498 lines
35 KiB
XML
498 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1081.71872" ID-GBIF-Dataset="1ca2a495-97f3-4226-8c61-df278e3a66e7" ID-PMC="PMC8786815" ID-Pensoft-Pub="1313-2970-1081-137" ID-Pensoft-UUID="B8C62CFD7F495CEE8F18C5BD0A8B8281" ID-PubMed="35087300" ID-ZooBank="8ED559A3F2D444F88A13AF880D6B5A6C" ModsDocID="1313-2970-1081-137" checkinTime="1642466624738" checkinUser="pensoft" docAuthor="Rasoamanana, Nicole & Fisher, Brian L." docDate="2022" docId="8F479C2D6B0958D4845A5D9316B32BA8" docLanguage="en" docName="ZooKeys 1081: 137-231" docOrigin="ZooKeys 1081" docPubDate="2022-01-17" docSource="http://dx.doi.org/10.3897/zookeys.1081.71872" docTitle="Camponotus jjacquia Rasoamanana & Fisher 2022, sp. nov." docType="treatment" docUuid="7B513036-F4AC-45A4-8B70-5FCCAF2C534D" docUuidSource="ZooBank" docVersion="4" id="B8C62CFD7F495CEE8F18C5BD0A8B8281" lastPageNumber="137" masterDocId="B8C62CFD7F495CEE8F18C5BD0A8B8281" masterDocTitle="A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses" masterLastPageNumber="231" masterPageNumber="137" pageNumber="137" updateTime="1668151311846" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasoamanana, Nicole</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9702-7231</mods:nameIdentifier>
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<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar</mods:affiliation>
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<mods:nameIdentifier type="email">rsoanarivo35@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-01-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1081</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>137</mods:start>
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<mods:end>231</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1081.71872</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1081.71872</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1081-137</mods:identifier>
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<mods:identifier type="ZooBank">8ED559A3F2D444F88A13AF880D6B5A6C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B8C62CFD7F495CEE8F18C5BD0A8B8281</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="192358981" LSID="urn:lsid:zoobank.org:act:7B513036-F4AC-45A4-8B70-5FCCAF2C534D" httpUri="http://treatment.plazi.org/id/8F479C2D6B0958D4845A5D9316B32BA8" lastPageNumber="137" pageId="0" pageNumber="137">
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<subSubSection pageId="0" pageNumber="137" type="nomenclature">
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<paragraph pageId="0" pageNumber="137">
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<taxonomicName LSID="http://zoobank.org/7B513036-F4AC-45A4-8B70-5FCCAF2C534D" authority="Rasoamanana & Fisher, 2022" authorityName="Rasoamanana & Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus jjacquia" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="jjacquia" status="sp. nov.">Camponotus jjacquia</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="description">
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<paragraph pageId="0" pageNumber="137">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological character of minor and major workers of C. repens species group A, B minor worker of Camponotus repens (CASENT 0217300), head in full-face view and body in lateral view C head of major worker of C. jjacquia (CASENT 0217295) D minor worker of C. jjacquia (CASENT 0445276)." figureDoi="10.3897/zookeys.1081.71872.figure10" httpUri="https://binary.pensoft.net/fig/634929" pageId="0" pageNumber="137">Figures 10C, D</figureCitation>
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<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Mesosoma in lateral view A C. Camponotus jjacquia (CASENT 0445276) B C. tsimelahy (CASENT 0446651)." figureDoi="10.3897/zookeys.1081.71872.figure45" httpUri="https://binary.pensoft.net/fig/634964" pageId="0" pageNumber="137">, 45A</figureCitation>
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<figureCitation captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Camponotus jjacquia minor worker (CASENT 0445276) 1 body in lateral view 2 head in full-face view 3 body in dorsal view." figureDoi="10.3897/zookeys.1081.71872.figure66" httpUri="https://binary.pensoft.net/fig/634985" pageId="0" pageNumber="137">, 66</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="holotype worker">
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<paragraph pageId="0" pageNumber="137">Holotype worker.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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Madagascar, Province Toliara, Parc National de Tsimanampetsotsa,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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de Bemanateza, 20.7 km 81°E Efoetse, 23.0 km 131°SE Beheloka,
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<geoCoordinate degrees="23.99222" direction="south" orientation="latitude" precision="1" value="-23.99222">-23.99222</geoCoordinate>
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,
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<geoCoordinate degrees="43.88067" direction="east" orientation="longitude" precision="1" value="43.88067">43.88067</geoCoordinate>
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, 90 m, spiny forest/thicket, under stone, 22-26 March 2002 (BLF), collection code: BLF06310, specimen code: CASENT0445276 (CAS).
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<emphasis bold="true" pageId="0" pageNumber="137">Paratypes.</emphasis>
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Four workers with the same data as holotype CASENT0445284, CASENT0445285, CASENT0445287, CASENT0445288 (CAS).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="worker diagnosis">
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<paragraph pageId="0" pageNumber="137">Worker diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="137">Integument black; entire body with plentiful, erect setae; anterior clypeal margin produced into a rounded triangular lobe; dorsal outline of mesosoma slightly impressed at the level of metanotum; petiole long and low.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="description">
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<paragraph pageId="0" pageNumber="137">Description of minor worker.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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Medium-sized. Absolute cephalic size (CS: 0.99
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.14; 0.91-1.09). In full-face view, head almost quadrate (CWb/CL: 0.32
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02; 0.31-0.34); posterior margin evenly straight, lateral margin of head anterior to eyes slightly convex. Eyes circular, placed above midline of head (PoOC/CL: 0.10
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02; 0.09-0.12). Mandibles triangular with six teeth. Clypeus carinate, its anterior border angularly produced in the middle (ClyL/GPD: 0.20
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.06, 0.16-0.23). Antennal scape long, surpassing the occiput by the length of one basal funiculus (SL/CS: 0.34
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02; 0.32-0.34). In lateral view, anterodorsal corner of pronotum rounded, mesonotum dorsum inclined toward nearly straight propodeum face (MW/ML: 0.19
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02, 0.18-0.20; MPH/ML: 0.13
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02, 0.12-0.14). In lateral view, petiolar node longer than wide, with flat summit, its anterior face leaning forward. Ants finely foveolate throughout, apparently shiny. Entire body, including mandibles and appendages, covered with plentiful, erect, whitish setae, uneven in length, some long. Head, mesosoma, and gaster dark brown to black; antennae, mandibles, tibiae, and tarsi reddish brown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="description">
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<paragraph pageId="0" pageNumber="137">Description of major worker.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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Characteristics of minor workers, except: head much wider posteriorly (CS: 1.68
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.07, 1.61-1.76; CWb/CL: 0.91
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02, 0.89-0.92); lateral margins convex and converging towards base of mandible. Eyes circular placed dorsolaterally midway from the occipital margin (PoOC/CL: 0.30
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.00, 0.29-0.30). Clypeus with a short rectangular lobe with rounded lateral angle (ClyL/GPD: 0.85
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.03, 0.81-0.88). Antennal scape short, not reaching the occipital margin (SL/CS: 0.57
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02, 0.56-0.59). Dorsal outline of mesosoma interrupted by sutures, metanotal suture forms a canal. Head dorsum and mesosoma coarsely reticulate-punctate, petiole finely striolate, gaster imbricate and shiny.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="distribution">
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<paragraph pageId="0" pageNumber="137">Distribution and biology.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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The known elevational range of
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<taxonomicName authorityName="Rasoamanana & Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus jjacquia" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="jjacquia">
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<emphasis italics="true" pageId="0" pageNumber="137">Camponotus jjacquia</emphasis>
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</taxonomicName>
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extends from 43 to 923 m. Specimens have been collected on tree trunks, on low vegetation, in leaf litter, and under stones in
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<emphasis italics="true" pageId="0" pageNumber="137">Bismarckia</emphasis>
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woodland, deciduous forest, gallery forest, savannah woodland, spiny forest, and tropical dry forest (Fig.
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<figureCitation captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Distribution map of C. repens species group A C. claveri B C. ihazofotsy C C. jjacquia D C. tsimelahy and C. robustus species group E C. zoro" figureDoi="10.3897/zookeys.1081.71872.figure72" httpUri="https://binary.pensoft.net/fig/634991" pageId="0" pageNumber="137">72C</figureCitation>
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). This species nests in the soil.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="discussion">
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<paragraph pageId="0" pageNumber="137">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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The unique petiolar shape, the density of its pilosity, and the shape of its metanotal dorsum make
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<taxonomicName lsidName="C. jjacquia" pageId="0" pageNumber="137" rank="species" species="jjacquia">
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<emphasis italics="true" pageId="0" pageNumber="137">C. jjacquia</emphasis>
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</taxonomicName>
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a very distinctive species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="etymology">
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<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="137">The new species is dedicated to Jean Jacques Rafanomezantsoa, who has been a Malagasy ant collector for more than twenty years.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="materials_examined">
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<paragraph pageId="0" pageNumber="137">Additional material examined.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" pageId="0" pageNumber="137">Province Fianarantsoa</emphasis>
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:
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Isalo IV National Park, Ampotoampoto IV National Park, Andohaela National Park Parcelle II, de Manatalinjo, B. L. Fisher, F. A. Esteves, Foret d' Ambohibory, de Bemanateza, SE Beheloka, Parcel I, R. Harin'Hala, Parcel II, M. E. Irwin, F. D. Parker" county="Tsimelahy" elevation="867" latitude="-24.93683" location="Ampotoampoto" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" specimenCount="1">
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||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:1B953F4A53FA8857460765BE53B96BF3" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Ampotoampoto">Ampotoampoto</location>
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||
III
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||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:F4904CAFD14A53C554D2DB6C97C69312" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="National Park">National Park</location>
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||
,
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||
<locationDeviation location="Ilakaka">
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||
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.91" unit="km" value="7.91">7.91 km</quantity>
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||
NW Ilakaka
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||
</locationDeviation>
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||
,
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||
<geoCoordinate degrees="22.62944" direction="south" orientation="latitude" precision="1" value="-22.62944">-22.62944</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="45.189" direction="east" orientation="longitude" precision="55" value="45.189">45.189</geoCoordinate>
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||
, 919 m, savannah woodland (ARA) (CAS);
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<collectorName>Isalo IV National Park</collectorName>
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,
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||
<locationDeviation location="Ranohira">
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||
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
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SW Ranohira
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||
</locationDeviation>
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||
,
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||
<geoCoordinate degrees="22.61472" direction="south" orientation="latitude" precision="1" value="-22.61472">-22.61472</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="45.31304" direction="east" orientation="longitude" precision="1" value="45.31304">45.31304</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">867 m</elevation>
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||
</quantity>
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, Bismarckia woodland (ARA) (CAS);
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<collectorName>Ampotoampoto IV National Park</collectorName>
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,
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||
<locationDeviation location="Ilakaka">
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||
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.83" unit="km" value="7.83">7.83 km</quantity>
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||
NW Ilakaka
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||
</locationDeviation>
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||
,
|
||
<geoCoordinate degrees="22.62944" direction="south" orientation="latitude" precision="1" value="-22.62944">-22.62944</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="45.1912" direction="east" orientation="longitude" precision="5" value="45.1912">45.1912</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.23" unit="m" value="923.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.23" unit="m" value="923.0">923 m</elevation>
|
||
</quantity>
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, savanna woodland (ARA) (CAS);
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||
<emphasis bold="true" pageId="0" pageNumber="137">Province Mahajanga</emphasis>
|
||
:
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||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:4CA83ADAACBAA5297D655ADB32FA4104" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Ampijoroa National Park">Ampijoroa National Park</location>
|
||
,
|
||
<locationDeviation location="Maevatanana">
|
||
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.6" unit="km" value="160.0">160 km</quantity>
|
||
N Maevatanana
|
||
</locationDeviation>
|
||
, deciduous forest,
|
||
<geoCoordinate degrees="16.31944" direction="south" orientation="latitude" precision="1" value="-16.31944">-16.31944</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.81333" direction="east" orientation="longitude" precision="1" value="46.81333">46.81333</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="4.3" unit="m" value="43.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="4.3" unit="m" value="43.0">43 m</elevation>
|
||
</quantity>
|
||
, deciduous forest (MG) (CAS).
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Province Toliara</emphasis>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:B80D288631E06FAE9A4CB70C772CAD57" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Anosy Region">Anosy Region</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:4EE6DE1DE37CB3AF89487B4ADDECDF4A" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="District of Amboasary">District of Amboasary</location>
|
||
,
|
||
<locationDeviation location="Fort Dauphin">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.8" unit="km" value="58.0">58 km</quantity>
|
||
SW of Fort Dauphin
|
||
</locationDeviation>
|
||
,
|
||
<locationDeviation location="Amboasary">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.0" unit="km" value="8.0">8 km</quantity>
|
||
NW of Amboasary
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:31620E6B4422314B6F106DA81DEA7EFA" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Berenty Special Reserve">Berenty Special Reserve</location>
|
||
,
|
||
<geoCoordinate degrees="25.00667" direction="south" orientation="latitude" precision="1" value="-25.00667">-25.00667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.30333" direction="east" orientation="longitude" precision="1" value="46.30333">46.30333</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">85 m</elevation>
|
||
</quantity>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:E9A6CAEB68383A25E426DC1907BB6BEE" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Galery forest">Galery forest</location>
|
||
(MG) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:18F23029089D8FAAC2BC3EB6BCF34960" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Anosy Region">Anosy Region</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:7C6B9D099EA2D370A0C8AC1A79CD0CFF" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="District of Fort-Dauphin">District of Fort-Dauphin</location>
|
||
,
|
||
<collectorName>
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:9D709DE2CB1F6F7F7CB496F936D9C126" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Andohaela National Park">Andohaela National Park</location>
|
||
Parcelle II
|
||
</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:707312C7EBCD0491C6514E4FDD6F179C" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Tsimelahy">Tsimelahy</location>
|
||
,
|
||
<locationDeviation location="Fort-Dauphin">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.2" unit="km" value="42.0">42 km</quantity>
|
||
W of Fort-Dauphin
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate degrees="24.93683" direction="south" orientation="latitude" precision="1" value="-24.93683">-24.93683</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.62667" direction="east" orientation="longitude" precision="1" value="46.62667">46.62667</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.76" unit="m" value="176.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.76" unit="m" value="176.0">176 m</elevation>
|
||
</quantity>
|
||
, transition forest (Mike Irwin, Frank Parker,
|
||
<normalizedToken originalValue="Rin’ha">Rin'ha</normalizedToken>
|
||
) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:BCA3FE614B305CA1B901C3C4A6DF1E74" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Anosy Region">Anosy Region</location>
|
||
, Parc National
|
||
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:65F8D941C8B6985CEF302E3E839464A5" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Foret de Manatalinjo">
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<collectorName>de Manatalinjo</collectorName>
|
||
</location>
|
||
,
|
||
<geoCoordinate degrees="24.82505" direction="south" orientation="latitude" precision="1" value="-24.82505">-24.82505</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.57811" direction="east" orientation="longitude" precision="1" value="46.57811">46.57811</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
|
||
</quantity>
|
||
, spiny forest/thicket (
|
||
<collectorName>B.L. Fisher</collectorName>
|
||
,
|
||
<collectorName>F.A. Esteves</collectorName>
|
||
et al.) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:9A603B882FDECE3E329E389D14A62051" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Atsimo Andrefana Region">Atsimo Andrefana Region</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:72242D45B68F9A77D00B8F2BD7A174F7" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="District of Betioky">District of Betioky</location>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:CABB73F043DB2EE02040C845A163CBF7" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Beza Mahafaly Special reserve">Beza Mahafaly Special reserve</location>
|
||
, Parcelle Belle vue
|
||
<locationDeviation location="Research Station">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
|
||
W of Research Station
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate degrees="23.68983" direction="south" orientation="latitude" precision="1" value="-23.68983">-23.68983</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44.5755" direction="east" orientation="longitude" precision="5" value="44.5755">44.5755</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.77" unit="m" value="177.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.77" unit="m" value="177.0">177 m</elevation>
|
||
</quantity>
|
||
, spiny forest (
|
||
<normalizedToken originalValue="Rin’ha">Rin'ha</normalizedToken>
|
||
) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:02B7C1DE31F04A9A8B9CD72E6A5C4E1E" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Atsimo Andrefana Region">Atsimo Andrefana Region</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:C466C29E36ED212908887228BBB1F131" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="District of Betioky">District of Betioky</location>
|
||
,
|
||
<locationDeviation location="Betioky">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
|
||
E Betioky
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:3C669D8301CFA112F8831737C1DE3898" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Beza Mahafaly Special Reserve">Beza Mahafaly Special Reserve</location>
|
||
(around
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:F047ADE8E1BBBA5F52B0A52E26F8BCBA" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Research Station">Research Station</location>
|
||
),
|
||
<geoCoordinate degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
|
||
, 165 m, gallery dry deciduous forest (MG); Beza-Mahafaly,
|
||
<locationDeviation location="Betioky">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.7" unit="km" value="27.0">27 km</quantity>
|
||
E Betioky
|
||
</locationDeviation>
|
||
, -23.65,4 4.63333,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">135 m</elevation>
|
||
</quantity>
|
||
, tropical dry forest (
|
||
<collectorName>B.L. Fisher</collectorName>
|
||
) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:E8EADEE57F533D8C3E09D0EE5BA5C894" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Foret de Mahavelo">
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
de Mahavelo
|
||
</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:0B484E7748E2ABB598AC18EDE360E1A2" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Isantoria River">Isantoria River</location>
|
||
,
|
||
<geoCoordinate degrees="24.75833" direction="south" orientation="latitude" precision="1" value="-24.75833">-24.75833</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.15717" direction="east" orientation="longitude" precision="1" value="46.15717">46.15717</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">110 m</elevation>
|
||
</quantity>
|
||
, spiny forest/thicket (FGAT) (CAS); Parc National
|
||
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
|
||
,
|
||
<collectorName>
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken originalValue="d’">d'</normalizedToken>
|
||
Ambohibory
|
||
</collectorName>
|
||
,
|
||
<locationDeviation location="Tsimelahy">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="km" value="1.7">1.7 km</quantity>
|
||
61° ENE Tsimelahy
|
||
</locationDeviation>
|
||
,
|
||
<locationDeviation location="Tolagnaro">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.61" unit="km" value="36.1">36.1 km</quantity>
|
||
308° NW Tolagnaro
|
||
</locationDeviation>
|
||
, -24.93, 46.6455,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
, tropical dry forest (FGAT) (CAS); Parc National
|
||
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:49C69A69B379D91F6CDD3B7A8DB2DA36" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Foret de Manatalinjo">
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<collectorName>de Manatalinjo</collectorName>
|
||
</location>
|
||
,
|
||
<locationDeviation location="Amboasary">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.36" unit="km" value="33.6">33.6 km</quantity>
|
||
63° ENE Amboasary
|
||
</locationDeviation>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.6" unit="km" value="7.6">7.6 km</quantity>
|
||
<geoCoordinate degrees="99" direction="east" orientation="longitude" precision="55555" value="99.0">99° E</geoCoordinate>
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:CF4E7F6EDC669358FFBA9E45435F22B4" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Hazofotsy">Hazofotsy</location>
|
||
, -24.81694, 46.61,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</elevation>
|
||
</quantity>
|
||
, spiny forest/thicket (FGAT) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:64BC41406789949EF8BC882D33C36A66" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Parc National de Kirindy Mite">Parc National de Kirindy Mite</location>
|
||
,
|
||
<locationDeviation location="Belo sur mer">
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.63" unit="km" value="16.3">16.3 km</quantity>
|
||
127° SE Belo sur mer
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate degrees="20.79528" direction="south" orientation="latitude" precision="1" value="-20.79528">-20.79528</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44.147" direction="east" orientation="longitude" precision="55" value="44.147">44.147</geoCoordinate>
|
||
, 80 m, tropical dry forest (FGAT) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:186FA8B8B075EE421C6F885BBBD844E0" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Parc National de Tsimanampetsotsa">Parc National de Tsimanampetsotsa</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:9EFC43288568BC2D3818E815559C7517" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Foret de Bemanateza">
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<collectorName>de Bemanateza</collectorName>
|
||
</location>
|
||
,
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.07" unit="km" value="20.7">20.7 km</quantity>
|
||
<geoCoordinate degrees="81" direction="east" orientation="longitude" precision="55555" value="81.0">81° E</geoCoordinate>
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:FB59F479AC7F63D2272A8EFA3BD2B3AE" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Efoetse">Efoetse</location>
|
||
, 23.0 km 131°
|
||
<collectorName>SE Beheloka</collectorName>
|
||
,
|
||
<geoCoordinate degrees="23.99222" direction="south" orientation="latitude" precision="1" value="-23.99222">-23.99222</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="43.88067" direction="east" orientation="longitude" precision="1" value="43.88067">43.88067</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
|
||
</quantity>
|
||
, spiny forest/thicket (FGAT) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:BC04DBCA64B7D9194EAB536B3362421E" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Beza Mahafaly Reserve">Beza Mahafaly Reserve</location>
|
||
, near research station,
|
||
<collectorName>Parcel I</collectorName>
|
||
,
|
||
<geoCoordinate degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
|
||
, 165 m, dry deciduous forest (
|
||
<collectorName>
|
||
R.
|
||
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||
</collectorName>
|
||
) (CAS);
|
||
<location LSID="urn:lsid:plazi:treatment:8F479C2D6B0958D4845A5D9316B32BA8:81EC009DED9E6425860C4B5D18BB60B1" county="Tsimelahy" latitude="-24.93683" longLatPrecision="1" longitude="46.62667" municipality="Andohahela National Park" name="Beza Mahafaly Reserve">Beza Mahafaly Reserve</location>
|
||
, near Bellevue,
|
||
<collectorName>Parcel II</collectorName>
|
||
,
|
||
<geoCoordinate degrees="23.68983" direction="south" orientation="latitude" precision="1" value="-23.68983">-23.68983</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44.5755" direction="east" orientation="longitude" precision="5" value="44.5755">44.5755</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">180 m</elevation>
|
||
</quantity>
|
||
, spiny forest (
|
||
<collectorName>
|
||
R.
|
||
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||
</collectorName>
|
||
) (CAS);
|
||
<collectingMunicipality>Andohahela National Park</collectingMunicipality>
|
||
,
|
||
<collectingCounty>Tsimelahy</collectingCounty>
|
||
-
|
||
<collectorName>Parcel II</collectorName>
|
||
, transitional forest,
|
||
<geoCoordinate degrees="24.93683" direction="south" orientation="latitude" precision="1" value="-24.93683">-24.93683</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.62667" direction="east" orientation="longitude" precision="1" value="46.62667">46.62667</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">180 m</elevation>
|
||
</quantity>
|
||
, transitional forest (
|
||
<collectorName>M.E. Irwin</collectorName>
|
||
,
|
||
<collectorName>F.D. Parker</collectorName>
|
||
,
|
||
<collectorName>
|
||
R.
|
||
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||
</collectorName>
|
||
) (CAS)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1081.71872.figure66" httpUri="https://binary.pensoft.net/fig/634985" pageId="0" pageNumber="137" start="Figure 66" startId="F66">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 66.</emphasis>
|
||
<taxonomicName authorityName="Rasoamanana & Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus jjacquia" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="jjacquia">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus jjacquia</emphasis>
|
||
</taxonomicName>
|
||
minor worker (CASENT0445276)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">1</emphasis>
|
||
body in lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">2</emphasis>
|
||
head in full-face view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">3</emphasis>
|
||
body in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |