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<mods:title id="7D03F639091F255CCF3BAB2FB2676091">Contribution to the knowledge of the genus Dendarus Dejean, 1821 (Coleoptera: Tenebrionidae: Blaptinae: Dendarini) from Iran, Turkmenistan and some adjacent territories</mods:title>
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<mods:namePart id="3D4E2F465D0F89F9D3D4ACB73C728BF3">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="D65C3A5E54E79807C7104BA27C1AE23F">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia.</mods:affiliation>
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<mods:namePart id="29804E4B234958874F4844B085CAAB2C">Poggi, Roberto</mods:namePart>
<mods:affiliation id="B712E7417C669F3087E6D637F17C0E5B">Museo Civico di Storia Naturale “ Giacomo Doria ”, Genova, 16121 Italy.</mods:affiliation>
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<treatment id="8F671405FFC6FFE9FF41FBA3FCA4FF5C" ID-DOI="http://doi.org/10.5281/zenodo.6669366" ID-GBIF-Taxon="196085049" ID-Zenodo-Dep="6669366" LSID="urn:lsid:plazi:treatment:8F671405FFC6FFE9FF41FBA3FCA4FF5C" httpUri="http://treatment.plazi.org/id/8F671405FFC6FFE9FF41FBA3FCA4FF5C" lastPageId="10" lastPageNumber="115" pageId="9" pageNumber="114">
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<taxonomicName id="C0CEDE90FFC6FFEAFF41FBA3FD82FB0A" authority="Brancsik, 1899" authorityName="Brancsik" authorityYear="1899" box="[151,636,1126,1153]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="114" phylum="Arthropoda" rank="species" species="transcaspicus">
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<emphasis id="35BA7901FFC6FFEAFF41FBA3FE45FB0A" bold="true" box="[151,443,1126,1153]" italics="true" pageId="9" pageNumber="114">Dendarus transcaspicus</emphasis>
<bibRefCitation id="635FD8E2FFC6FFEAFE17FBA3FD82FB0A" author="Brancsik, C." box="[449,636,1126,1152]" pageId="9" pageNumber="114" pagination="97 - 105" refId="ref10330" refString="Brancsik, C. (1899) Aliquot Coleoptera novae Russiae asiaticae. Evkonyve, A Trencsen Varmegyei Termeszettudomanyi Egylet [Jahresheft des Naturwissenschaftlichen Vereines des Trencsiner Comitates], 21 - 22, 97 - 105 + [1], pl. 4. [1898 - 1899]" type="journal article" year="1899">Brancsik, 1899</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="4FD4F698FFC6FFEAFF41FB4FFC82FA8F" pageId="9" pageNumber="114" type="description">
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(
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Figs
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,
<figureCitation id="9FF5B996FFC6FFEAFF22FB4FFEDDFB2E" box="[244,291,1162,1188]" captionStart="FIGURE 10" captionStartId="13.[151,250,1678,1702]" captionTargetBox="[158,1433,1022,1645]" captionTargetId="figure-65@13.[151,1437,1017,1655]" captionTargetPageId="13" captionText="FIGURE 10. Dendarus spp., male tergite VIII. A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus." figureDoi="http://doi.org/10.5281/zenodo.6669278" httpUri="https://zenodo.org/record/6669278/files/figure.png" pageId="9" pageNumber="114">10E</figureCitation>
,
<figureCitation id="9FF5B996FFC6FFEAFEF9FB4FFEA3FB2E" box="[303,349,1162,1188]" captionStart="FIGURE 11" captionStartId="14.[151,250,1173,1197]" captionTargetBox="[156,1424,385,1146]" captionTargetId="figure-108@14.[151,1436,381,1150]" captionTargetPageId="14" captionText="FIGURE 11. Dendarus spp., male inner sternite VIII.A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus. sVIII—inner sternite VIII, tVIII—inner tergite VIII, tIX—inner tergite IX." figureDoi="http://doi.org/10.5281/zenodo.6669284" httpUri="https://zenodo.org/record/6669284/files/figure.png" pageId="9" pageNumber="114">11E</figureCitation>
,
<figureCitation id="9FF5B996FFC6FFEAFEBFFB4FFE66FB2E" box="[361,408,1162,1188]" captionStart="FIGURE 12" captionStartId="14.[151,250,1967,1991]" captionTargetBox="[155,1430,1261,1940]" captionTargetId="figure-164@14.[151,1436,1257,1943]" captionTargetPageId="14" captionText="FIGURE 12. Dendarus spp., spiculum gastrale.A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus." figureDoi="http://doi.org/10.5281/zenodo.6669288" httpUri="https://zenodo.org/record/6669288/files/figure.png" pageId="9" pageNumber="114">12E</figureCitation>
,
<figureCitation id="9FF5B996FFC6FFEAFE72FB4FFE2FFB2E" box="[420,465,1162,1188]" captionStart="FIGURE 13" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[182,1391,190,1909]" captionTargetId="figure-17@15.[181,1406,181,1920]" captionTargetPageId="15" captionText="FIGURE 13. Dendarus spp., aedeagus, ventrally. A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = the same, turned forward; F = D. transcaspicus; G = D. vagabundus." figureDoi="http://doi.org/10.5281/zenodo.6669294" httpUri="https://zenodo.org/record/6669294/files/figure.png" pageId="9" pageNumber="114">13F</figureCitation>
)
</paragraph>
<paragraph id="0771A513FFC6FFEAFF41FB0AFC82FA8F" blockId="9.[151,1436,1231,1285]" pageId="9" pageNumber="114">
<bibRefCitation id="635FD8E2FFC6FFEAFF41FB0AFE96FB6D" author="Brancsik, C." box="[151,360,1231,1255]" pageId="9" pageNumber="114" pagination="97 - 105" refId="ref10330" refString="Brancsik, C. (1899) Aliquot Coleoptera novae Russiae asiaticae. Evkonyve, A Trencsen Varmegyei Termeszettudomanyi Egylet [Jahresheft des Naturwissenschaftlichen Vereines des Trencsiner Comitates], 21 - 22, 97 - 105 + [1], pl. 4. [1898 - 1899]" type="journal article" year="1899">Brancsik, 1899: 101</bibRefCitation>
;
<bibRefCitation id="635FD8E2FFC6FFEAFEA5FB0AFDDEFB6D" author="Reitter, E." box="[371,544,1231,1255]" pageId="9" pageNumber="114" pagination="25 - 189" refId="ref11342" refString="Reitter, E. (1904) Bestimmungs-Tabelle der Tenebrioniden-Unterfamilien: Lachnogyini, Akidini, Pedinini, Opatrini und Trachyscelini aus Europa und den angrenzenden Landern. Verhandlungen des Naturforschenden Vereines in Brunn, 42, 25 - 189." type="journal article" year="1904">Reitter, 1904: 84</bibRefCitation>
;
<bibRefCitation id="635FD8E2FFC6FFEAFDFAFB0AFD00FB6D" author="Medvedev, G. S." box="[556,766,1231,1255]" pageId="9" pageNumber="114" refId="ref11059" refString="Medvedev, G. S. (1968) Fauna of the USSR. Coleoptera. Vol. 19. Issue. 2. Darkling beetles (Tenebrionidae). Subfamily Opatrinae. Tribes Platynotini, Dendarini, Pedinini, Dissonomini, Pachypterini, Opatrini (part) and Heterotarsini. Nauka, Leningrad, 285 pp. [in Russian]" type="book" year="1968">Medvedev, 1968: 98</bibRefCitation>
, 99, 110, figs 109111;
<bibRefCitation id="635FD8E2FFC6FFEAFC25FB0AFAA4FB6D" author="Medvedev, G. S. &amp; Nepesova, M. G." box="[1011,1370,1231,1255]" pageId="9" pageNumber="114" refId="ref11121" refString="Medvedev, G. S. &amp; Nepesova, M. G. (1985) Key to darkling beetles of Turkmenistan. Ylym, Ashgabat, 180 pp. [in Russian]" type="book" year="1985">Medvedev &amp; Nepesova, 1985: 123</bibRefCitation>
;
<bibRefCitation id="635FD8E2FFC6FFEAFAB3FB0AFE02FA8F" author="Medvedev, G. S. &amp; Nepesova, M. G." pageId="9" pageNumber="114" pagination="23 - 30" refId="ref11155" refString="Medvedev, G. S. &amp; Nepesova, M. G. (1990 a) Review of darkling beetles (Coleoptera, Tenebrionidae) of Kopetdagh (communication 2). Izvestiya Akademii nauk Turkmenskoy SSR. Seria biologicheskikh nauk, 3, 23 - 30. [in Russian]" type="journal article" year="1990">Medvedev &amp; Nepesova, 1990a: 25</bibRefCitation>
;
<bibRefCitation id="635FD8E2FFC6FFEAFDD1FB28FC86FA8F" author="Medvedev, G. S. &amp; Nepesova, M. G." box="[519,888,1261,1285]" pageId="9" pageNumber="114" pagination="879 - 889" refId="ref11208" refString="Medvedev, G. S. &amp; Nepesova, M. G. (1990 b) Composition and geographic distribution of the fauna of tenebrionid beetles (Coleoptera, Tenebrionidae) of Kopetdagh. Entomologicheskoe obozrenie, 69, 879 - 889. [in Russian]" type="journal article" year="1990">Medvedev &amp; Nepesova, 1990b: 886</bibRefCitation>
.
</paragraph>
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<emphasis id="35BA7901FFC6FFEAFF41FAF1FEB8FAC7" bold="true" box="[151,326,1331,1358]" pageId="9" pageNumber="114">
<typeStatus id="D8751BB1FFC6FFEAFF41FAF1FF2CFAC4" box="[151,210,1332,1358]" pageId="9" pageNumber="114">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D8751BB1FFC6FFEAFE98FAF1FE6EFAC4" box="[334,400,1332,1358]" pageId="9" pageNumber="114">Types</typeStatus>
were not found in the collection of K. Brancsik in Field Museum, Chicago,
<collectingCountry id="7FD9E583FFC6FFEAFB20FAF1FAD2FAC4" box="[1270,1324,1332,1358]" name="United States of America" pageId="9" pageNumber="114">USA</collectingCountry>
(personal communication of the curator Maureen Turcatel). Here we interpreted the species according to the original description of
<bibRefCitation id="635FD8E2FFC6FFEAFF3EFABEFE63FA1C" author="Brancsik, C." box="[232,413,1403,1430]" pageId="9" pageNumber="114" pagination="97 - 105" refId="ref10330" refString="Brancsik, C. (1899) Aliquot Coleoptera novae Russiae asiaticae. Evkonyve, A Trencsen Varmegyei Termeszettudomanyi Egylet [Jahresheft des Naturwissenschaftlichen Vereines des Trencsiner Comitates], 21 - 22, 97 - 105 + [1], pl. 4. [1898 - 1899]" type="journal article" year="1899">Brancsik (1899)</bibRefCitation>
(especially slender and dull body; only one species has this combination of characters) and the redescription of
<bibRefCitation id="635FD8E2FFC6FFEAFEAEFA5AFDBAFA30" author="Medvedev, G. S." box="[376,580,1439,1466]" pageId="9" pageNumber="114" refId="ref11059" refString="Medvedev, G. S. (1968) Fauna of the USSR. Coleoptera. Vol. 19. Issue. 2. Darkling beetles (Tenebrionidae). Subfamily Opatrinae. Tribes Platynotini, Dendarini, Pedinini, Dissonomini, Pachypterini, Opatrini (part) and Heterotarsini. Nauka, Leningrad, 285 pp. [in Russian]" type="book" year="1968">Medvedev (1968)</bibRefCitation>
.
</paragraph>
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<emphasis id="35BA7901FFC6FFEAFF11FA06FEC8FA57" bold="true" box="[199,310,1475,1501]" pageId="9" pageNumber="114">Material.</emphasis>
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<collectingCountry id="7FD9E583FFC6FFEAFEE9FA06FE23FA57" box="[319,477,1475,1501]" name="Turkmenistan" pageId="9" pageNumber="114">Turkmenistan</collectingCountry>
,
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(
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):
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[now Bereket] environs, mountains,
<quantity id="C03608F6FFC6FFEAFB45FA06FB20FA54" box="[1171,1246,1475,1502]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="114" unit="m" value="600.0">
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,
<date id="737083D3FFC6FFEAFB3DFA06FA9FFA54" box="[1259,1377,1475,1502]" pageId="9" pageNumber="114" value="1952-04-21">
<collectingDate id="63347A3BFFC6FFEAFB3DFA06FA9FFA54" box="[1259,1377,1475,1502]" pageId="9" pageNumber="114" value="1952-04-21">21.iv.1952</collectingDate>
</date>
(leg.
<collectorName id="AA3BC0C5FFC6FFEAFF41FA2DFE83F988" box="[151,381,1511,1538]" pageId="9" pageNumber="114">O.L. Kryzhanovskij</collectorName>
)
</materialsCitation>
;
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<specimenCount id="11C86E9AFFC6FFEAFE44FA2DFE4BF988" box="[402,437,1512,1538]" pageId="9" pageNumber="114" type="female">1♀</specimenCount>
(
<collectionCode id="61DF3DD6FFC6FFEAFE10FA2DFE05F988" box="[454,507,1512,1538]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="114" type="Museum">ZIN</collectionCode>
):
<location id="0211F3C8FFC6FFEAFDC6FA22FD7EF98B" LSID="urn:lsid:plazi:treatment:8F671405FFC6FFE9FF41FBA3FCA4FF5C:0211F3C8FFC6FFEAFDC6FA22FD7EF98B" box="[528,640,1511,1537]" country="Turkmenistan" name="Kopetdag" pageId="9" pageNumber="114">Kopetdag</location>
,
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<quantity id="C03608F6FFC6FFEAFD5BFA22FD28F98B" box="[653,726,1511,1538]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="114" unit="km" value="15.0">15 km</quantity>
S Iskander station
</locationDeviation>
,
<date id="737083D3FFC6FFEAFC6DFA22FBCDF988" box="[955,1075,1511,1538]" pageId="9" pageNumber="114" value="1953-06-13">
<collectingDate id="63347A3BFFC6FFEAFC6DFA22FBCDF988" box="[955,1075,1511,1538]" pageId="9" pageNumber="114" value="1953-06-13">13.vi.1953</collectingDate>
</date>
(leg.
<collectorName id="AA3BC0C5FFC6FFEAFBA1FA2DFAA3F988" box="[1143,1373,1511,1538]" pageId="9" pageNumber="114">O.L. Kryzhanovskij</collectorName>
)
</materialsCitation>
;
<materialsCitation id="B7A6AF4EFFC6FFEAFAA4FA2DFB7CF9AC" ID-GBIF-Occurrence="3816430324" collectingDate="1952-04-26" collectionCode="ZIN" collectorName="Shteinberg" country="Turkmenistan" location="Serdar" pageId="9" pageNumber="114" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="11C86E9AFFC6FFEAFAA4FA2DFA69F988" box="[1394,1431,1512,1538]" pageId="9" pageNumber="114" type="male">1♂</specimenCount>
,
<specimenCount id="11C86E9AFFC6FFEAFF41F9C9FF44F9AC" box="[151,186,1548,1574]" pageId="9" pageNumber="114" type="female">1♀</specimenCount>
(
<collectionCode id="61DF3DD6FFC6FFEAFF1CF9C9FF01F9AC" box="[202,255,1548,1574]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="114" type="Museum">ZIN</collectionCode>
): mountains
<locationDeviation id="16C3BC31FFC6FFEAFE58F9C9FD62F9AF" box="[398,668,1547,1574]" location="Kyzyl-Arvat" pageId="9" pageNumber="114">
<quantity id="C03608F6FFC6FFEAFE58F9C9FE28F9AF" box="[398,470,1547,1574]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="114" unit="km" value="12.0">12 km</quantity>
SW Kyzyl-Arvat
</locationDeviation>
[now
<location id="0211F3C8FFC6FFEAFD32F9CEFCCDF9AC" LSID="urn:lsid:plazi:treatment:8F671405FFC6FFE9FF41FBA3FCA4FF5C:0211F3C8FFC6FFEAFD32F9CEFCCDF9AC" box="[740,819,1547,1574]" country="Turkmenistan" name="Serdar" pageId="9" pageNumber="114">Serdar</location>
],
<date id="737083D3FFC6FFEAFC92F9CEFC45F9AC" box="[836,955,1547,1574]" pageId="9" pageNumber="114" value="1952-04-26">
<collectingDate id="63347A3BFFC6FFEAFC92F9CEFC45F9AC" box="[836,955,1547,1574]" pageId="9" pageNumber="114" value="1952-04-26">26.iv.1952</collectingDate>
</date>
(leg.
<collectorName id="AA3BC0C5FFC6FFEAFC2AF9CEFB84F9AC" box="[1020,1146,1547,1574]" pageId="9" pageNumber="114">Shteinberg</collectorName>
)
</materialsCitation>
;
<materialsCitation id="B7A6AF4EFFC6FFEAFB5BF9C9FC5FF9C0" ID-GBIF-Occurrence="3816430356" collectingDate="1952-04-23" collectionCode="ZIN" collectorName="K. Romadina" country="Turkmenistan" elevation="900" location="Kopetdag" pageId="9" pageNumber="114" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="11C86E9AFFC6FFEAFB5BF9C9FB4CF9AC" box="[1165,1202,1548,1574]" pageId="9" pageNumber="114" type="male">1♂</specimenCount>
,
<specimenCount id="11C86E9AFFC6FFEAFB69F9C9FB08F9AC" box="[1215,1270,1548,1574]" pageId="9" pageNumber="114" type="female">3♀♀</specimenCount>
(
<collectionCode id="61DF3DD6FFC6FFEAFAD0F9C9FAC5F9AC" box="[1286,1339,1548,1574]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="114" type="Museum">ZIN</collectionCode>
):
<location id="0211F3C8FFC6FFEAFA98F9C9FF3CF9C3" LSID="urn:lsid:plazi:treatment:8F671405FFC6FFE9FF41FBA3FCA4FF5C:0211F3C8FFC6FFEAFA98F9C9FF3CF9C3" country="Turkmenistan" name="Kopetdag" pageId="9" pageNumber="114">Kopetdag</location>
,
<locationDeviation id="16C3BC31FFC6FFEAFF19F9F5FE1EF9C3" box="[207,480,1583,1610]" location="Kyzyl-Arvat" pageId="9" pageNumber="114">
<quantity id="C03608F6FFC6FFEAFF19F9F5FEE9F9C3" box="[207,279,1583,1610]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="114" unit="km" value="12.0">12 km</quantity>
SW Kyzyl-Arvat
</locationDeviation>
,
<quantity id="C03608F6FFC6FFEAFE3BF9F5FDC9F9C0" box="[493,567,1584,1610]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="114" unit="m" value="900.0">
<elevation id="8CE34220FFC6FFEAFE3BF9F5FDC9F9C0" box="[493,567,1584,1610]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="114" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<date id="737083D3FFC6FFEAFD92F9EAFD44F9C0" box="[580,698,1583,1610]" pageId="9" pageNumber="114" value="1952-04-23">
<collectingDate id="63347A3BFFC6FFEAFD92F9EAFD44F9C0" box="[580,698,1583,1610]" pageId="9" pageNumber="114" value="1952-04-23">23.iv.1952</collectingDate>
</date>
(leg.
<collectorName id="AA3BC0C5FFC6FFEAFD2BF9F5FC66F9C0" box="[765,920,1583,1610]" pageId="9" pageNumber="114">K. Romadina</collectorName>
)
</materialsCitation>
;
<materialsCitation id="B7A6AF4EFFC6FFEAFC7AF9F5FE4FF9E4" ID-GBIF-Occurrence="3816430349" collectingDate="1952-04-25" collectionCode="ZIN" collectorName="Il'ichev" country="Turkmenistan" location="Kyzyl-Arvat" pageId="9" pageNumber="114" specimenCount="1" specimenCount-male="1">
<specimenCount id="11C86E9AFFC6FFEAFC7AF9F5FC31F9C0" box="[940,975,1584,1610]" pageId="9" pageNumber="114" type="male">1♂</specimenCount>
(
<collectionCode id="61DF3DD6FFC6FFEAFC36F9F5FBEBF9C0" box="[992,1045,1584,1610]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="114" type="Museum">ZIN</collectionCode>
): footnotes,
<quantity id="C03608F6FFC6FFEAFB76F9F5FB16F9C3" box="[1184,1256,1583,1610]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="114" unit="km" value="13.0">13 km</quantity>
S
<location id="0211F3C8FFC6FFEAFADEF9EAFA66F9C3" LSID="urn:lsid:plazi:treatment:8F671405FFC6FFE9FF41FBA3FCA4FF5C:0211F3C8FFC6FFEAFADEF9EAFA66F9C3" box="[1288,1432,1583,1609]" country="Turkmenistan" name="Kyzyl-Arvat" pageId="9" pageNumber="114">Kyzyl-Arvat</location>
,
<date id="737083D3FFC6FFEAFF41F996FEF3F9E4" box="[151,269,1619,1646]" pageId="9" pageNumber="114" value="1952-04-25">
<collectingDate id="63347A3BFFC6FFEAFF41F996FEF3F9E4" box="[151,269,1619,1646]" pageId="9" pageNumber="114" value="1952-04-25">25.iv.1952</collectingDate>
</date>
(leg.
<collectorName id="AA3BC0C5FFC6FFEAFE98F996FE54F9E4" box="[334,426,1619,1646]" pageId="9" pageNumber="114">Ilichev</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FD4F698FFC6FFE9FF11F9B2FBB5FF3B" lastPageId="10" lastPageNumber="115" pageId="9" pageNumber="114" type="diagnosis">
<paragraph id="0771A513FFC6FFEAFF11F9B2FC15F873" blockId="9.[151,1437,1331,2042]" pageId="9" pageNumber="114">
<emphasis id="35BA7901FFC6FFEAFF11F9B2FEBCF91B" bold="true" box="[199,322,1655,1681]" pageId="9" pageNumber="114">Diagnosis.</emphasis>
Male. Head and pronotum dorsally shiny, elytra matt. Body slender, legs and antennae slender, long. Head with moderately coarse, sparse puncturation (puncture diameter subequal to interpuncture distance). Head ventrally with simple (not impressed) longitudinal furrow beyond prementum. Pronotum strongly convex. Lateral margins of pronotum evenly rounded, not emarginated at basal quarter; anterior angles not protruding; base slightly bisinuate, posterior angles rectangular or weakly obtuse, not protruding. Puncturation of pronotum uneven: fine and sparse, with round punctures at middle and sharply coarse and dense elongate, often merged punctures on lateral sides. Prosternal process comparatively wide, rounded at apex. Elytra widest at middle or slightly beyond middle. Strial punctures small, moderately sparse (3034 punctures in one row), round, weakly impressed. Interstriae with very fine and sparse, poorly visible puncturation. Protibiae narrow, with a deep and wide notch on inner side and protruding rectangular tooth before indentation; protibia widest at level of tooth and apex. Mesotibiae straight. Mesotarsomeres 14 not widened, tarsomere 2 with reduced hair brush on sole.
</paragraph>
<paragraph id="0771A513FFC5FFE9FF11FF52FBB5FF3B" blockId="10.[199,1099,151,214]" box="[199,1099,151,177]" pageId="10" pageNumber="115">Female. Body more robust, protibiae weakly bent. Elytra widest beyond middle.</paragraph>
</subSubSection>
<subSubSection id="4FD4F698FFC5FFE9FF11FF7FFCA4FF5C" box="[199,858,186,214]" pageId="10" pageNumber="115" type="distribution">
<paragraph id="0771A513FFC5FFE9FF11FF7FFCA4FF5C" blockId="10.[199,1099,151,214]" box="[199,858,186,214]" pageId="10" pageNumber="115">
<emphasis id="35BA7901FFC5FFE9FF11FF7FFE9FFF5E" bold="true" box="[199,353,186,212]" pageId="10" pageNumber="115">Distribution.</emphasis>
S
<collectingCountry id="7FD9E583FFC5FFE9FEABFF7EFDE6FF5F" box="[381,536,187,213]" name="Turkmenistan" pageId="10" pageNumber="115">Turkmenistan</collectingCountry>
(Kopetdag, low mountains).
</paragraph>
</subSubSection>
</treatment>
</document>