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<document ID-CLB-Dataset="55905" ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.83916" ID-GBIF-Dataset="e036b25d-83e3-4247-9c3b-41ad2a84b72a" ID-PMC="PMC9043006" ID-Pensoft-Pub="1313-2970-1097-1" ID-Pensoft-UUID="6593D2730A295F46B63A738B12E43A22" ID-PubMed="35837583" ID-ZooBank="66F17436DF364FFDB2D3021F14D40D62" ModsDocID="1313-2970-1097-1" checkinTime="1650371395093" checkinUser="pensoft" docAuthor="Antic, Dragan, Vagalinski, Boyan, Stoev, Pavel &amp; Akkari, Nesrine" docDate="2022" docId="8FBCC310088A5EAE90DCB67E2AF03D7A" docLanguage="en" docName="ZooKeys 1097: 1-46" docOrigin="ZooKeys 1097" docPubDate="2022-04-19" docSource="http://dx.doi.org/10.3897/zookeys.1097.83916" docTitle="Balkanodesminus serbicus gen. nov. et 2022, sp. nov." docType="treatment" docUuid="2684D82A-21C8-4235-8B2C-39886C7440B1" docUuidSource="ZooBank" docVersion="6" id="6593D2730A295F46B63A738B12E43A22" lastPageNumber="1" masterDocId="6593D2730A295F46B63A738B12E43A22" masterDocTitle="A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="1" updateTime="1732687488807" updateUser="ExternalLinkService">
<mods:mods id="07C76F9B227F4FA9C8C5F11D956E10CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2D71204F64C912586ED39172BEBB68EF">
<mods:title id="FFFC8261573F48101AEAFB3C5BD7E6E4">A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species</mods:title>
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<mods:name id="ACC5037F593E10D55A0948D4210D5DA2" type="personal">
<mods:role id="B53A705F6D19E70C03CFE3EEC08FDB0E">
<mods:roleTerm id="E1678E57D3154779A3417598D3C3D8CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9F10A8CA7AD5064E6DEF522454E205B">Antic, Dragan</mods:namePart>
<mods:nameIdentifier id="C3A5F569BAE2BAF10BEF85C5C44BB3DD" type="ORCID">https://orcid.org/0000-0002-1231-4213</mods:nameIdentifier>
<mods:affiliation id="442A654E05A62FEC1A203B08AC5C4EA4">University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski trg 16, 11000 Belgrade, Serbia &amp; Serbian Biospeleological Society, Trg Dositeja Obradovica 2, 21000 Novi Sad, Serbia</mods:affiliation>
<mods:nameIdentifier id="8C7887B1F9A81F7D8DC20C52D9735B5F" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:name id="832A5C16670EF19CC04F381C11918FB2" type="personal">
<mods:role id="2DCA909BDAE49B34F80517B009CBB759">
<mods:roleTerm id="25A77C082850624388BDEC738F579B42">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9DCFD8945CBE16FD94A8018087A5B33">Vagalinski, Boyan</mods:namePart>
<mods:affiliation id="2D971E9EC6F229E17208894B548D02F0">Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name id="A6BEAFD702B83853726CA69E0D7F9DEE" type="personal">
<mods:role id="EE8FBD5964FED0A8ECAD69DD50728F08">
<mods:roleTerm id="1AD3A46D81DB3980AFEB5E047E757BC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93B6C76B40273F98FA642A7BD972AD38">Stoev, Pavel</mods:namePart>
<mods:nameIdentifier id="AF7259D2D80A5EE86692F65D7CBD6243" type="ORCID">https://orcid.org/0000-0002-5702-5677</mods:nameIdentifier>
<mods:affiliation id="4A4C826D7199EDCD2755F6E8F3F53227">National Museum of Natural History, Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd. 1, Sofia 1000, Bulgaria &amp; Pensoft Publishers, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name id="569A1DBCE6D5ACC45075A4619E204785" type="personal">
<mods:role id="5DCBBF178B852B647A2368FB1E1BEEF0">
<mods:roleTerm id="50B32B30E4C809B3874ABD9954EAAE30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D28D6AFCE95B3B08B7CB9081C56CDB81">Akkari, Nesrine</mods:namePart>
<mods:nameIdentifier id="5F6CC21ABAE015F97908823FCC76768C" type="ORCID">https://orcid.org/0000-0001-5019-4833</mods:nameIdentifier>
<mods:affiliation id="4C1C4847C7B92B28984CFD68CB7CD6DF">3</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A2A7A172EC3392E093EA6DEC3F279E20">text</mods:typeOfResource>
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<mods:titleInfo id="CC798CE1FCD3AB4526B548050AE44EA8">
<mods:title id="162AA75B4C0AFE0ADB00BC83AB09DC51">ZooKeys</mods:title>
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<mods:part id="9944C6E98CDEAE4232636CED08F6DD75">
<mods:date id="C803D26FF877AA2F762DDE59AB1EF0B1">2022</mods:date>
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<mods:number id="6EEC0B1EA82A497575571901991F5239">2022-04-19</mods:number>
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<mods:detail id="95CEBFA3AFC3AC9CDE5B3E47B9E69BE5" type="volume">
<mods:number id="2B73A758F0643871C0FF3AB889F2EB75">1097</mods:number>
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<mods:start id="AE08B67B50FAA71A0A38F661F4721351">1</mods:start>
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<mods:url id="ECAB1A369B676160E48A38CD30D46D85">http://dx.doi.org/10.3897/zookeys.1097.83916</mods:url>
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<mods:classification id="9E0DEB2F7A58DBF0E4394CB70A73E742">journal article</mods:classification>
<mods:identifier id="1409B09EC93E3869A7849071DEA699F8" type="DOI">http://dx.doi.org/10.3897/zookeys.1097.83916</mods:identifier>
<mods:identifier id="48892A5C03FAC24B857E56EE090256E6" type="Pensoft-Pub">1313-2970-1097-1</mods:identifier>
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<treatment id="8FBCC310088A5EAE90DCB67E2AF03D7A" ID-GBIF-Taxon="194660822" LSID="urn:lsid:zoobank.org:act:2684D82A-21C8-4235-8B2C-39886C7440B1" httpUri="http://treatment.plazi.org/id/8FBCC310088A5EAE90DCB67E2AF03D7A" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Diplopoda" scope_family="Trichopolydesmidae" scope_order="Polydesmida">
<subSubSection id="6990DFFE43EBB776A4E73EAB750270A8" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="6B934B916DDE34F513627FF3DC1EE842" pageId="0" pageNumber="1">
<taxonomicName id="DF3114C8CC6493DB76CFA0BDE56AF1C2" ID-CoL="8ZDS5" LSID="http://zoobank.org/2684D82A-21C8-4235-8B2C-39886C7440B1" authority="gen. nov. et" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus" status="sp. nov.">Balkanodesminus serbicus gen. nov. et</taxonomicName>
<taxonomicNameLabel id="CD60C8CE84C63F303BD32A42B52CE6CF" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A8F96DB80D05E3832B3A8A778AA57CA7" pageId="0" pageNumber="1" type="description">
<paragraph id="5B561C340AFEC350D94C2DAE0EBE4E2E" pageId="0" pageNumber="1">
<figureCitation id="804FF58CEBE954DC5863D05055737B8A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">Figs 10</figureCitation>
<figureCitation id="2F3A9B01255D42107A364E533164D9C9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">, 11</figureCitation>
<figureCitation id="DDD324D5254715A024A8AFA676F22982" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">, 12</figureCitation>
<figureCitation id="032CD53C8279EA260EE4D56A9AFD8237" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♂ left gonopod (NHMW MY 10263) A mesal view B anterior view C distal view D lateral view E postro-lateral view F posterior view. Abbreviations: a acropodite, bl basal lamellar lobe of solenophore, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, pf prefemorite, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore. Scale bars: 0.02 mm." figureDoi="10.3897/zookeys.1097.83916.figure13" httpUri="https://binary.pensoft.net/fig/674291" pageId="0" pageNumber="1">, 13</figureCitation>
<figureCitation id="4B226F57EE8955A1F1E65D53ED4C8A60" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">, 17J</figureCitation>
<figureCitation id="AA46DB38D7257661C01F2F1D38FEB5CA" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="900FCD1A849E8288AFF6CEAFC556C7F5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="34DCF8CB70BF7975345458EAD892ABC8" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="7672C669283C7C8D87790197A9AEBC3E" pageId="0" pageNumber="1">
Differs from
<taxonomicName id="57A57B4510A413B1AF635F66E4CC53C3" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
<emphasis id="92F631EE4835C32BDDC8A0E6A13CABF7" italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
</taxonomicName>
gen. nov. et sp. nov. and
<taxonomicName id="B8F9A07FF9313C64DC72CFCA6B10E28D" lsidName="B. dentatus" pageId="0" pageNumber="1" rank="species" species="dentatus">
<emphasis id="FA485FC06E3C4AA257E6A96754620028" italics="true" pageId="0" pageNumber="1">B. dentatus</emphasis>
</taxonomicName>
gen. nov., comb. nov. by the presence of longer (vs. shorter) metatergal setae and their smaller (vs. greater) number of rows, as well as by the presence of more simplified gonopods, with uniramous (vs. biramous) distal solenomeral process and smaller and smooth (vs. larger and denticulated) lamella of solenophore. From
<taxonomicName id="ACC35C6BC12012A9B99F57DE96CE1D61" lsidName="B. bulgaricus" pageId="0" pageNumber="1" rank="species" species="bulgaricus">
<emphasis id="A73611ED1F760493BEE75DC2D601E273" italics="true" pageId="0" pageNumber="1">B. bulgaricus</emphasis>
</taxonomicName>
gen. nov., comb. nov., with which it shares similar habitus and similar gonopods, it differs by the presence of smaller (vs. larger) lamella of solenophore, more robust and sigmoid (vs. slender, almost straight) distal projection of solenophore, distal projection with (vs. without) basal lobe, and more robust (vs. slenderer) solenomere, exceeding the distal solenomeral process by
<normalizedToken id="E18AF8B742FF2AC82414718795E03341" originalValue="¼">1/4</normalizedToken>
(vs.
<normalizedToken id="13C7E127DC2ED58A4313E9107858C9B7" originalValue="½">1/2</normalizedToken>
) of its length.
</paragraph>
</subSubSection>
<subSubSection id="51EB3606A174066523CE7F56B464F547" pageId="0" pageNumber="1" type="name">
<paragraph id="4880D50E1C5CD58C087D9EBEC44CA91B" pageId="0" pageNumber="1">Name.</paragraph>
<paragraph id="BE8DCA714D256DF3128888470371C75A" pageId="0" pageNumber="1">The specific name is an adjective derived from the type locality.</paragraph>
</subSubSection>
<subSubSection id="07C92C1D1FAF6F168306EF01DBB3507F" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="03FF0EA064E6F41B020CE98CF6F88C92" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="2BF66FCA9AEDBC5F54C650AF7F1867C3" pageId="0" pageNumber="1">
<materialsCitation id="E67567B0708E4CBAFDE5FF814C4D77CF" collectingDate="2017-10-29" collectorName="D. Antic." country="Serbia" county="Mt. Kalafat" location="Cerjanska Cave" municipality="Cerje" specimenCount="♂" stateProvince="Nis" typeStatus="Holotype">
<emphasis id="06283CDE6CD9578B061CE6910B96787E" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="3332A806C8327F24AF465147DC8C4A30">Holotype</typeStatus>
</emphasis>
<specimenCount id="040CF9C1B38822158B0361BF5F3D906E">♂</specimenCount>
(NHMW MY10262),
<collectingCountry id="629253FFC49CC9D32DB415DA2F4F9A9A" name="Serbia">Serbia</collectingCountry>
,
<normalizedToken id="6A4B038CFC6F25E30803466931DC27A0" originalValue="Niš">
<collectingRegion id="325FBB60CB149458EADBA803E30FD0E3" country="Serbia" name="Nis">Nis</collectingRegion>
</normalizedToken>
,
<collectingCounty id="EF4AD65C5481B07EB54A2C8F69F1B18D">Mt. Kalafat</collectingCounty>
, village of
<collectingMunicipality id="7DC78538C2D4654852A7865C16E40B10">Cerje</collectingMunicipality>
,
<location id="9834E05ACE9CDB7BEEAFDF46393036EC" LSID="urn:lsid:plazi:treatment:8FBCC310088A5EAE90DCB67E2AF03D7A:9834E05ACE9CDB7BEEAFDF46393036EC" country="Serbia" county="Mt. Kalafat" municipality="Cerje" name="Cerjanska Cave" stateProvince="Nis">Cerjanska Cave</location>
(=
<location id="205C08CAEA7A17E23B9CD3BCE60D460E" LSID="urn:lsid:plazi:treatment:8FBCC310088A5EAE90DCB67E2AF03D7A:205C08CAEA7A17E23B9CD3BCE60D460E" country="Serbia" county="Mt. Kalafat" municipality="Cerje" name="Provalija Cave" stateProvince="Nis">Provalija Cave</location>
),
<collectingDate id="089BB29726B3675EDB5223EE350B4F00" value="2017-10-29">29.X.2017</collectingDate>
, leg.
<collectorName id="466E0DBB1DA2F001482174F1C3C49343">
D.
<normalizedToken id="937D114D77E3924F1BE1DA1FF942E98B" originalValue="Antić">Antic</normalizedToken>
.
</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="1DB389086E661A571E5C50AE09B825E6" pageId="0" pageNumber="1">
<emphasis id="33A56B4BA0FA8EF91481262FB0AD12F5" bold="true" pageId="0" pageNumber="1">
<emphasis id="72423BD0D3ED010CB1BB30873ED7FEAD" italics="true" pageId="0" pageNumber="1">
<typeStatus id="344D785D4EA1C8E3B903B1DB29B90830">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="C0C56740571FF01026066A93290B6D4E" type="male">1 ♂</specimenCount>
(NHMW MY10263),
<specimenCount id="C785D817D2D38C49B86384A764AC2401" type="female">1 ♀</specimenCount>
(used for SEM, NHMW MY10264),
<materialsCitation id="B10E2DE2ECF1C8493C9C330FE04AFC4E" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="D. Antic." country="Serbia" county="Mt. Kalafat" location="Cerjanska Cave" municipality="Cerje" specimenCount="8" specimenCount-female="1" specimenCount-juvenile="7" stateProvince="Nis" typeStatus="paratype">
<specimenCount id="3E1898DD183DF004615FF67568667EA9" type="female">1 ♀</specimenCount>
,
<specimenCount id="EEF9F55FB9AA6C88503428298E380406" type="juvenile">7 juveniles</specimenCount>
(NHMW MY10265), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7A26EAAE4183049F3EC81D2E53F3516" pageId="0" pageNumber="1" type="description">
<paragraph id="802A53F4EC41B15DDA9AD1B7A5949993" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="F0455A7A2BDD86FADD11DC0F6C1FAF3F" pageId="0" pageNumber="1">
<emphasis id="D6D236856AC3A17A60D649BBA1D84ED9" bold="true" italics="true" pageId="0" pageNumber="1">Number of body rings and measurements</emphasis>
: Body with 19 rings (including telson) in adults, moniliform (Fig.
<figureCitation id="5C9E37327942C90F7EA0D32535DD84D0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10</figureCitation>
). Holotype male and paratype male 4.8 mm and 4.7 mm long, respectively; width of midbody pro- and metazonae 0.30 mm and 0.45 mm, respectively. Paratype females 5.0 mm and 5.2 mm long, width of midbody pro- and metazonae 0.35 mm and 0.50 mm, respectively.
</paragraph>
<caption id="A56B018D3081C4DBCD26960CD5337547" doi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph id="63430070E425FA102D589C79D6C85885" pageId="0" pageNumber="1">
<emphasis id="8EA9046C11EC3900EFC7E6222498CA0E" bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName id="80340D66FCA44D314CCB590EADEDE760" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="28A42C618C7B5DF8F61CB6CA3AB06CDC" italics="true" pageId="0" pageNumber="1">Balkanodesminus serbicus</emphasis>
</taxonomicName>
gen. nov. et sp. nov., habitus (NHMW)
<emphasis id="375DB89B961017478370DC9F285D1B1C" bold="true" pageId="0" pageNumber="1">A</emphasis>
mating in situ (photo D.
<normalizedToken id="878018E162F83571EB34A01E8C209F31" originalValue="Antić">Antic</normalizedToken>
)
<emphasis id="F3553CAA5E76B0E5BF8F95EFE969EF85" bold="true" pageId="0" pageNumber="1">B</emphasis>
holotype ♂ (NHMW MY10262) and paratype ♀ (NHMW MY10265), respectively, lateral views. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="084ACDA669D06FAF92D627E1DDA3817B" pageId="0" pageNumber="1">
<emphasis id="2501A108BBD67DC6C692621EE3E5E762" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
: Entirely pallid, slightly translucent (Fig.
<figureCitation id="AD03538EA2102685CF9125D5FB5B9723" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10</figureCitation>
).
</paragraph>
<paragraph id="B9356A3C1712EAD13B41288088F27E2B" pageId="0" pageNumber="1">
<emphasis id="3560F2D25336C765D6C367B5BE7EC600" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: Broader than collum, setose; epicranial suture poorly developed; isthmus between antennae ≈ 1.7
<normalizedToken id="CFB73B34F0320D58739A012ABD079B07" originalValue="×">x</normalizedToken>
broader than diameter of antennal socket (Fig.
<figureCitation id="B234815B55464E7CAAA2571E2CCDC744" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11A, B</figureCitation>
). Labrum with three labral teeth, and with 3+3 labral and five supralabral setae (Fig.
<figureCitation id="73E4E593F432104A18AF3C1203E23B35" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11A</figureCitation>
). Gnathochilarium without peculiarities. Antennae rather short, clavate (Fig.
<figureCitation id="D6FB66E86F900C22DA076ABA421467AF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11</figureCitation>
). Antennomere length 6&gt; 2 = 3 = 4&gt; 5&gt; 7 = 1. Antennae 0.7 mm long in the holotype male; length/breadth ratios of antennomeres 1-7: 1 (1), 2 (2), 3 (2), 4 (2), 5 (1), 6 (1.5) and 7 (1). Antennomere 6 with four sensilla trichoidea and with strongly developed disto-dorsal pit with numerous long sensilla basiconica partially exposed outside the pit (Fig.
<figureCitation id="CC2FBEABBAF5B75AD278968B40737240" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11E, F</figureCitation>
). Antennomere 7 with one sensillum trichodeum and a small bulge with three sensilla basiconica spiniformia (Fig.
<figureCitation id="B1F1A0450E547F89959011E3975A3535" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11E</figureCitation>
). Four apical cones (Fig.
<figureCitation id="16C88CCDE47607D14D8DF6C17C8EF80A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A head, anterior view B head and collum, lateral view C, D left antenna, semi-anterior and anterior views, respectively E right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) F sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (A-D), 0.02 mm (E, F)." figureDoi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1">11C</figureCitation>
).
</paragraph>
<caption id="7F233B653C5AD16020877AA8544D7B21" doi="10.3897/zookeys.1097.83916.figure11" httpUri="https://binary.pensoft.net/fig/674289" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph id="1A83C69D018BB0700164B1B95E42822F" pageId="0" pageNumber="1">
<emphasis id="DFD3D759CBECA704B44067093BE3C4E2" bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<taxonomicName id="E454E578F724D2DB60E345F3C8143472" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="514598898AA50E2AF7B1A9ECB585403B" italics="true" pageId="0" pageNumber="1">Balkanodesminus serbicus</emphasis>
</taxonomicName>
gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264)
<emphasis id="F86D5C1B4D4DC4ABD6AE147DC4AF711F" bold="true" pageId="0" pageNumber="1">A</emphasis>
head, anterior view
<emphasis id="5E71C4A5BF3F49C7472E9BEEEC34DE7D" bold="true" pageId="0" pageNumber="1">B</emphasis>
head and collum, lateral view
<emphasis id="DF58C592B05803B602AC3833FA4A9CB8" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
left antenna, semi-anterior and anterior views, respectively
<emphasis id="2D041D2FB500401D3B87616E0D907F8F" bold="true" pageId="0" pageNumber="1">E</emphasis>
right antennomeres 6 and 7, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia)
<emphasis id="40BCF5BA480F69BF2733A43A8F3CB86F" bold="true" pageId="0" pageNumber="1">F</emphasis>
sensilla basiconica of left antennomere 6, disto-dorsal view. Scale bars: 0.1 mm (
<emphasis id="EF79576A5A2BAA356A25ED0C3E16924C" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
), 0.02 mm (
<emphasis id="033CA9BF47C6DF3BBA348FC0FB058CCC" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
).
</paragraph>
</caption>
<paragraph id="BE93F9D77E7D5DAD76462A1222A5AF00" pageId="0" pageNumber="1">
<emphasis id="5CCFFFB0A9E87A633361C817828B0BEF" bold="true" italics="true" pageId="0" pageNumber="1">Collum</emphasis>
: Semi-circular, with one or two lateral incisions and ≈ 5 irregular rows of relatively long and trichoid setae.
</paragraph>
<paragraph id="B039C117DDD202F7739BA339BA3ACDED" pageId="0" pageNumber="1">
<emphasis id="FAB8755A639809132C4A090239EA3CD0" bold="true" italics="true" pageId="0" pageNumber="1">Body rings</emphasis>
: Tegument shining, texture alveolate, reticulate and scaly. Setae relatively long and trichoid, originating from small tubercules (Figs
<figureCitation id="8CC220F31F0D9C32204A2ED4C0D1AD3F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10B</figureCitation>
,
<figureCitation id="CBAAA176B4BEE94E56C9F5E098B6FDE7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12</figureCitation>
). Posteriormost tubercules mostly with a small thorn (Fig.
<figureCitation id="E7F6F73F94DEBF77F56FB02C236EE4D0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12A, E</figureCitation>
). Rings 2-4 with three mostly regular rows of setae, one anterior and two posterior (Fig.
<figureCitation id="4CBD686FF05FB3543F9DF2D54253337B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10B</figureCitation>
). Rings 5-18 with ≈ 4-6 irregular rows of setae (Figs
<figureCitation id="8A27AEBC6B9CCBA15ABC3E9AD8F4081C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10B</figureCitation>
,
<figureCitation id="87C1E42952C3725C81DA8C5280A0D707" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12</figureCitation>
). Paraterga serrated, with four or five teeth (Fig.
<figureCitation id="D93AF48F3C8A5323324999B27F7C2CBC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12A, D, E</figureCitation>
). Pore formula normal: 5, 7, 9, 10, 12, 13, 15-18. Poriferous metazonae with an enlarged posterolateral cone bearing an ozopore and three medium-sized setae (Fig.
<figureCitation id="9708B0FDDA607A1B105AB06568B32DE5" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12C</figureCitation>
). Epiproct blunt, directed slightly ventrad (Fig.
<figureCitation id="11B48E0CCC426CD54C9411DF16BE64F1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12G, H</figureCitation>
). Paraprocts semi-spherical, each with 2 long setae originating from small tubercules and ≈ 10 shorter setae without tubercules (Fig.
<figureCitation id="13BD4AA10DD8E4FC42F96A5B23DFFF35" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12H</figureCitation>
). Hypoproct trapeziform, with two long distal setae and numerous shorter setae throughout (Fig.
<figureCitation id="2B5538D0F9904AC6E734AE01A04335A1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10264) A, B midbody rings 10 and 11, dorsal and lateral views respectively C right ozopore 10, lateral view D right half of body ring 4, ventral view E, F body ring 17, dorsal and lateral views, respectively G telson, dorsal view H body ring 18 and telson, lateral view. Scale bars: 0.1 mm (A, B, E, F, H), 0.05 mm (G), 0.02 mm (D), 0.01 mm (C)." figureDoi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1">12H</figureCitation>
). Sterna unmodified, poorly setose. Pleurosternal carinae absent, only a few small teeth sometimes present on rings 2 and 3.
<normalizedToken id="C0A1756D3C1FD9DB6D11DFDC75B4EDF7" originalValue="Gonopod’s">Gonopod's</normalizedToken>
aperture large, subsemi-circular.
</paragraph>
<caption id="60CAF24F57836E376BD12CCB6A03D7C7" doi="10.3897/zookeys.1097.83916.figure12" httpUri="https://binary.pensoft.net/fig/674290" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph id="A26DA6D6EDD20B6D51D18B1B88753F7D" pageId="0" pageNumber="1">
<emphasis id="F1232428447E98C6473A61C5B789101B" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName id="B418D5A027D6EED4C5AC1D9CE49DD3AB" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="47758FF3A7AB09B62C785AE28883E63C" italics="true" pageId="0" pageNumber="1">Balkanodesminus serbicus</emphasis>
</taxonomicName>
gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY10264)
<emphasis id="6AA05145F45F4132B3599B8B8733EE5D" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
midbody rings 10 and 11, dorsal and lateral views respectively
<emphasis id="10B98144BFA85078762556AD9ED858A9" bold="true" pageId="0" pageNumber="1">C</emphasis>
right ozopore 10, lateral view
<emphasis id="92700BA6CFC8C5BDA5306B22784564C5" bold="true" pageId="0" pageNumber="1">D</emphasis>
right half of body ring 4, ventral view
<emphasis id="C4025BE1D8EF10FA7412F9C9E3CED545" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
body ring 17, dorsal and lateral views, respectively
<emphasis id="B22F4B62410E2AA4E03B71B2F030234A" bold="true" pageId="0" pageNumber="1">G</emphasis>
telson, dorsal view
<emphasis id="5E345E1D7FE281A5395D427B1B6D0628" bold="true" pageId="0" pageNumber="1">H</emphasis>
body ring 18 and telson, lateral view. Scale bars: 0.1 mm (
<emphasis id="8A103F30C14C4D8419DDFA7D666234D1" bold="true" pageId="0" pageNumber="1">A, B, E, F, H</emphasis>
), 0.05 mm (
<emphasis id="5CF63F1993851060DD05F754E32DE6D3" bold="true" pageId="0" pageNumber="1">G</emphasis>
), 0.02 mm (
<emphasis id="ED45912A51F70AE2F9B44F9D71E1B520" bold="true" pageId="0" pageNumber="1">D</emphasis>
), 0.01 mm (
<emphasis id="6A624A2F8CCFE250E9BAE4F2A2209CEF" bold="true" pageId="0" pageNumber="1">C</emphasis>
).
</paragraph>
</caption>
<paragraph id="B649AE7DE481BF466F369D821B901168" pageId="0" pageNumber="1">
<emphasis id="BBBC31CEAFD7B9E71A859CC582533AEE" bold="true" italics="true" pageId="0" pageNumber="1">Walking legs</emphasis>
: Legs 1-3 in males with swollen femur; coxa 2 with a short mesal apophysis (cf.
<bibRefCitation id="D6C6875317ADB390D341A50FBC1E9F80" author="Strasser, K" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="1" pagination="325 - 385" refId="B54" refString="Strasser, K, 1966a. Ueber Diplopoden Bulgariens. Annales Zoologici 23 (12): 325 - 385" title="Ueber Diplopoden Bulgariens." volume="23" year="1966 a">Strasser 1966a</bibRefCitation>
: 341, fig. 13). No other peculiarities.
</paragraph>
<caption id="8AA320EAB1ECC9FFF1F4F330986B7301" doi="10.3897/zookeys.1097.83916.figure13" httpUri="https://binary.pensoft.net/fig/674291" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph id="B7A7D243219ED1C984631864BD1B3CB3" pageId="0" pageNumber="1">
<emphasis id="3F6E952075210ECC2569321F8AC2A577" bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName id="52110836023396C674606F74D7E3F685" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="294CF66E7EFC5329D5EDC92FCEEA83B8" italics="true" pageId="0" pageNumber="1">Balkanodesminus serbicus</emphasis>
</taxonomicName>
gen. nov. et sp. nov., paratype ♂ left gonopod (NHMW MY10263)
<emphasis id="C1FED39D5BC7600ABD0A846674445D71" bold="true" pageId="0" pageNumber="1">A</emphasis>
mesal view
<emphasis id="9FC4E3795DB7EDA7BE9A8CB60B8A1940" bold="true" pageId="0" pageNumber="1">B</emphasis>
anterior view
<emphasis id="3F812DB3329CE98D3B2758A577CFC3A0" bold="true" pageId="0" pageNumber="1">C</emphasis>
distal view
<emphasis id="5145141FA80ACAF34033AE54B25EACA5" bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral view
<emphasis id="35C27370FBED47CE4B9114CA0F118226" bold="true" pageId="0" pageNumber="1">E</emphasis>
postro-lateral view
<emphasis id="49509E1413117F4EC48A166C929BCFB4" bold="true" pageId="0" pageNumber="1">F</emphasis>
posterior view. Abbreviations:
<emphasis id="C5327209FCC8CFDC89325DB5D166CC48" bold="true" pageId="0" pageNumber="1">a</emphasis>
acropodite,
<emphasis id="FD550C535585E7B43FD0680FFB81B2EA" bold="true" pageId="0" pageNumber="1">bl</emphasis>
basal lamellar lobe of solenophore,
<emphasis id="8E108112C96ABD417D8072CD1D4DA594" bold="true" pageId="0" pageNumber="1">ca</emphasis>
cannula,
<emphasis id="428F2B614F1EC2D41C4BF6B127B8E405" bold="true" pageId="0" pageNumber="1">cx</emphasis>
coxa,
<emphasis id="19EA9C77B26BD7704C6F7E6D0654F268" bold="true" pageId="0" pageNumber="1">dp</emphasis>
distal projection of solenophore,
<emphasis id="D62EEDB0AD7A2CA90598E517E49225DA" bold="true" pageId="0" pageNumber="1">dsp</emphasis>
distal solenomeral process,
<emphasis id="EDB33D831EA597DE7E341189C6EAEF58" bold="true" pageId="0" pageNumber="1">ll</emphasis>
lamella of solenophore,
<emphasis id="8C9E6B82E03F8B4C38361403C7BE278D" bold="true" pageId="0" pageNumber="1">pf</emphasis>
prefemorite,
<emphasis id="916FBBE766CA41BE03DE23A99B33907D" bold="true" pageId="0" pageNumber="1">s</emphasis>
solenomere,
<emphasis id="14DCE754916D7DDB126B59A7786DBC2C" bold="true" pageId="0" pageNumber="1">sb</emphasis>
solenomeral branch,
<emphasis id="1A783FC507B59C57D3C4301429C4899A" bold="true" pageId="0" pageNumber="1">sf</emphasis>
seminal fossa,
<emphasis id="7CDCFFCF16E39E87E37CBADCD0F373B7" bold="true" pageId="0" pageNumber="1">sg</emphasis>
seminal groove,
<emphasis id="054B8DC040634111BAC5494EDF5A003C" bold="true" pageId="0" pageNumber="1">sph</emphasis>
solenophore. Scale bars: 0.02 mm.
</paragraph>
</caption>
<paragraph id="E40B7831921928CE36339F1BDDCEB67E" pageId="0" pageNumber="1">
<taxonomicName id="E607EF99CDB48E89B38B191684424EB3" genus="Gonopods" lsidName="Gonopods" pageId="0" pageNumber="1" rank="genus">
<emphasis id="6BF4361070AFAE113F5C3BD3E28528A6" bold="true" italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
</taxonomicName>
(Figs
<figureCitation id="9F34280540EEE9A7D294C3C952900B92" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Balkanodesminus serbicus gen. nov. et sp. nov., paratype ♂ left gonopod (NHMW MY 10263) A mesal view B anterior view C distal view D lateral view E postro-lateral view F posterior view. Abbreviations: a acropodite, bl basal lamellar lobe of solenophore, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, pf prefemorite, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore. Scale bars: 0.02 mm." figureDoi="10.3897/zookeys.1097.83916.figure13" httpUri="https://binary.pensoft.net/fig/674291" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation id="9770D2195EEA41B58FB72546C509E86A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">17J</figureCitation>
): Coxa (cx) large, semi-circular in ventral and lateral views, with differentiated gonocoel mesally; lateral part swollen, alveolate, with three long setae near mesal ridge. Cannula (ca) long, C-shaped. Telopodite long compared to coxa, consisting of a transverse, setose prefemorite (pf) and a somewhat C-shaped (in lateral and mesal views) acropodite (a) longitudinally divided in its distal half into two branches, solenomeral branch (sb) and solenophore (sph). Solenomeral branch positioned mesally, with a narrow base, then abruptly expands and transversely divides into two processes, solenomere (s) and distal solenomeral process (dsp). Solenomere (s) long, slender, distally expanded (in lateral and mesal views), forming U-shaped rift with distal solenomeral process. Distal solenomeral process (dsp) extends in the same direction as solenomere,
<normalizedToken id="990A8F6D7404CCD6B78456BB9882EC39" originalValue="¾">3/4</normalizedToken>
the length of solenomere; ending with a small expansion (in lateral and mesal views). Solenophore (sph) longer and more robust than solenomeral branch, characterized by a lateral, ear-shaped lamella (ll) and a distal projection (dp). Lateral lamella (ll) begins at bifurcation of solenomeral branch and solenophore, surrounding laterally solenophore up to beginning of distal projection; lateral margins of lamella smooth. Distal projection (dp) sigmoid (in lateral and mesal views), with well-developed, basal lamellar lobe (bl). Seminal groove (sg) starts from seminal fossa (sf) mesally on prefemorite, extends along mesal side of acropodite up to bifurcation of solenomeral branch and solenophore, then passes on lateral side of solenomeral branch, further on solenomere, ending subdistally.
</paragraph>
</subSubSection>
<subSubSection id="716711E887072921878F566231F3A96A" pageId="0" pageNumber="1" type="habitat">
<paragraph id="D27E0F15BDB194112AF07F2A00536A66" pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph id="0F7FB9DDE35D2871B9CA7CED22EDF5E1" pageId="0" pageNumber="1">
With its 6131 m of explored channels, the Cerjanska Cave represents one of the longest and most significant fluviokarst underground systems in Serbia. This is a relatively simple speleological object, consisting of one main river channel in two levels with a length of 4903 m, as well as several side channels with a total length of 1228 m (
<bibRefCitation id="D6845EE53645E5C4EAB1E3C581FA522F" author="Nesic, D" editor="Nesic, D" journalOrPublisher="Surroundings, Natural History, Protection. Institute for Nature Conservation in Nis, Nis, Serbia" pageId="0" pageNumber="1" pagination="47 - 62" refId="B40" refString="Nesic, D, 2016. Provalija cave (Cerjanska cave), morphology and formation. In: Nesic, D, Jovic, D, Eds., Natural monument Cerjanska Cave. Surroundings, Natural History, Protection. Institute for Nature Conservation in Nis, Nis, Serbia: 47 - 62" title="Provalija cave (Cerjanska cave), morphology and formation." volumeTitle="Natural monument Cerjanska Cave." year="2016">
<normalizedToken id="1ECB1BCC0B02153EAE76ED3BA772925F" originalValue="Nešić">Nesic</normalizedToken>
2016
</bibRefCitation>
). Numerous arthropod taxa have been registered in the cave, from epigean, guanophiles, trogloxenes, and troglophiles to troglobionts (
<bibRefCitation id="8D9FEBB6FD3BBDAC3C071B13986E0486" author="Pavicevic, D" editor="Nesic, D" journalOrPublisher="Surroundings, Natural History, Protection. Institute for Nature Conservation in Nis, Nis, Serbia" pageId="0" pageNumber="1" pagination="90 - 99" refId="B41" refString="Pavicevic, D, Zatezalo, A, Popovic, M, 2016. Biospeleological research of Cerjanska Cave and the northern part of Kalafat area. In: Nesic, D, Jovic, D, Eds., Natural monument Cerjanska Cave. Surroundings, Natural History, Protection. Institute for Nature Conservation in Nis, Nis, Serbia: 90 - 99" title="Biospeleological research of Cerjanska Cave and the northern part of Kalafat area." volumeTitle="Natural monument Cerjanska Cave." year="2016">
<normalizedToken id="5E939A42A0CAAC60A5F531B5BA7323FA" originalValue="Pavićević">Pavicevic</normalizedToken>
et al. 2016
</bibRefCitation>
). The troglobionts include the endemic Balkan harvestman
<taxonomicName id="D40AF09FBBC36D003B82D7798431A223" baseAuthorityName="Roewer" baseAuthorityYear="1926" class="Arachnida" family="Nemastomatidae" genus="Paranemastoma" kingdom="Animalia" lsidName="Paranemastoma bureschi" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bureschi">
<emphasis id="8A6C6C967B61222EDCB7B6C657381D2F" italics="true" pageId="0" pageNumber="1">Paranemastoma bureschi</emphasis>
</taxonomicName>
(Roewer, 1926), the millipede
<taxonomicName id="42B4F74D090AB8A36CB7A53EAB692D96" authorityName="Makarov &amp; Curcic" authorityYear="2012" class="Diplopoda" family="Anthroleucosomatidae" genus="Dazbogosoma" kingdom="Animalia" lsidName="Dazbogosoma naissi" order="Chordeumatida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="naissi">
<emphasis id="5A23E12444673B2DA84B74687977101D" italics="true" pageId="0" pageNumber="1">Dazbogosoma naissi</emphasis>
</taxonomicName>
Makarov &amp;
<normalizedToken id="A97039A5D420B2B95D2096B019E7677F" originalValue="Ćurčić">Curcic</normalizedToken>
in
<bibRefCitation id="B17FB7F4760C1FE46231ED212D12B912" DOI="https://doi.org/10.2298/ABS1202793M" author="Makarov, SE" journalOrPublisher="Archives of Biological Sciences" pageId="0" pageNumber="1" pagination="793 - 798" refId="B37" refString="Makarov, SE, Curcic, BPM, Milincic, M, Pecelj, MM, Antic, DZ, Mitic, BM, 2012. A new genus and species of the family Anthroleucosomatidae from Serbia (Myriapoda, Diplopoda, Chordeumatida). Archives of Biological Sciences 64 (2): 793 - 798, DOI: https://doi.org/10.2298/ABS1202793M" title="A new genus and species of the family Anthroleucosomatidae from Serbia (Myriapoda, Diplopoda, Chordeumatida)." url="https://doi.org/10.2298/ABS1202793M" volume="64" year="2012">Makarov et al. 2012</bibRefCitation>
, and the carabid beetle
<taxonomicName id="C79474B90AD2968DE473303DBD187991" authorityName="Pavicevic, Zatezalo &amp; Popovic" authorityYear="2016" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" lsidName="Duvalius rtanjensis subsp. provalijae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="rtanjensis" subSpecies="provalijae">
<emphasis id="57C2651AB48AC9E81A3D50F50C04FCCD" italics="true" pageId="0" pageNumber="1">Duvalius rtanjensis provalijae</emphasis>
</taxonomicName>
<normalizedToken id="C8146B007ABC1837A3D72A09C572F543" originalValue="Pavićević">Pavicevic</normalizedToken>
, Zatezalo &amp;
<normalizedToken id="A113735E9F5241E1CA813CC64A9D29BE" originalValue="Popović">Popovic</normalizedToken>
, 2016, the latter two endemics of Cerjanska Cave.
</paragraph>
<paragraph id="7D0135FEE78FE68677E3743252F048E5" pageId="0" pageNumber="1">
Despite many years of speleological and biospeleological research in the Cerjanska Cave, the new taxon was not registered until the first Biospeleological Expedition of the Serbian Biospeleological Society, organized at the end of October 2017. All 11 specimens were found in a small area, in the initial part of the cave. One male, one female and seven juveniles were found on the left side of the river, on a small branch of a tree lying on the wet sand. Another male and female were found just on the opposite side of the river, on the wall, in copulation (Fig.
<figureCitation id="C3AF93ACE4F04D24EC6031D51485BAA1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Balkanodesminus serbicus gen. nov. et sp. nov., habitus (NHMW) A mating in situ (photo D. Antic) B holotype ♂ (NHMW MY 10262) and paratype ♀ (NHMW MY 10265), respectively, lateral views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure10" httpUri="https://binary.pensoft.net/fig/674288" pageId="0" pageNumber="1">10A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D46881906E7EEC36BA50DDF3134D54EC" pageId="0" pageNumber="1" type="distribution">
<paragraph id="576D1BFC767FBFDB9FCD2C8B0580082E" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="138DA19339C317176CCEA2D17481B35A" pageId="0" pageNumber="1">
So far, known only from its type locality, the Cerjanska Cave, Serbia (Fig.
<figureCitation id="60F34344D64F473958EB71B9C7000894" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77EAA800B92C99AE1C244637BBD72218" pageId="0" pageNumber="1" type="remarks">
<paragraph id="A6123627F3684475C2D10E461C6119C3" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="89BEC0D7BD9443C35E10C1F8D81E1FDE" pageId="0" pageNumber="1">
This is the first representative of the family
<taxonomicName id="F07C0B8F503E7991C34B183C4EE2C012" authorityName="Verhoeff" authorityYear="1910" class="Diplopoda" family="Trichopolydesmidae" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
in Serbia.
</paragraph>
</subSubSection>
</treatment>
</document>
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