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<document id="6074764781DDF2888024921BC8A48A64" ID-CLB-Dataset="37318" ID-DOI="10.5281/zenodo.245827" ID-GBIF-Dataset="69cf893c-7b35-48e8-ad78-d9775626f287" ID-ISSN="1175-5326" ID-Zenodo-Dep="245827" ID-ZooBank="9345C596-8656-4B5C-AD8C-2FACF4E9240C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484506177574" checkinUser="GgServerImporter" docAuthor="Blake, James A." docDate="2017" docId="8F2387DD06530945FF31FEF6FBEDFEDC" docLanguage="en" docName="Zootaxa.4218.1.001-145" docOrigin="Zootaxa 4218 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leodamas cochleatus Ehlers 1900" docType="treatment" docVersion="14" lastPageNumber="73" masterDocId="731AFFA50615090DFFA6FFEBFFDCFFC0" masterDocTitle="Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America" masterLastPageNumber="145" masterPageNumber="1" pageNumber="71" updateTime="1698435671898" updateUser="ExternalLinkService">
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<mods:title id="B6BACB57FEC6694B29B90F51613D43D5">Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America</mods:title>
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<mods:namePart id="5A9DC7CE52C4E9B747E2DC38A4A7392E">Blake, James A.</mods:namePart>
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<treatment id="8F2387DD06530945FF31FEF6FBEDFEDC" ID-DOI="http://doi.org/10.5281/zenodo.4901783" ID-GBIF-Taxon="127422014" ID-Zenodo-Dep="4901783" LSID="urn:lsid:plazi:treatment:8F2387DD06530945FF31FEF6FBEDFEDC" httpUri="http://treatment.plazi.org/id/8F2387DD06530945FF31FEF6FBEDFEDC" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71">
<subSubSection id="4F9065400653094BFF31FEF6FEE4FE98" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="073536CB0653094BFF31FEF6FD36FEF7" blockId="70.[151,746,285,344]" box="[151,746,285,311]" pageId="70" pageNumber="71">
<heading id="5C7D81A70653094BFF31FEF6FD36FEF7" bold="true" box="[151,746,285,311]" fontSize="11" level="1" pageId="70" pageNumber="71" reason="1">
<emphasis id="35FEEAD90653094BFF31FEF6FD36FEF7" bold="true" box="[151,746,285,311]" pageId="70" pageNumber="71">
<taxonomicName id="C08A4D480653094BFF31FEF6FD8DFEF7" ID-CoL="3T6BB" authority="Ehlers, 1900" authorityName="Ehlers" authorityYear="1900" box="[151,593,285,311]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90653094BFF31FEF6FE46FEF7" bold="true" box="[151,410,285,311]" italics="true" pageId="70" pageNumber="71">Leodamas cochleatus</emphasis>
(
<bibRefCitation id="631B4B3A0653094BFE0CFEF6FD96FEF7" author="Ehlers" box="[426,586,285,311]" pageId="70" pageNumber="71" refString="Ehlers, E. (1900) Magellanische Anneliden gesammelt wahrend der schwedischen Expedition nach den Magellanslander. Nachrichten von der Gesellschaft der Wissenschaften zu Gottingen, Mathematisch-Physikalische Klasse, 1900, 206 - 223." type="journal article" year="1900">Ehlers, 1900</bibRefCitation>
)
</taxonomicName>
. New Status
</emphasis>
</heading>
</paragraph>
<paragraph id="073536CB0653094BFF31FED4FEE4FE98" blockId="70.[151,746,285,344]" box="[151,312,319,344]" pageId="70" pageNumber="71">
<figureCitation id="9FB12A4E0653094BFF31FED4FEE4FE98" box="[151,312,319,344]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="71.[151,249,1820,1842]" captionStartId-1="72.[151,249,1512,1534]" captionTargetBox-0="[168,1419,709,1773]" captionTargetBox-1="[159,1431,347,1487]" captionTargetId-0="figure@71.[151,1436,697,1798]" captionTargetId-1="figure@72.[151,1436,339,1491]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." captionText-1="FIGURE 32. Leodamas cochleatus (Ehlers, 1900). Phase Contrast images from Argentinean specimens (AHF Poly 5043): A B, anterior thoracic uncini from setiger 2; C, detail of anterior thoracic uncinus showing longitudinal notch; D E, posterior thoracic uncini from setiger 8 showing expanded apex and notch; F, abdominal notopodial spine; G, abdominal neuropodium with spine and capillaries; H, abdominal notosetae including capillaries, spines (broken) and furcate seta." httpUri-0="https://zenodo.org/record/245857/files/figure.png" httpUri-1="https://zenodo.org/record/245858/files/figure.png" pageId="70" pageNumber="71">Figures 3132</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F9065400653094BFF31FE62FCB8FE5E" box="[151,868,392,414]" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="073536CB0653094BFF31FE62FCB8FE5E" blockId="70.[151,868,392,414]" box="[151,868,392,414]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="279A11E50653094BFF31FE62FCB8FE5E" box="[151,868,392,414]" pageId="70" pageNumber="71">
<taxonomicName id="C08A4D480653094BFF31FE62FE29FE5E" ID-CoL="67QXC" authority="Ehlers, 1900:217" authorityName="Ehlers" authorityPageNumber="217" authorityYear="1900" box="[151,501,392,414]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="35FEEAD90653094BFF31FE62FEE3FE5E" box="[151,319,392,414]" italics="true" pageId="70" pageNumber="71">Aricia cochleata</emphasis>
<treatmentCitation id="862B10DA0653094BFEE3FE63FE29FE5E" author="Ehlers" box="[325,501,392,414]" page="217" pageId="70" pageNumber="71" year="1900">
<bibRefCitation id="631B4B3A0653094BFEE3FE63FE15FE5E" author="Ehlers" box="[325,457,392,414]" pageId="70" pageNumber="71" refString="Ehlers, E. (1900) Magellanische Anneliden gesammelt wahrend der schwedischen Expedition nach den Magellanslander. Nachrichten von der Gesellschaft der Wissenschaften zu Gottingen, Mathematisch-Physikalische Klasse, 1900, 206 - 223." type="journal article" year="1900">Ehlers, 1900</bibRefCitation>
:217
</treatmentCitation>
</taxonomicName>
; 1901: 166167, pl. 21, figs. 1421.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4F9065400653094BFF31FE21FB78FDEB" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="073536CB0653094BFF31FE21FB78FDEB" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
<materialsCitation id="B7E23C960653094BFF31FE21FF0DFDEB" ID-GBIF-Occurrence="1424727936" collectingDate="1917-05" collectionCode="SMNH" collectorName="E. Nordenskold &amp; Plata, R" country="Chile" elevation="46" latitude="-52.633335" location="Chile" longLatPrecision="1082" longitude="-70.0" pageId="70" pageNumber="71" specimenCode="SMNH 549" specimenCount="1" typeStatus="holotype">
<emphasis id="35FEEAD90653094BFF31FE21FEA6FE23" bold="true" box="[151,378,458,483]" pageId="70" pageNumber="71">Material examined</emphasis>
.
<collectingCountry id="7F9D765B0653094BFE2CFE21FE15FE23" box="[394,457,458,483]" name="Chile" pageId="70" pageNumber="71">Chile</collectingCountry>
, Straits of Magellan, Tribune Bank,
<geoCoordinate id="62BE500C0653094BFCC1FE21FC1FFE23" box="[871,963,458,483]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="925" value="-52.633335">52°38S</geoCoordinate>
,
<geoCoordinate id="62BE500C0653094BFC76FE21FBE8FE23" box="[976,1076,458,483]" direction="west" orientation="longitude" pageId="70" pageNumber="71" precision="925" value="-70.0">70°00W</geoCoordinate>
,
<quantity id="C0729B2E0653094BFBE6FE21FBA0FE22" box="[1088,1148,458,483]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="70" pageNumber="71" unit="m" value="46.0">
<elevation id="8CA7D1F80653094BFBE6FE21FBA0FE22" box="[1088,1148,458,483]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="70" pageNumber="71" unit="m" value="46.0">46 m</elevation>
</quantity>
, sand and gravel, coll.
<collectorName id="AA7F531D0653094BFA22FE20FEF7FDC7" pageId="70" pageNumber="71">E. Nordenskold</collectorName>
,
<emphasis id="35FEEAD90653094BFE90FE05FE41FDC7" bold="true" box="[310,413,494,519]" pageId="70" pageNumber="71">
<typeStatus id="D83188690653094BFE90FE05FE41FDC7" box="[310,413,494,519]" pageId="70" pageNumber="71" type="holotype">holotype</typeStatus>
</emphasis>
of
<taxonomicName id="C08A4D480653094BFE62FE04FDA3FDC6" box="[452,639,494,518]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="35FEEAD90653094BFE62FE04FDA3FDC6" box="[452,639,494,518]" italics="true" pageId="70" pageNumber="71">Aricia cochleata</emphasis>
</taxonomicName>
(
<specimenCode id="572C9EB00653094BFD28FE04FCCFFDC7" box="[654,787,494,519]" collectionCode="SMNH" country="Canada" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="70" pageNumber="71">
<collectionCode id="619BAE0E0653094BFD28FE04FD3DFDC7" box="[654,737,495,519]" country="Canada" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="70" pageNumber="71">SMNH</collectionCode>
549
</specimenCode>
).—Off
<collectingCountry id="7F9D765B0653094BFCCBFE05FC03FDC7" box="[877,991,494,519]" name="Argentina" pageId="70" pageNumber="71">Argentina</collectingCountry>
, E of Mar del
<collectorName id="AA7F531D0653094BFB22FE05FB02FDC7" box="[1156,1246,494,519]" pageId="70" pageNumber="71">Plata, R</collectorName>
/V
<taxonomicName id="C08A4D480653094BFAA4FE04FAE2FDC6" box="[1282,1342,495,518]" class="Monoplacophora" family="Neopilinidae" genus="Vema" kingdom="Animalia" order="Tryblidiida" pageId="70" pageNumber="71" phylum="Mollusca" rank="genus">
<emphasis id="35FEEAD90653094BFAA4FE04FAE2FDC6" box="[1282,1342,495,518]" italics="true" pageId="70" pageNumber="71">Vema</emphasis>
</taxonomicName>
Station,
<date id="7334100B0653094BFF31FDF8FF0DFDEB" box="[151,209,531,555]" pageId="70" pageNumber="71" value="1917-05">
<collectingDate id="6370E9E30653094BFF31FDF8FF0DFDEB" box="[151,209,531,555]" pageId="70" pageNumber="71" value="1917-05">V-17</collectingDate>
</date>
</materialsCitation>
-101, 450
<quantity id="C0729B2E0653094BFEE1FDF9FE4CFDEA" box="[327,400,530,555]" metricMagnitude="2" metricUnit="m" metricValue="4.54" pageId="70" pageNumber="71" unit="m" value="454.0">454 m</quantity>
,
<date id="7334100B0653094BFE3AFDF8FDF9FDEB" box="[412,549,530,555]" pageId="70" pageNumber="71" value="1961-06-19">19 Jun 1961</date>
(5, LACM-AHF Poly 5043; 3, LACM-AHF Poly 5045).
</paragraph>
</subSubSection>
<subSubSection id="4F9065400653094BFF61FDDDFEA7F9DB" pageId="70" pageNumber="71" type="description">
<paragraph id="073536CB0653094BFF61FDDDFAB5FD1F" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
<emphasis id="35FEEAD90653094BFF61FDDDFE85FD8F" bold="true" box="[199,345,566,591]" pageId="70" pageNumber="71">Description.</emphasis>
<typeStatus id="D83188690653094BFEC7FDDDFE15FD8F" box="[353,457,566,591]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
(SMNH 549) incomplete,
<quantity id="C0729B2E0653094BFD59FDDCFC8CFD8E" box="[767,848,567,591]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="70" pageNumber="71" unit="mm" value="32.0">32 mm</quantity>
long,
<quantity id="C0729B2E0653094BFC3DFDDCFC02FD8E" box="[923,990,567,591]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="70" pageNumber="71" unit="mm" value="2.0">2 mm</quantity>
wide for 110 setigers, color in alcohol pale yellow. Argentinean specimens all incomplete, smaller,
<quantity id="C0729B2E0653094BFCEEFDB0FC68FDB2" box="[840,948,603,627]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="70" pageNumber="71" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long,
<quantity id="C0729B2E0653094BFC5BFDB0FBADFDB2" box="[1021,1137,603,627]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="70" pageNumber="71" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
wide, with 2430 setigers (LACM-AHF Poly 5043); juveniles also present, one mostly complete,
<quantity id="C0729B2E0653094BFC41FD95FB9EFD56" box="[999,1090,638,663]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="70" pageNumber="71" unit="mm" value="4.5">4.5 mm</quantity>
long,
<quantity id="C0729B2E0653094BFB35FD94FB32FD56" box="[1171,1262,639,663]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="70" pageNumber="71" unit="mm" value="0.3">0.3 mm</quantity>
wide, with 40 setigers (LACM-AHF Poly 5045), color in alcohol tan. Body of all specimens flattened dorsally throughout, rounded ventrally, more so in the abdominal segments; body widest in thoracic segments, tapering posteriorly.
</paragraph>
<paragraph id="073536CB0653094BFF61FD01FB91FC8B" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
Prostomium triangular in outline, tapering anteriorly to narrow, blunt tip (
<figureCitation id="9FB12A4E0653094BFBA8FD01FB87FCC3" box="[1038,1115,746,771]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Fig 31</figureCitation>
A); without eyespots; paired nuchal organs narrow slits, dorsolateral at border with peristomium. Peristomium with one achaetous ring only vaguely separated from prostomium, but well separated from first setiger (
<figureCitation id="9FB12A4E0653094BFC7EFCD9FBF6FC8B" box="[984,1066,818,843]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Fig. 31</figureCitation>
A).
</paragraph>
<paragraph id="073536CB0653094BFF61FCBDFCFBFBAB" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
<typeStatus id="D83188690653094BFF61FCBDFEF3FCAF" box="[199,303,854,879]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
with 29 setigers in thoracic region; smaller Argentinean specimens with 1011 thoracic setigers. Notopodial postsetal lobes from setiger 1, fingerlike in shape, well developed from the first, increasing in size over thoracic region (
<figureCitation id="9FB12A4E0653094BFEFCFC75FE6CFC77" box="[346,432,926,951]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Fig. 31</figureCitation>
A), continuing through abdominal segments. Thoracic neuropodia elongate thickened lobes, with no postsetal lobe or lamella present; thoracic uncini arranged in three elongate rows, with the first appearing to curve ventrally forming a fourth row behind the third, similar to that of
<taxonomicName id="C08A4D480653094BFBEAFC0CFB34FC3E" box="[1100,1256,999,1022]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD90653094BFBEAFC0CFB34FC3E" box="[1100,1256,999,1022]" italics="true" pageId="70" pageNumber="71">L. marginatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C08A4D480653094BFAB8FC0CFA49FC3E" box="[1310,1429,999,1022]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90653094BFAB8FC0CFA49FC3E" box="[1310,1429,999,1022]" italics="true" pageId="70" pageNumber="71">L. cirratus</emphasis>
</taxonomicName>
. Abdominal neuropodia thickened elongated lobes, with short ventral cirrus on apex of neuropodium and prominent cirriform subpodial lobe at base of neuropodium (
<figureCitation id="9FB12A4E0653094BFD5FFBC5FC8DFB87" box="[761,849,1070,1095]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Fig. 31</figureCitation>
B) present from first present abdominal setigers, continuing posteriorly through on all abdominal segments.
</paragraph>
<paragraph id="073536CB0653094BFF61FB9DFF04FA13" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
Thoracic notosetae including fascicles of crenulated capillaries; abdominal notosetae including capillaries, 12 straight spines with narrow pointed tips (
<figureCitation id="9FB12A4E0653094BFDC3FB71FD64FB73" box="[613,696,1178,1203]" captionStart="FIGURE 32" captionStartId="72.[151,249,1512,1534]" captionTargetBox="[159,1431,347,1487]" captionTargetId="figure@72.[151,1436,339,1491]" captionTargetPageId="72" captionText="FIGURE 32. Leodamas cochleatus (Ehlers, 1900). Phase Contrast images from Argentinean specimens (AHF Poly 5043): A B, anterior thoracic uncini from setiger 2; C, detail of anterior thoracic uncinus showing longitudinal notch; D E, posterior thoracic uncini from setiger 8 showing expanded apex and notch; F, abdominal notopodial spine; G, abdominal neuropodium with spine and capillaries; H, abdominal notosetae including capillaries, spines (broken) and furcate seta." httpUri="https://zenodo.org/record/245858/files/figure.png" pageId="70" pageNumber="71">Fig. 32</figureCitation>
F), and 13 furcate setae (
<figureCitation id="9FB12A4E0653094BFC7BFB71FBE6FB73" box="[989,1082,1178,1203]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">
Figs.
<date id="7334100B0653094BFBBBFB70FBE6FB73" box="[1053,1082,1179,1203]" pageId="70" pageNumber="71" value="1931-01">31</date>
</figureCitation>
I, 32H); furcate setae each with unequal tynes bearing fine needles; shaft smooth (
<figureCitation id="9FB12A4E0653094BFD7CFB55FCE5FB17" box="[730,825,1214,1239]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">
Figs.
<date id="7334100B0653094BFCBAFB54FCE5FB17" box="[796,825,1215,1239]" pageId="70" pageNumber="71" value="1931-01">31</date>
</figureCitation>
I, 32H). Thoracic neuropodia with uncini arranged in three long vertical rows and one short posterior row, similar to
<taxonomicName id="C08A4D480653094BFC66FB08FBBAFB3A" box="[960,1126,1251,1274]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD90653094BFC66FB08FBBAFB3A" box="[960,1126,1251,1274]" italics="true" pageId="70" pageNumber="71">L. marginatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C08A4D480653094BFB08FB08FAF2FB3A" box="[1198,1326,1251,1274]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90653094BFB08FB08FAF2FB3A" box="[1198,1326,1251,1274]" italics="true" pageId="70" pageNumber="71">L. cirratus</emphasis>
</taxonomicName>
; without accompanying capillaries; uncini of anteriormost 23 setigers with straight shaft bearing weakly developed transverse ribs and with a rounded narrowing apex; with elongate narrow groove on one side (
<figureCitation id="9FB12A4E0653094BFB67FAC1FAC0FA83" box="[1217,1308,1322,1347]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Figs. 31</figureCitation>
CE, 32A C); subsequent setigers with shaft of uncini developing prominent transverse ribs or rows of blunt barbs; with tip of uncini becoming expanded, curved, blunt, with subapical pocket or notch on concave side (
<figureCitation id="9FB12A4E0653094BFB06FA99FB27FA4B" box="[1184,1275,1394,1419]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Figs. 31</figureCitation>
FH, 32DE). Abdominal neurosetae including capillaries and 12 acicular spines with tip narrowing to curved blunt tip (
<figureCitation id="9FB12A4E0653094BFAC9FA7DFF69FA13" captionStart="FIGURE 32" captionStartId="72.[151,249,1512,1534]" captionTargetBox="[159,1431,347,1487]" captionTargetId="figure@72.[151,1436,339,1491]" captionTargetPageId="72" captionText="FIGURE 32. Leodamas cochleatus (Ehlers, 1900). Phase Contrast images from Argentinean specimens (AHF Poly 5043): A B, anterior thoracic uncini from setiger 2; C, detail of anterior thoracic uncinus showing longitudinal notch; D E, posterior thoracic uncini from setiger 8 showing expanded apex and notch; F, abdominal notopodial spine; G, abdominal neuropodium with spine and capillaries; H, abdominal notosetae including capillaries, spines (broken) and furcate seta." httpUri="https://zenodo.org/record/245858/files/figure.png" pageId="70" pageNumber="71">Fig. 32</figureCitation>
G).
</paragraph>
<paragraph id="073536CB0653094BFF61FA35FEA7F9DB" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
Branchiae from setiger 6, each broad, basally tapering to prolonged, narrow apical extension (
<figureCitation id="9FB12A4E0653094BFAA2FA35FA8BFA37" box="[1284,1367,1502,1527]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="70" pageNumber="71">Fig. 31</figureCitation>
AB). Pygidium unknown.
</paragraph>
</subSubSection>
<subSubSection id="4F90654006530945FF61F9CDFBE5FF38" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71" type="discussion">
<paragraph id="073536CB0653094BFF61F9CDFF33F82F" blockId="70.[151,1437,458,2031]" pageId="70" pageNumber="71">
<emphasis id="35FEEAD90653094BFF61F9CDFEE7F9FF" bold="true" box="[199,315,1574,1599]" pageId="70" pageNumber="71">Remarks.</emphasis>
<taxonomicName id="C08A4D480653094BFEE4F9CCFE20F9FE" box="[322,508,1574,1598]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="35FEEAD90653094BFEE4F9CCFE20F9FE" box="[322,508,1574,1598]" italics="true" pageId="70" pageNumber="71">Aricia cochleata</emphasis>
</taxonomicName>
and
<taxonomicName id="C08A4D480653094BFD95F9CCFD4FF9FE" box="[563,659,1574,1598]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="ohlini">
<emphasis id="35FEEAD90653094BFD95F9CCFD4FF9FE" box="[563,659,1574,1598]" italics="true" pageId="70" pageNumber="71">A. ohlini</emphasis>
</taxonomicName>
were both briefly described but not illustrated by
<bibRefCitation id="631B4B3A0653094BFB63F9CDFA82F9FF" author="Ehlers" box="[1221,1374,1574,1599]" pageId="70" pageNumber="71" refString="Ehlers, E. (1900) Magellanische Anneliden gesammelt wahrend der schwedischen Expedition nach den Magellanslander. Nachrichten von der Gesellschaft der Wissenschaften zu Gottingen, Mathematisch-Physikalische Klasse, 1900, 206 - 223." type="journal article" year="1900">Ehlers (1900)</bibRefCitation>
from the same locality in the Strait of Magellan and subsequently fully described and illustrated in
<bibRefCitation id="631B4B3A0653094BFB51F9A1FA45F9A3" author="Ehlers" box="[1271,1433,1610,1635]" pageId="70" pageNumber="71" refString="Ehlers, E. (1901) Die Polychaeten des magellanischen und chilenischen Strandes. Ein faunistischer Versuch. Festschrift zur Feier des Hundertfunfzigjahrigen Bestehens der koniglichen Gesellschaft der Wissenschaften zu Gottingen. pp. 1 - 232, 25 plates." type="book chapter" year="1901">Ehlers (1901)</bibRefCitation>
.
<bibRefCitation id="631B4B3A0653094BFF31F984FE90F947" author="Augener" box="[151,332,1646,1671]" pageId="70" pageNumber="71" refString="Augener, H. (1926) Polychaeten von Neuseeland. II. Sedentaria. Videnskabelige Meddelelser Naturhistorisk Forening i KObenhavn, 81, 157 - 294." type="journal article" year="1926">Augener (1926)</bibRefCitation>
synonymized
<taxonomicName id="C08A4D480653094BFE51F984FD5BF946" box="[503,647,1646,1670]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="35FEEAD90653094BFE51F984FD5BF946" box="[503,647,1646,1670]" italics="true" pageId="70" pageNumber="71">A. cochleata</emphasis>
</taxonomicName>
with
<taxonomicName id="C08A4D480653094BFD68F984FCEDF946" box="[718,817,1646,1670]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="ohlini">
<emphasis id="35FEEAD90653094BFD68F984FCEDF946" box="[718,817,1646,1670]" italics="true" pageId="70" pageNumber="71">A. ohlini</emphasis>
</taxonomicName>
based on collections from
<collectingCountry id="7F9D765B0653094BFBD5F984FACDF947" box="[1139,1297,1646,1671]" name="New Zealand" pageId="70" pageNumber="71">
New
<collectingRegion id="C54EF8290653094BFB15F985FACDF947" box="[1203,1297,1646,1671]" country="Netherlands" name="Zeeland" pageId="70" pageNumber="71">Zealand</collectingRegion>
</collectingCountry>
. He did not examine the original collections and provided little justification for the synonymy.
<bibRefCitation id="631B4B3A0653094BFB2AF978FA9DF96B" author="Hartman" box="[1164,1345,1682,1707]" pageId="70" pageNumber="71" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15 (3), 211 - 393, plates 20 - 44, 1 chart." type="journal article" year="1957">Hartman (1957</bibRefCitation>
,
<bibRefCitation id="631B4B3A0653094BFAF2F979FA4CF96B" author="Hartman" box="[1364,1424,1682,1707]" pageId="70" pageNumber="71" refString="Hartman, O. (1966) Polychaeta Myzostomidae and Sedentaria of Antarctica. Antarctic Research Series, 7, 1 - 158, 46 plates, 5 charts. American Geophysical Union, Washington, D. C." type="book chapter" year="1966">1966</bibRefCitation>
),
<bibRefCitation id="631B4B3A0653094BFF31F95DFEACF90F" author="Rozbaczylo" box="[151,368,1718,1743]" pageId="70" pageNumber="71" refString="Rozbaczylo, N. (1985) Los Anelidos Poliquetos de Chile. Indice Sinonimico y distribucion geografica de especies. Facultad de Ciencias Biologicas Pontificia Universidad Catolica de Chile, Monografias Biologicas, No. 3, 1 - 284." type="book chapter" year="1985">Rozbaczylo (1985)</bibRefCitation>
and others, however, recognized and perpetuated this synonymy with little or no discussion despite there being clear differences between the two species in Ehlers (1901) published descriptions and illustrations. Further, if these two species were synonyms, then
<taxonomicName id="C08A4D480653094BFCDAF914FBD7F8D6" box="[892,1035,1790,1814]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="35FEEAD90653094BFCDAF914FBD7F8D6" box="[892,1035,1790,1814]" italics="true" pageId="70" pageNumber="71">A. cochleata</emphasis>
</taxonomicName>
would have to be the valid name, having been described first in the same paper by
<bibRefCitation id="631B4B3A0653094BFD72F8C9FCAFF8FB" author="Ehlers" box="[724,883,1826,1851]" pageId="70" pageNumber="71" refString="Ehlers, E. (1900) Magellanische Anneliden gesammelt wahrend der schwedischen Expedition nach den Magellanslander. Nachrichten von der Gesellschaft der Wissenschaften zu Gottingen, Mathematisch-Physikalische Klasse, 1900, 206 - 223." type="journal article" year="1900">Ehlers (1900)</bibRefCitation>
. Both species names were referred to
<taxonomicName id="C08A4D480653094BFA89F8C8FEC0F89F" authority="(Leodamas)" baseAuthorityName="Leodamas" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD90653094BFA89F8C8FA40F8FB" box="[1327,1436,1827,1851]" italics="true" pageId="70" pageNumber="71">Scoloplos</emphasis>
(
<emphasis id="35FEEAD90653094BFF06F8ADFECFF89E" box="[160,275,1862,1886]" italics="true" pageId="70" pageNumber="71">Leodamas</emphasis>
)
</taxonomicName>
by
<bibRefCitation id="631B4B3A0653094BFEECF8ACFE23F89F" author="Hartman" box="[330,511,1862,1887]" pageId="70" pageNumber="71" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15 (3), 211 - 393, plates 20 - 44, 1 chart." type="journal article" year="1957">Hartman (1957)</bibRefCitation>
and are clearly species of
<taxonomicName id="C08A4D480653094BFC95F8ADFC7AF89E" box="[819,934,1862,1886]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD90653094BFC95F8ADFC7AF89E" box="[819,934,1862,1886]" italics="true" pageId="70" pageNumber="71">Leodamas</emphasis>
</taxonomicName>
as defined in the present paper. Despite the assumption of synonymy, an inspection of the descriptions of these two species suggests that they are different. This was verified by examining the
<typeStatus id="D83188690653094BFD98F864FDB2F867" box="[574,622,1935,1959]" pageId="70" pageNumber="71">type</typeStatus>
specimens and additional collections. In the present paper,
<taxonomicName id="C08A4D480653094BFA8FF865FF04F80B" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="ohlini">
<emphasis id="35FEEAD90653094BFA8FF865FF04F80B" italics="true" pageId="70" pageNumber="71">Leodamas ohlini</emphasis>
</taxonomicName>
becomes a synonym of
<taxonomicName id="C08A4D480653094BFE54F858FDB6F80A" box="[498,618,1971,1994]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90653094BFE54F858FDB6F80A" box="[498,618,1971,1994]" italics="true" pageId="70" pageNumber="71">L. cirratus</emphasis>
</taxonomicName>
and
<taxonomicName id="C08A4D480653094BFD02F858FCE0F80B" box="[676,828,1971,1995]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90653094BFD02F858FCE0F80B" box="[676,828,1971,1995]" italics="true" pageId="70" pageNumber="71">L. cochleatus</emphasis>
</taxonomicName>
is redescribed and resurrected as a distinct and valid species.
</paragraph>
<paragraph id="073536CB0652094AFF61FF7CFE6AFD44" blockId="71.[151,1436,151,645]" pageId="71" pageNumber="72">
The main difference between
<taxonomicName id="C08A4D480652094AFDBDFF73FD2CFF6F" box="[539,752,152,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90652094AFDBDFF73FD2CFF6F" box="[539,752,152,176]" italics="true" pageId="71" pageNumber="72">Leodamas cirratus</emphasis>
</taxonomicName>
(with
<taxonomicName id="C08A4D480652094AFC9AFF72FC40FF70" box="[828,924,152,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="ohlini">
<emphasis id="35FEEAD90652094AFC9AFF72FC40FF70" box="[828,924,152,176]" italics="true" pageId="71" pageNumber="72">L. ohlini</emphasis>
</taxonomicName>
as a synonym) and
<taxonomicName id="C08A4D480652094AFB22FF72FAC0FF70" box="[1156,1308,152,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90652094AFB22FF72FAC0FF70" box="[1156,1308,152,176]" italics="true" pageId="71" pageNumber="72">L. cochleatus</emphasis>
</taxonomicName>
is with the nature of the thoracic neuropodial uncini. In
<taxonomicName id="C08A4D480652094AFD35FF56FCD7FF14" box="[659,779,189,212]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90652094AFD35FF56FCD7FF14" box="[659,779,189,212]" italics="true" pageId="71" pageNumber="72">L. cirratus</emphasis>
</taxonomicName>
, these uncini are relatively simple, straight-shafted with a weakly developed subapical notch or groove; in SEM weakly developed transverse ridges were observed. In contrast, the thoracic neuropodial uncini of
<taxonomicName id="C08A4D480652094AFD2AFEEEFCF8FEDC" box="[652,804,260,284]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90652094AFD2AFEEEFCF8FEDC" box="[652,804,260,284]" italics="true" pageId="71" pageNumber="72">L. cochleatus</emphasis>
</taxonomicName>
exhibit a distinct change in structure from those of the anteriormost 23 setigers where uncini have a relatively smooth, straight shaft bearing weakly developed transverse ridges and a narrow, rounded apex and an elongate narrow groove on one side. These uncini are replaced in middle and posterior thoracic neuropodia by spines with distinct transverse ribs or rows of blunt barbs on the shaft and with the tip of the uncini becoming expanded, curved, blunt, and bearing a subapical pocket or notch on the concave side. These differences with the thoracic uncini between
<taxonomicName id="C08A4D480652094AFC1EFE52FB8CFE10" box="[952,1104,440,464]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90652094AFC1EFE52FB8CFE10" box="[952,1104,440,464]" italics="true" pageId="71" pageNumber="72">L. cochleatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C08A4D480652094AFB2AFE52FAD9FE0F" box="[1164,1285,440,464]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90652094AFB2AFE52FAD9FE0F" box="[1164,1285,440,464]" italics="true" pageId="71" pageNumber="72">L. cirratus</emphasis>
</taxonomicName>
(as
<taxonomicName id="C08A4D480652094AFA9EFE52FA4EFE10" box="[1336,1426,440,464]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="ohlini">
<emphasis id="35FEEAD90652094AFA9EFE52FA4EFE10" box="[1336,1426,440,464]" italics="true" pageId="71" pageNumber="72">L ohlini</emphasis>
</taxonomicName>
) were clearly described and illustrated by
<bibRefCitation id="631B4B3A0652094AFDD7FE37FCD3FE34" author="Ehlers" box="[625,783,476,501]" pageId="71" pageNumber="72" refString="Ehlers, E. (1901) Die Polychaeten des magellanischen und chilenischen Strandes. Ein faunistischer Versuch. Festschrift zur Feier des Hundertfunfzigjahrigen Bestehens der koniglichen Gesellschaft der Wissenschaften zu Gottingen. pp. 1 - 232, 25 plates." type="book chapter" year="1901">Ehlers (1901)</bibRefCitation>
;
<taxonomicName id="C08A4D480652094AFCBBFE36FC4AFE34" box="[797,918,477,500]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90652094AFCBBFE36FC4AFE34" box="[797,918,477,500]" italics="true" pageId="71" pageNumber="72">L. cirratus</emphasis>
</taxonomicName>
is most closely related
<emphasis id="35FEEAD90652094AFB0EFE36FAB9FE34" box="[1192,1381,477,500]" italics="true" pageId="71" pageNumber="72">
to
<taxonomicName id="C08A4D480652094AFB61FE36FAB9FE34" box="[1223,1381,477,500]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="marginatus">L. marginatus</taxonomicName>
</emphasis>
(see above). Additional features of
<taxonomicName id="C08A4D480652094AFE5AFDEAFD4AFDD8" box="[508,662,512,536]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90652094AFE5AFDEAFD4AFDD8" box="[508,662,512,536]" italics="true" pageId="71" pageNumber="72">L. cochleatus</emphasis>
</taxonomicName>
not seen in
<taxonomicName id="C08A4D480652094AFC8AFDEAFC7AFDD7" box="[812,934,512,536]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cirratus">
<emphasis id="35FEEAD90652094AFC8AFDEAFC7AFDD7" box="[812,934,512,536]" italics="true" pageId="71" pageNumber="72">L. cirratus</emphasis>
</taxonomicName>
are with the presence of abdominal spines (aciculae) in the neuropodia and uniquely, in the notopodia (
<figureCitation id="9FB12A4E0652094AFC9EFDCFFC56FDFC" box="[824,906,548,573]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="71" pageNumber="72">Fig. 31</figureCitation>
B). The neuropodial spines number 12 and are weakly curved and blunt-tipped apically (
<figureCitation id="9FB12A4E0652094AFDCEFDACFD66FDA0" box="[616,698,583,608]" captionStart="FIGURE 32" captionStartId="72.[151,249,1512,1534]" captionTargetBox="[159,1431,347,1487]" captionTargetId="figure@72.[151,1436,339,1491]" captionTargetPageId="72" captionText="FIGURE 32. Leodamas cochleatus (Ehlers, 1900). Phase Contrast images from Argentinean specimens (AHF Poly 5043): A B, anterior thoracic uncini from setiger 2; C, detail of anterior thoracic uncinus showing longitudinal notch; D E, posterior thoracic uncini from setiger 8 showing expanded apex and notch; F, abdominal notopodial spine; G, abdominal neuropodium with spine and capillaries; H, abdominal notosetae including capillaries, spines (broken) and furcate seta." httpUri="https://zenodo.org/record/245858/files/figure.png" pageId="71" pageNumber="72">Fig. 32</figureCitation>
FG). The notopodial spines number 12 and are straight and also bluntly tipped (
<figureCitation id="9FB12A4E0652094AFEE5FD87FE49FD44" box="[323,405,620,645]" captionStart="FIGURE 31" captionStartId="71.[151,249,1820,1842]" captionTargetBox="[168,1419,709,1773]" captionTargetId="figure@71.[151,1436,697,1798]" captionTargetPageId="71" captionText="FIGURE 31. Leodamas cochleatus (Ehlers, 1900). Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; C E, anterior thoracic neuropodial uncini from setiger 2; F H, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta." httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="71" pageNumber="72">Fig. 31</figureCitation>
B).
</paragraph>
<caption id="53F566430652094AFF31F8F7FD5BF8AF" httpUri="https://zenodo.org/record/245857/files/figure.png" pageId="71" pageNumber="72" targetBox="[168,1419,709,1773]" targetPageId="71">
<paragraph id="073536CB0652094AFF31F8F7FD5BF8AF" blockId="71.[151,1436,1820,1903]" pageId="71" pageNumber="72">
<emphasis id="35FEEAD90652094AFF31F8F7FEC3F8F2" bold="true" box="[151,287,1820,1842]" pageId="71" pageNumber="72">FIGURE 31.</emphasis>
<taxonomicName id="C08A4D480652094AFE80F8F7FD40F8F2" authority="(Ehlers, 1900)" authorityName="Ehlers" authorityYear="1900" baseAuthorityName="Ehlers" baseAuthorityYear="1900" box="[294,668,1820,1842]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD90652094AFE80F8F7FDDDF8F2" box="[294,513,1820,1842]" italics="true" pageId="71" pageNumber="72">Leodamas cochleatus</emphasis>
(Ehlers, 1900)
</taxonomicName>
. Argentinean specimens (AHF Poly 5043). A, Anterior end dorsal view; B, abdominal parapodium, anterior view; CE, anterior thoracic neuropodial uncini from setiger 2; FH, posterior thoracic neuropodial uncini from setiger 8; I, furcate seta.
</paragraph>
</caption>
<paragraph id="073536CB06520945FF61F877FBE5FF38" blockId="71.[151,1436,1948,2009]" lastBlockId="72.[151,1436,151,285]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<taxonomicName id="C08A4D480652094AFF61F877FE7CF874" box="[199,416,1948,1972]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="71" pageNumber="72" phylum="Annelida" rank="species" species="hamatus">
<emphasis id="35FEEAD90652094AFF61F877FE7CF874" box="[199,416,1948,1972]" italics="true" pageId="71" pageNumber="72">Leodamas hamatus</emphasis>
</taxonomicName>
recently described from off
<collectingCountry id="7F9D765B0652094AFD44F877FC80F875" box="[738,860,1948,1973]" name="Costa Rica" pageId="71" pageNumber="72">
<collectingRegion id="C54EF8290652094AFD44F877FCFEF874" box="[738,802,1948,1972]" country="New Zealand" name="West Coast" pageId="71" pageNumber="72">Costa</collectingRegion>
Rica
</collectingCountry>
by
<bibRefCitation id="631B4B3A0652094AFC2EF876FBA1F875" author="Dean" box="[904,1149,1948,1973]" pageId="71" pageNumber="72" refString="Dean, H. K. &amp; Blake, J. A. (2015) The Orbiniidae (Annelida: Polychaeta) of Pacific Costa Rica. Zootaxa, 3956, 183 - 198. http: // dx. doi. org / 10.11646 / zootaxa. 3956.2.2" type="journal article" year="2015">Dean &amp; Blake (2015)</bibRefCitation>
, is another species where the neuropodial uncini change morphologically from anterior to posterior thoracic setigers. However, in contrast to
<taxonomicName id="C08A4D48065D0945FF31FF72FEF2FF70" box="[151,302,152,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="72" pageNumber="73" phylum="Annelida" rank="species" species="cochleatus">
<emphasis id="35FEEAD9065D0945FF31FF72FEF2FF70" box="[151,302,152,176]" italics="true" pageId="72" pageNumber="73">L. cochleatus</emphasis>
</taxonomicName>
, the posterior neuropodial spines of
<taxonomicName id="C08A4D48065D0945FD77FF72FC93FF70" box="[721,847,152,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="72" pageNumber="73" phylum="Annelida" rank="species" species="hamatus">
<emphasis id="35FEEAD9065D0945FD77FF72FC93FF70" box="[721,847,152,176]" italics="true" pageId="72" pageNumber="73">L. hamatus</emphasis>
</taxonomicName>
develop a prominent, thickened hood-like structure on the convex side of the curved spines. In addition, the abdominal neuropodial uncini of
<taxonomicName id="C08A4D48065D0945FB0FFF56FAF4FF14" box="[1193,1320,188,212]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="72" pageNumber="73" phylum="Annelida" rank="species" species="hamatus">
<emphasis id="35FEEAD9065D0945FB0FFF56FAF4FF14" box="[1193,1320,188,212]" italics="true" pageId="72" pageNumber="73">L. hamatus</emphasis>
</taxonomicName>
are large, heavy and curve to a pointed tip instead of being of being narrow and blunt-tipped.
</paragraph>
</subSubSection>
<subSubSection id="4F906540065D0945FF61FEEFFBEDFEDC" box="[199,1073,260,285]" pageId="72" pageNumber="73" type="distribution">
<paragraph id="073536CB065D0945FF61FEEFFBEDFEDC" blockId="72.[151,1436,151,285]" box="[199,1073,260,285]" pageId="72" pageNumber="73">
<emphasis id="35FEEAD9065D0945FF61FEEFFEBCFEDD" bold="true" box="[199,352,260,285]" pageId="72" pageNumber="73">Distribution.</emphasis>
Offshore
<collectingCountry id="7F9D765B065D0945FE77FEEFFD9FFEDD" box="[465,579,260,285]" name="Argentina" pageId="72" pageNumber="73">Argentina</collectingCountry>
, to
<quantity id="C0729B2E065D0945FDCDFEEFFD69FEDC" box="[619,693,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.54" pageId="72" pageNumber="73" unit="m" value="454.0">454 m</quantity>
;
<collectingCountry id="7F9D765B065D0945FD67FEEFFCDDFEDD" box="[705,769,260,285]" name="Chile" pageId="72" pageNumber="73">Chile</collectingCountry>
, Straits of Magellan;
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<quantity id="C0729B2E065D0945FC56FEEFFBF0FEDC" box="[1008,1068,260,285]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="72" pageNumber="73" unit="m" value="46.0">46 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>