; a paramedian pair of rather small, but evident tubercles in front of gonopod aperture; a small central lobe with a paramedian pair of evident, setose, apical cones between ♂ coxae 4 (Fig. 100I & J). Legs long and slender, slightly incrassate in ♂, midbody ones ca 1.3-1.4 (♂) or 1.1-1.2 times (♀) as long as body height, prefemora without modifications; ♂ tarsal brushes visible on legs 1-3, thereafter gradually thinning out.</paragraph>
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<paragraph pageId="92" pageNumber="93">
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<pageBreakToken pageId="92" pageNumber="93" start="start">Gonopods</pageBreakToken>
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(Figs 101 & 102) simple. Coxa long and slender, with several strong setae distodorsally. Prefemur densely setose, more than 2 times shorter than femorite +
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<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite slightly curved, slender, nearly not enlarged distad, with a
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<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus. Solenophore tip distinctly trifid, terminal prong being sharp and longest, middle denticle nearly as long as subterminal lobule; solenomere long and flagelliform.
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</paragraph>
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<caption pageId="92" pageNumber="93">
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<paragraph pageId="92" pageNumber="93">
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Figure 100.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha elevata" order="Polydesmida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="elevata">Orthomorpha elevata</taxonomicName>
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sp. n., ♂ holotype. A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption pageId="92" pageNumber="93">
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<paragraph pageId="92" pageNumber="93">
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Figure 101.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha elevata" order="Polydesmida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="elevata">Orthomorpha elevata</taxonomicName>
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sp. n., ♂ holotype. A, B right gonopod, mesal and lateral views, respectively C-F distal part of right gonopod, mesal, lateral, suboral and subcaudal views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="92" pageNumber="93">
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<paragraph pageId="92" pageNumber="93">
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Figure 102.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha elevata" order="Polydesmida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="elevata">Orthomorpha elevata</taxonomicName>
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sp. n., ♂ holotype. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="92" pageNumber="93" type="remark">
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<paragraph pageId="92" pageNumber="93">Remark.</paragraph>
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<paragraph pageId="92" pageNumber="93">This new species is rather widespread in northern Malaysia (Map 2).</paragraph>
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</subSubSection>
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</treatment>
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</document> |