474 lines
75 KiB
XML
474 lines
75 KiB
XML
<document id="F8B22D6EA83EFEABE7AEEB706D0CFFA4" ID-CLB-Dataset="172790" ID-DOI="10.11646/zootaxa.5198.1.1" ID-GBIF-Dataset="17babbe7-1645-47c8-ac5f-2299f1ad436c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251602" ID-ZooBank="019FE488-4263-4BC2-8606-446E599E226A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666769542164" checkinUser="plazi" docAuthor="Hou, Yanmeng, Gao, Zhizhong & Zhang, Feng" docDate="2022" docId="8A6487A5FFCBA921FF11FF72FD05FC96" docLanguage="en" docName="zootaxa.5198.1.1.pdf" docOrigin="Zootaxa 5198 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5198.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Lagynochthonius crassus Hou & Gao & Zhang 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="765DFFDDFFCEA92BFF86FFEAFFEBFFCB" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="6" updateTime="1701997268366" updateUser="plazi">
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<mods:title id="B7B7E43140A83817221D9D595EB6FC20">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="31C82F7B9ADC44F4C300149EA2086137">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="529027CBD8C6BBA7C364643D83CDFCC4">Gao, Zhizhong</mods:namePart>
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<mods:namePart id="ED9D4A92984044A22A7E6DD2D5254763">Zhang, Feng</mods:namePart>
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<mods:title id="B203B88A4E7372340B5B0706157A36FD">Zootaxa</mods:title>
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<mods:part id="5D93B95980659B235761855C4BB6068E">
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<mods:date id="C822EB0C5194A88E9B029E7F78B259DB">2022</mods:date>
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<mods:number id="5B014CE854B06202EA9624FE866E805B">2022-10-25</mods:number>
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<treatment id="8A6487A5FFCBA921FF11FF72FD05FC96" ID-DOI="http://doi.org/10.5281/zenodo.7255273" ID-GBIF-Taxon="203411090" ID-Zenodo-Dep="7255273" LSID="urn:lsid:plazi:treatment:8A6487A5FFCBA921FF11FF72FD05FC96" httpUri="http://treatment.plazi.org/id/8A6487A5FFCBA921FF11FF72FD05FC96" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6">
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<subSubSection id="4AD76538FFCBA92EFF11FF72FD03FF7B" box="[151,744,152,179]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="027236B3FFCBA92EFF11FF72FD03FF7B" blockId="5.[151,744,152,215]" box="[151,744,152,179]" pageId="5" pageNumber="6">
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<heading id="593A81DFFFCBA92EFF11FF72FD03FF7B" bold="true" box="[151,744,152,179]" fontSize="11" level="2" pageId="5" pageNumber="6" reason="4">
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<emphasis id="30B9EAA1FFCBA92EFF11FF72FD03FF7B" bold="true" box="[151,744,152,179]" pageId="5" pageNumber="6">
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<taxonomicName id="C5CD4D30FFCBA92EFF11FF72FE27FF78" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[151,460,152,179]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
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<emphasis id="30B9EAA1FFCBA92EFF11FF72FE27FF78" bold="true" box="[151,460,152,179]" italics="true" pageId="5" pageNumber="6">Lagynochthonius crassus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2B8A57DAFFCBA92EFE55FF72FDDAFF79" box="[467,561,152,178]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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(DḆIJ伪oi)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AD76538FFCBA92EFF11FF56FF12FF1C" box="[151,249,188,215]" pageId="5" pageNumber="6" type="description">
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<paragraph id="027236B3FFCBA92EFF11FF56FF12FF1C" blockId="5.[151,744,152,215]" box="[151,249,188,215]" pageId="5" pageNumber="6">
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<figureCitation id="9AF62A36FFCBA92EFF11FF56FF12FF1C" box="[151,249,188,215]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1944,1968]" captionStartId-1="7.[151,250,858,882]" captionStartId-2="8.[151,250,1907,1931]" captionStartId-3="9.[151,250,1829,1853]" captionTargetBox-0="[151,1436,273,1921]" captionTargetBox-1="[181,1434,189,830]" captionTargetBox-2="[151,1436,202,1883]" captionTargetBox-3="[186,1403,183,1797]" captionTargetId-0="figure-80@6.[151,1436,273,1921]" captionTargetId-1="figure-19@7.[151,1436,181,834]" captionTargetId-2="figure-20@8.[151,1436,181,1883]" captionTargetId-3="figure-19@9.[185,1403,181,1805]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 2. Unnamed Cave 1, type locality of Lagynochthonius crassus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where L. crassus sp. nov. specimens were collected; D–E. Live individuals of L. crassus sp. nov. (D: male, E: female) in their natural environment." captionText-1="FIGURE 3. Lagynochthonius crassus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." captionText-3="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi-0="http://doi.org/10.5281/zenodo.7251608" figureDoi-1="http://doi.org/10.5281/zenodo.7251610" figureDoi-2="http://doi.org/10.5281/zenodo.7251612" figureDoi-3="http://doi.org/10.5281/zenodo.7251616" httpUri-0="https://zenodo.org/record/7251608/files/figure.png" httpUri-1="https://zenodo.org/record/7251610/files/figure.png" httpUri-2="https://zenodo.org/record/7251612/files/figure.png" httpUri-3="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Figs 2–5</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="4AD76538FFCBA92EFF11FEEEFE95FE65" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="027236B3FFCBA92EFF11FEEEFE95FE65" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
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<emphasis id="30B9EAA1FFCBA92EFF11FEEEFEA0FED5" bold="true" box="[151,331,260,286]" pageId="5" pageNumber="6">Type material.</emphasis>
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<materialsCitation id="B2A53CEEFFCBA92EFEDEFEEEFBB0FEAD" ID-GBIF-Occurrence="3946861304" collectingDate="2021-04-21" collectionCode="MHBU-HBUARA" collectorName="Zegang Feng" country="China" county="Maguan County" elevation="1508" latitude="22.892042" location="Laomenzhai Village" longLatPrecision="1" longitude="104.09709" municipality="Miechang Township" pageId="5" pageNumber="6" specimenCode="MHBU-HBUARA#2021-28801" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="DD768811FFCBA92EFEDEFEEEFE2BFED5" box="[344,448,260,286]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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<specimenCount id="14CBFD3AFFCBA92EFE48FEEFFE08FED4" box="[462,483,261,287]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
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(Ps.-
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<specimenCode id="526B9EC8FFCBA92EFD9BFEEEFC78FED5" box="[541,915,260,286]" collectionCode="MHBU-HBUARA" pageId="5" pageNumber="6">MHBU-HBUARA#2021-28801</specimenCode>
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):
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<collectingCountry id="7ADA7623FFCBA92EFC2DFEEEFC1AFED5" box="[939,1009,260,286]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
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,
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||
<collectingRegion id="C009F851FFCBA92EFB87FEEFFB21FED5" box="[1025,1226,260,287]" country="China" name="Yunnan" pageId="5" pageNumber="6">Yunnan Province</collectingRegion>
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,
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<collectingCounty id="EB134E3FFFCBA92EFB5AFEEFFA72FED5" box="[1244,1433,260,287]" pageId="5" pageNumber="6">Maguan County</collectingCounty>
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,
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<collectingMunicipality id="E216ACC9FFCBA92EFF11FEC2FE95FE89" box="[151,382,296,322]" pageId="5" pageNumber="6">Miechang Township</collectingMunicipality>
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,
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<location id="07126068FFCBA92EFE0EFEC2FD87FE89" LSID="urn:lsid:plazi:treatment:8A6487A5FFCBA921FF11FF72FD05FC96:07126068FFCBA92EFE0EFEC2FD87FE89" box="[392,620,296,322]" country="China" county="Maguan County" latitude="22.892042" longLatPrecision="1" longitude="104.09709" municipality="Miechang Township" name="Laomenzhai Village" pageId="5" pageNumber="6" stateProvince="Yunnan">Laomenzhai Village</location>
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, cave without name (
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<location id="07126068FFCBA92EFCD0FEC2FBEEFE89" LSID="urn:lsid:plazi:treatment:8A6487A5FFCBA921FF11FF72FD05FC96:07126068FFCBA92EFCD0FEC2FBEEFE89" box="[854,1029,296,322]" country="China" county="Maguan County" latitude="22.892042" longLatPrecision="1" longitude="104.09709" municipality="Miechang Township" name="Unnamed Cave" pageId="5" pageNumber="6" stateProvince="Yunnan">Unnamed Cave</location>
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1), under stones and detritus in deep zone [
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<geoCoordinate id="67F95074FFCBA92EFF5FFEA6FE6FFEAC" box="[217,388,332,359]" degrees="22" direction="north" minutes="53" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="31.35" value="22.892042">22°53'31.35"N</geoCoordinate>
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,
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<geoCoordinate id="67F95074FFCBA92EFE09FEA6FDACFEAC" box="[399,583,332,359]" degrees="104" direction="east" minutes="05" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="49.52" value="104.09709">104°05'49.52"E</geoCoordinate>
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],
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<elevation id="89E0D180FFCBA92EFDDEFEA6FD0CFEAD" box="[600,743,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.508" pageId="5" pageNumber="6" unit="m" value="1508.0">
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<quantity id="C5359B56FFCBA92EFDDEFEA6FD47FEAC" box="[600,684,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.508" pageId="5" pageNumber="6" unit="m" value="1508.0">1508 m</quantity>
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a.s.l.
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</elevation>
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,
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<date id="76731073FFCBA92EFD75FEA6FC64FEAD" box="[755,911,332,358]" pageId="5" pageNumber="6" value="2021-04-21">
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||
<collectingDate id="6637E99BFFCBA92EFD75FEA6FC64FEAD" box="[755,911,332,358]" pageId="5" pageNumber="6" value="2021-04-21">21 April 2021</collectingDate>
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||
</date>
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||
,
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<collectorName id="AF385365FFCBA92EFC1FFEA7FBC7FEAC" box="[921,1068,333,359]" pageId="5" pageNumber="6">Zegang Feng</collectorName>
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leg.
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</materialsCitation>
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<typeStatus id="DD768811FFCBA92EFBE7FEA7FB39FEAC" box="[1121,1234,333,359]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="14CBFD3AFFCBA92EFB5AFEA6FAEEFEAC" box="[1244,1285,332,359]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
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(Ps.-MHBU-HBUARA#2021-28802),
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<materialsCitation id="B2A53CEEFFCBA92EFE3DFE9AFE91FE65" ID-GBIF-Occurrence="3946861324" collectingDate="2021-04-21" collectionCode="MSWU-HBUARA" collectorName="Zegang Feng" country="China" county="Maguan County" elevation="1508" latitude="22.892042" location="Laomenzhai Village" longLatPrecision="1" longitude="104.09709" municipality="Miechang Township" pageId="5" pageNumber="6" specimenCode="MSWU-HBUARA#2021-28803" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="14CBFD3AFFCBA92EFE3DFE9AFE0CFE40" box="[443,487,368,395]" pageId="5" pageNumber="6" type="female">2 ♀</specimenCount>
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(Ps.-
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<specimenCode id="526B9EC8FFCBA92EFD9AFE9AFC64FE41" box="[540,911,368,394]" collectionCode="MSWU-HBUARA" pageId="5" pageNumber="6">MSWU-HBUARA#2021-28803</specimenCode>
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& HBUARA#2021-28804), all with the same data as the holotype
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4AD76538FFCBA92EFF41FE52FDD9FE3D" pageId="5" pageNumber="6" type="etymology">
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<paragraph id="027236B3FFCBA92EFF41FE52FDD9FE3D" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
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<emphasis id="30B9EAA1FFCBA92EFF41FE52FEA5FE19" bold="true" box="[199,334,440,466]" pageId="5" pageNumber="6">Etymology.</emphasis>
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The specific name is derived from the Latin word “
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<taxonomicName id="C5CD4D30FFCBA92EFC23FE53FC12FE19" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[933,1017,441,466]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassus">
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<emphasis id="30B9EAA1FFCBA92EFC23FE53FC12FE19" box="[933,1017,441,466]" italics="true" pageId="5" pageNumber="6">crassus</emphasis>
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</taxonomicName>
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”, meaning thick, which refers to the character of a thickened chelal hand.
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</paragraph>
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</subSubSection>
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<subSubSection id="4AD76538FFCBA92EFF41FDEAFACFFD04" pageId="5" pageNumber="6" type="diagnosis">
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||
<paragraph id="027236B3FFCBA92EFF41FDEAFACFFD04" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
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<emphasis id="30B9EAA1FFCBA92EFF41FDEAFE63FDD5" bold="true" box="[199,392,512,542]" pageId="5" pageNumber="6">
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Diagnosis (
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<specimenCount id="14CBFD3AFFCBA92EFEC9FDE8FE8FFDD5" box="[335,356,514,542]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
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<specimenCount id="14CBFD3AFFCBA92EFEE5FDE8FE93FDD5" box="[355,376,514,542]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
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).
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</emphasis>
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Moderately sized troglomorphic species with elongated appendages; carapace with a pair of anterior eyespots only; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with 2 setae; tergites I–Ⅴ each with 4 setae. Pedipalps slender, femur 8.21–8.50 (
|
||
<specimenCount id="14CBFD3AFFCBA92EFAF1FDA2FA65FDA9" box="[1399,1422,584,610]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
|
||
), 7.50–7.87 (
|
||
<specimenCount id="14CBFD3AFFCBA92EFE9DFD86FEC6FD4D" box="[283,301,620,646]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
) times longer than broad; chela 7.00–7.14 (
|
||
<specimenCount id="14CBFD3AFFCBA92EFCA3FD86FCD7FD4D" box="[805,828,620,646]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
|
||
), 6.40–6.46 (
|
||
<specimenCount id="14CBFD3AFFCBA92EFC50FD86FC03FD4D" box="[982,1000,620,646]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
) times longer than broad; chelal hand strong and thick; both chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (
|
||
<emphasis id="30B9EAA1FFCBA92EFF27FD5EFF5CFD05" box="[161,183,692,718]" italics="true" pageId="5" pageNumber="6">td</emphasis>
|
||
) on dorso-antiaxial face; chemosensory setae (
|
||
<emphasis id="30B9EAA1FFCBA92EFD43FD5FFD37FD05" box="[709,732,693,718]" italics="true" pageId="5" pageNumber="6">sc</emphasis>
|
||
) present on dorsum of chelal hand; sensilla present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFCBA921FF41FD32FE06FE8A" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="027236B3FFCBA92EFF41FD32FC24FD38" blockId="5.[151,1437,260,2015]" box="[199,975,728,755]" pageId="5" pageNumber="6">
|
||
<emphasis id="30B9EAA1FFCBA92EFF41FD32FEB2FD39" bold="true" box="[199,345,728,754]" pageId="5" pageNumber="6">Description.</emphasis>
|
||
Males (
|
||
<typeStatus id="DD768811FFCBA92EFE35FD32FDFCFD39" box="[435,535,728,754]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="DD768811FFCBA92EFDCBFD32FD44FD39" box="[589,687,728,754]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<figureCitation id="9AF62A36FFCBA92EFD43FD32FCC8FD39" box="[709,803,728,754]" captionStart="FIGURE 2" captionStartId="6.[151,250,1944,1968]" captionTargetBox="[151,1436,273,1921]" captionTargetId="figure-80@6.[151,1436,273,1921]" captionTargetPageId="6" captionText="FIGURE 2. Unnamed Cave 1, type locality of Lagynochthonius crassus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where L. crassus sp. nov. specimens were collected; D–E. Live individuals of L. crassus sp. nov. (D: male, E: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251608" httpUri="https://zenodo.org/record/7251608/files/figure.png" pageId="5" pageNumber="6">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFCA8FD32FCB9FD39" box="[814,850,728,754]" captionStart="FIGURE 3" captionStartId="7.[151,250,858,882]" captionTargetBox="[181,1434,189,830]" captionTargetId="figure-19@7.[151,1436,181,834]" captionTargetPageId="7" captionText="FIGURE 3. Lagynochthonius crassus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251610" httpUri="https://zenodo.org/record/7251610/files/figure.png" pageId="5" pageNumber="6">3A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFCD9FD32FC4DFD39" box="[863,934,728,754]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="5" pageNumber="6">4A–D</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFC37FD32FC2AFD38" box="[945,961,728,755]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFCBA92EFF41FD16FF26FCF1" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
|
||
Colour (
|
||
<figureCitation id="9AF62A36FFCBA92EFEA5FD16FE6BFCDD" box="[291,384,764,790]" captionStart="FIGURE 2" captionStartId="6.[151,250,1944,1968]" captionTargetBox="[151,1436,273,1921]" captionTargetId="figure-80@6.[151,1436,273,1921]" captionTargetPageId="6" captionText="FIGURE 2. Unnamed Cave 1, type locality of Lagynochthonius crassus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where L. crassus sp. nov. specimens were collected; D–E. Live individuals of L. crassus sp. nov. (D: male, E: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251608" httpUri="https://zenodo.org/record/7251608/files/figure.png" pageId="5" pageNumber="6">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFE0CFD16FE44FCDD" box="[394,431,764,790]" captionStart="FIGURE 3" captionStartId="7.[151,250,858,882]" captionTargetBox="[181,1434,189,830]" captionTargetId="figure-19@7.[151,1436,181,834]" captionTargetPageId="7" captionText="FIGURE 3. Lagynochthonius crassus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251610" httpUri="https://zenodo.org/record/7251610/files/figure.png" pageId="5" pageNumber="6">3A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFE3CFD16FDEFFCDD" box="[442,516,764,790]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="5" pageNumber="6">4A–D</figureCitation>
|
||
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFCBA92EFF41FCAEFC5EFB68" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
|
||
Cephalothorax (
|
||
<figureCitation id="9AF62A36FFCBA92EFEFCFCAEFE3FFC95" box="[378,468,836,862]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="5" pageNumber="6">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFE59FCAEFDEEFC94" box="[479,517,836,863]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">5A</figureCitation>
|
||
): carapace 0.90–0.93 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior eyes reduced into eyespots, posterior eyes reduced completely; anterior margin slightly serrate; epistome obtuse and small, round, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, preocular setae absent in
|
||
<specimenCount id="14CBFD3AFFCBA92EFE34FC5BFDF1FC01" box="[434,538,944,971]" pageId="5" pageNumber="6" type="male">one male</specimenCount>
|
||
, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 4–5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
|
||
<figureCitation id="9AF62A36FFCBA92EFEA1FB62FE94FB68" box="[295,383,1160,1187]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5C</figureCitation>
|
||
); intercoxal tubercle absent; without sub-oral seta.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFCBA92EFF41FB46FD04FA08" blockId="5.[151,1437,260,2015]" pageId="5" pageNumber="6">
|
||
Chelicera (
|
||
<figureCitation id="9AF62A36FFCBA92EFEC2FB46FE49FB0D" box="[324,418,1196,1222]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="5" pageNumber="6">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFE36FB46FE3FFB0C" box="[432,468,1196,1223]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">5B</figureCitation>
|
||
): large, about as long as carapace, 2.33–2.35 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation dorsal side. Both fingers well provided with teeth, fixed finger with 13–15 teeth, distal one largest; movable finger with 11 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (
|
||
<figureCitation id="9AF62A36FFCBA92EFAB5FAD6FA64FA9C" box="[1331,1423,1340,1367]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5B</figureCitation>
|
||
). Serrula exterior with 21–22 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
|
||
<figureCitation id="9AF62A36FFCBA92EFD0EFA42FD0AFA08" box="[648,737,1448,1475]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFCBA92DFF41FA26FB42FF32" blockId="5.[151,1437,260,2015]" lastBlockId="6.[151,1436,151,249]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Pedipalp (
|
||
<figureCitation id="9AF62A36FFCBA92EFEBFFA26FE73FA2D" box="[313,408,1484,1510]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="5" pageNumber="6">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFCBA92EFE23FA26FE07FA2C" box="[421,492,1484,1511]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">5E–G</figureCitation>
|
||
): long and slender, trochanter 1.94–2.20, femur 8.21–8.50, patella 2.69–2.71, chela 7.00–7.14, hand 2.73–2.82 times longer than broad; femur 2.59–2.67 times longer than patella; movable chelal finger 1.53–1.57 times longer than hand and 0.61 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
|
||
<figureCitation id="9AF62A36FFCBA92EFDAEF9D2FD94F998" box="[552,639,1592,1619]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5E</figureCitation>
|
||
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
|
||
<emphasis id="30B9EAA1FFCBA92EFB0EF96AFB75F951" box="[1160,1182,1664,1690]" italics="true" pageId="5" pageNumber="6">ib</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFCBA92EFB5FF96AFB11F951" box="[1241,1274,1664,1690]" italics="true" pageId="5" pageNumber="6">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis id="30B9EAA1FFCBA92EFD34F94EFD27F975" box="[690,716,1700,1726]" italics="true" pageId="5" pageNumber="6">eb</emphasis>
|
||
,
|
||
<emphasis id="30B9EAA1FFCBA92EFD5FF94EFD15F975" box="[729,766,1700,1726]" italics="true" pageId="5" pageNumber="6">esb</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFCBA92EFCB4F94FFCA7F975" box="[818,844,1701,1726]" italics="true" pageId="5" pageNumber="6">ist</emphasis>
|
||
forming a nearly straight oblique row at base of fixed chelal finger;
|
||
<emphasis id="30B9EAA1FFCBA92EFEB2F923FEAFF929" box="[308,324,1737,1762]" italics="true" pageId="5" pageNumber="6">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="30B9EAA1FFCBA92EFD8FF923FDC3F929" box="[521,552,1737,1762]" italics="true" pageId="5" pageNumber="6">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis id="30B9EAA1FFCBA92EFC98F923FCD9F929" box="[798,818,1737,1762]" italics="true" pageId="5" pageNumber="6">et</emphasis>
|
||
slightly near to tip of fixed chelal finger, very close to chelal teeth;
|
||
<emphasis id="30B9EAA1FFCBA92EFEA1F906FEAAF8CD" box="[295,321,1772,1798]" italics="true" pageId="5" pageNumber="6">dx</emphasis>
|
||
situated distal to
|
||
<emphasis id="30B9EAA1FFCBA92EFD8EF907FDF7F8CD" box="[520,540,1773,1798]" italics="true" pageId="5" pageNumber="6">et</emphasis>
|
||
;
|
||
<emphasis id="30B9EAA1FFCBA92EFDADF906FDAFF8CD" box="[555,580,1772,1798]" italics="true" pageId="5" pageNumber="6">sb</emphasis>
|
||
closer to
|
||
<emphasis id="30B9EAA1FFCBA92EFD35F907FD2DF8CD" box="[691,710,1773,1798]" italics="true" pageId="5" pageNumber="6">st</emphasis>
|
||
than to
|
||
<emphasis id="30B9EAA1FFCBA92EFCA4F906FCDBF8CD" box="[802,816,1772,1798]" italics="true" pageId="5" pageNumber="6">b</emphasis>
|
||
;
|
||
<emphasis id="30B9EAA1FFCBA92EFCB9F906FCA6F8CD" box="[831,845,1772,1798]" italics="true" pageId="5" pageNumber="6">b</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFCBA92EFC02F907FC67F8CD" box="[900,908,1773,1798]" italics="true" pageId="5" pageNumber="6">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis id="30B9EAA1FFCBA92EFBF1F906FB6EF8CD" box="[1143,1157,1772,1798]" italics="true" pageId="5" pageNumber="6">b</emphasis>
|
||
situated at same level as
|
||
<emphasis id="30B9EAA1FFCBA92EFF11F8FBFF5DF8E1" box="[151,182,1809,1834]" italics="true" pageId="5" pageNumber="6">est</emphasis>
|
||
;
|
||
<emphasis id="30B9EAA1FFCBA92EFF43F8FBFF26F8E1" box="[197,205,1809,1834]" italics="true" pageId="5" pageNumber="6">t</emphasis>
|
||
situated distal to
|
||
<emphasis id="30B9EAA1FFCBA92EFE16F8FBFE4BF8E1" box="[400,416,1809,1834]" italics="true" pageId="5" pageNumber="6">it</emphasis>
|
||
(
|
||
<figureCitation id="9AF62A36FFCBA92EFE29F8FAFDEFF8E0" box="[431,516,1808,1835]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5F</figureCitation>
|
||
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
|
||
<emphasis id="30B9EAA1FFCBA92EFF11F8DFFF5AF885" box="[151,177,1845,1870]" italics="true" pageId="5" pageNumber="6">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30–31 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on dorso-antiaxial face (
|
||
<emphasis id="30B9EAA1FFCBA92EFF27F84AFF5CF871" box="[161,183,1952,1978]" italics="true" pageId="5" pageNumber="6">td</emphasis>
|
||
, slightly distal to
|
||
<emphasis id="30B9EAA1FFCBA92EFE04F84AFE77F871" box="[386,412,1952,1978]" italics="true" pageId="5" pageNumber="6">dx</emphasis>
|
||
); movable chelal finger with 15 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 13–15 small, rounded and contiguous basal teeth,
|
||
<quantity id="C5359B56FFCBA92EFB15F82EFB1EF815" box="[1171,1269,1988,2014]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="7.62" metricValueMin="7.112" pageId="5" pageNumber="6" unit="in" value="29.0" valueMax="30.0" valueMin="28.0">28–30 in</quantity>
|
||
total (
|
||
<figureCitation id="9AF62A36FFCBA92EFABDF82EFA7BF814" box="[1339,1424,1988,2015]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="5" pageNumber="6">Fig. 5F</figureCitation>
|
||
). Fixed chelal finger with sensilla
|
||
<emphasis id="30B9EAA1FFC8A92DFD8FFF7DFDF4FF7A" box="[521,543,151,177]" italics="true" pageId="6" pageNumber="7">af</emphasis>
|
||
<subScript id="9E4934F6FFC8A92DFD98FF43FDDCFF7C" attach="left" box="[542,567,169,183]" fontSize="6" pageId="6" pageNumber="7">1–2</subScript>
|
||
close together, near tip; movable chelal finger with 4 sensilla:
|
||
<emphasis id="30B9EAA1FFC8A92DFB7BFF7DFAF4FF7B" box="[1277,1311,151,176]" italics="true" pageId="6" pageNumber="7">am</emphasis>
|
||
<subScript id="9E4934F6FFC8A92DFAA6FF43FAD3FF7C" attach="left" box="[1312,1336,169,183]" fontSize="6" pageId="6" pageNumber="7">1–2</subScript>
|
||
near tip,
|
||
<emphasis id="30B9EAA1FFC8A92DFF11FF51FF4EFF1F" box="[151,165,187,212]" italics="true" pageId="6" pageNumber="7">p</emphasis>
|
||
<subScript id="9E4934F6FFC8A92DFF23FF27FF46FF10" attach="left" box="[165,173,205,219]" fontSize="6" pageId="6" pageNumber="7">1</subScript>
|
||
slightly distad of
|
||
<emphasis id="30B9EAA1FFC8A92DFEF0FF51FE64FF1E" box="[374,399,187,213]" italics="true" pageId="6" pageNumber="7">sb</emphasis>
|
||
and very close to chelal teeth,
|
||
<emphasis id="30B9EAA1FFC8A92DFD61FF51FD1EFF1F" box="[743,757,187,212]" italics="true" pageId="6" pageNumber="7">p</emphasis>
|
||
<subScript id="9E4934F6FFC8A92DFD73FF27FD16FF10" attach="left" box="[757,765,205,219]" fontSize="6" pageId="6" pageNumber="7">2</subScript>
|
||
slightly distad of
|
||
<emphasis id="30B9EAA1FFC8A92DFC40FF51FC3FFF1F" box="[966,980,187,212]" italics="true" pageId="6" pageNumber="7">p</emphasis>
|
||
<subScript id="9E4934F6FFC8A92DFC52FF27FC37FF10" attach="left" box="[980,988,205,219]" fontSize="6" pageId="6" pageNumber="7">1</subScript>
|
||
(
|
||
<figureCitation id="9AF62A36FFC8A92DFC6FFF51FBD5FF1E" box="[1001,1086,186,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="6" pageNumber="7">Fig. 5F</figureCitation>
|
||
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
|
||
<figureCitation id="9AF62A36FFC8A92DFBC4FF35FB77FF32" box="[1090,1180,222,249]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="6" pageNumber="7">Fig. 5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="56B2663BFFC8A92DFF11F872FE4BF833" ID-DOI="http://doi.org/10.5281/zenodo.7251608" ID-Zenodo-Dep="7251608" httpUri="https://zenodo.org/record/7251608/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1944,1968]" targetBox="[151,1436,273,1921]" targetPageId="6">
|
||
<paragraph id="027236B3FFC8A92DFF11F872FE4BF833" blockId="6.[151,1437,1944,2040]" pageId="6" pageNumber="7">
|
||
<emphasis id="30B9EAA1FFC8A92DFF11F872FEF8F87B" bold="true" box="[151,275,1944,1968]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
|
||
Unnamed Cave 1, type locality of
|
||
<taxonomicName id="C5CD4D30FFC8A92DFDFEF872FC91F87B" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[632,890,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC8A92DFDFEF872FC91F87B" box="[632,890,1944,1968]" italics="true" pageId="6" pageNumber="7">Lagynochthonius crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC8A92DFC06F872FC3AF87B" bold="true" box="[896,977,1944,1968]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="2B8A57DAFFC8A92DFC06F872FC3AF87B" box="[896,977,1944,1968]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Entrance; B. Inside the cave entrance; C. Area where
|
||
<taxonomicName id="C5CD4D30FFC8A92DFE95F857FE91F81F" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[275,378,1981,2004]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC8A92DFE95F857FE91F81F" box="[275,378,1981,2004]" italics="true" pageId="6" pageNumber="7">L. crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC8A92DFE07F856FE3AF81F" bold="true" box="[385,465,1980,2004]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="2B8A57DAFFC8A92DFE07F856FE3AF81F" box="[385,465,1980,2004]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimens were collected; D–E. Live individuals of
|
||
<taxonomicName id="C5CD4D30FFC8A92DFC6AF857FBB8F81F" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[1004,1107,1981,2004]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC8A92DFC6AF857FBB8F81F" box="[1004,1107,1981,2004]" italics="true" pageId="6" pageNumber="7">L. crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC8A92DFBDCF856FB41F81F" bold="true" box="[1114,1194,1980,2004]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="2B8A57DAFFC8A92DFBDCF856FB41F81F" box="[1114,1194,1980,2004]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(D: male, E: female) in their natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="56B2663BFFC9A92CFF11FCB0FF33FC5D" ID-DOI="http://doi.org/10.5281/zenodo.7251610" ID-Zenodo-Dep="7251610" httpUri="https://zenodo.org/record/7251610/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,858,882]" targetBox="[181,1434,189,830]" targetPageId="7">
|
||
<paragraph id="027236B3FFC9A92CFF11FCB0FF33FC5D" blockId="7.[151,1436,858,918]" pageId="7" pageNumber="8">
|
||
<emphasis id="30B9EAA1FFC9A92CFF11FCB0FEFFFCB9" bold="true" box="[151,276,858,882]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFC9A92CFE9AFCB0FDF5FCB9" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[284,542,858,882]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC9A92CFE9AFCB0FDF5FCB9" box="[284,542,858,882]" italics="true" pageId="7" pageNumber="8">Lagynochthonius crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC9A92CFDA0FCB0FD93FCB9" bold="true" box="[550,632,858,882]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="2B8A57DAFFC9A92CFDA0FCB0FD93FCB9" box="[550,632,858,882]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="027236B3FFC9A92CFF41FC2EFEECFBA5" blockId="7.[151,1437,964,2035]" pageId="7" pageNumber="8">
|
||
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 5: 5: 5–6: 4: T2T: 0; tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 9–11: 10–12: 8–9: 8–10: 8–9: 9–10: 9–11: 7–8: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 14–16 marginal setae on each side,
|
||
<quantity id="C5359B56FFC9A92CFA85FBDAFA8EFB81" box="[1283,1381,1072,1098]" metricMagnitude="0" metricUnit="m" metricValue="1.0287" metricValueMax="1.0413999999999999" metricValueMin="1.016" pageId="7" pageNumber="8" unit="in" value="40.5" valueMax="41.0" valueMin="40.0">40–41 in</quantity>
|
||
total (
|
||
<figureCitation id="9AF62A36FFC9A92CFF19FBBEFF11FBA5" box="[159,250,1108,1134]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="7" pageNumber="8">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFC9A92CFF41FB93FBF1FAA1" blockId="7.[151,1437,964,2035]" pageId="7" pageNumber="8">
|
||
Legs (
|
||
<figureCitation id="9AF62A36FFC9A92CFE8AFB92FE97FB59" box="[268,380,1144,1171]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="7" pageNumber="8">Fig. 5H, I</figureCitation>
|
||
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur I, IV and patella I, IV. Femur of leg I 1.83–1.84 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.28–2.29 times longer than tibia. Femoropatella of leg IV 4.26–5.30 times longer than deep; tibia 6.60–7.44 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.00–4.25 times longer than deep (TS= 0.34–0.35), telotarsus 14.67–17.40 times longer than deep and 2.56–2.75 times longer than basitarsus (TS= 0.34–0.36). Setae of leg I (trochanter to tibia) 3: 11: 8–10: 10–11, setae of leg IV (trochanter to basitarsus) 2–3: 4: 4: 11: 10–11. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFC9A92CFF41FA9EFCD5F989" blockId="7.[151,1437,964,2035]" pageId="7" pageNumber="8">
|
||
<materialsCitation id="B2A53CEEFFC9A92CFF41FA9EFCD1F989" ID-GBIF-Occurrence="3946861316" collectionCode="T, IV" pageId="7" pageNumber="8" specimenCode="III 5, IV 5, T2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
Adult females (
|
||
<typeStatus id="DD768811FFC9A92CFEF0FA9EFE0EFA45" box="[374,485,1396,1422]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="9AF62A36FFC9A92CFE7AFA9EFDB3FA45" box="[508,600,1396,1422]" captionStart="FIGURE 2" captionStartId="6.[151,250,1944,1968]" captionTargetBox="[151,1436,273,1921]" captionTargetId="figure-80@6.[151,1436,273,1921]" captionTargetPageId="6" captionText="FIGURE 2. Unnamed Cave 1, type locality of Lagynochthonius crassus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where L. crassus sp. nov. specimens were collected; D–E. Live individuals of L. crassus sp. nov. (D: male, E: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251608" httpUri="https://zenodo.org/record/7251608/files/figure.png" pageId="7" pageNumber="8">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFC9A92CFDE3FA9EFD63FA45" box="[613,648,1396,1422]" captionStart="FIGURE 3" captionStartId="7.[151,250,858,882]" captionTargetBox="[181,1434,189,830]" captionTargetId="figure-19@7.[151,1436,181,834]" captionTargetPageId="7" captionText="FIGURE 3. Lagynochthonius crassus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251610" httpUri="https://zenodo.org/record/7251610/files/figure.png" pageId="7" pageNumber="8">3B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFC9A92CFD13FA9EFD53FA45" box="[661,696,1396,1422]" captionStart="FIGURE 4" captionStartId="8.[151,250,1907,1931]" captionTargetBox="[151,1436,202,1883]" captionTargetId="figure-20@8.[151,1436,181,1883]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius crassus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="7" pageNumber="8">4E</figureCitation>
|
||
). Mostly same as males, but a little larger; preocular setae absent in
|
||
<specimenCount id="14CBFD3AFFC9A92CFF30FA73FEDCFA79" box="[182,311,1432,1459]" pageId="7" pageNumber="8" type="female">one female</specimenCount>
|
||
; chaetotaxy of coxae: P 3, I 3, II 4,
|
||
<specimenCode id="526B9EC8FFC9A92CFD50FA73FCE1FA78" box="[726,778,1432,1459]" collectionCode="III" pageId="7" pageNumber="8">III 5</specimenCode>
|
||
,
|
||
<specimenCode id="526B9EC8FFC9A92CFC9EFA73FCA5FA78" box="[792,846,1432,1459]" collectionCode="IV" pageId="7" pageNumber="8">IV 5</specimenCode>
|
||
; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4–5: 5: 6: 5: 4:
|
||
<specimenCode id="526B9EC8FFC9A92CFF33FA56FF3EFA1D" box="[181,213,1468,1494]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="8">T2</specimenCode>
|
||
<collectionCode id="64DCAE76FFC9A92CFF50FA56FF03FA1D" box="[214,232,1468,1494]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="8">T</collectionCode>
|
||
: 0; sternal chaetotaxy
|
||
<collectionCode id="64DCAE76FFC9A92CFE61FA57FDEEFA1C" box="[487,517,1469,1495]" pageId="7" pageNumber="8">IV</collectionCode>
|
||
–XII: 8–11: 9–11: 9–10: 9–11: 10: 11: 7–8: 0: 2; anterior genital operculum with 10 setae, posterior margin with 8–10 marginal setae,
|
||
<quantity id="C5359B56FFC9A92CFD73FA0AFCB1FA31" box="[757,858,1504,1530]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="7" pageNumber="8" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
|
||
total; leg
|
||
<collectionCode id="64DCAE76FFC9A92CFC49FA0BFC07FA30" box="[975,1004,1505,1531]" pageId="7" pageNumber="8">IV</collectionCode>
|
||
with a long tactile seta on both tarsal segments: basitarsus 4.25 times longer than deep (TS= 0.29–0.35), telotarsus 15.00–16.60 times longer than deep and 2.44–2.65 times longer than basitarsus (TS= 0.31–0.36)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFC9A92CFF41F9A6FED6F84D" blockId="7.[151,1437,964,2035]" pageId="7" pageNumber="8">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.84–1.94. Pedipalps: trochanter 0.31–0.33/0.15–0.16 (1.94–2.20), femur 1.15–1.19/0.14 (8.21–8.50), patella 0.43–0.46/0.16–0.17 (2.69–2.71), chela 1.54–1.57/0.22 (7.00–7.14), hand 0.60–0.62/0.22 (2.73–2.82), movable chelal finger length 0.94–0.95. Chelicera 0.61–0.63/0.26–0.27 (2.33–2.35), movable finger length 0.34–0.35. Carapace 0.54/0.58–0.60 (0.90–0.93). Leg I: trochanter 0.18–0.21/0.11–0.13 (1.62–1.64), femur 0.66–0.68/0.07– 0.08 (8.50–9.43), patella 0.36–0.37/0.06–0.07 (5.29–6.00), tibia 0.31–0.32/0.07 (4.43–4.57), tarsus 0.71–0.73/0.06 (11.83–12.17). Leg IV: trochanter 0.22–0.26/0.16 (1.38–1.63), femoropatella 0.98–1.06/0.20–0.23 (4.26–5.30), tibia 0.66–0.67/0.09–0.10 (6.60–7.44), basitarsus 0.32–0.34/0.08 (4.00–4.25), telotarsus 0.87–0.88/0.05–0.06 (14.67–17.40).</paragraph>
|
||
<paragraph id="027236B3FFC9A921FF41F87AFE06FE8A" blockId="7.[151,1437,964,2035]" lastBlockId="10.[151,1437,150,861]" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">Females: body length 1.95–2.01. Pedipalps: trochanter 0.31–0.32/0.16–0.17 (1.82–2.20), femur 1.18– 1.20/0.15–0.16 (7.50–7.87), patella 0.42–0.47/0.17–0.19 (2.47), chela 1.55–1.60/0.24–0.25 (6.40–6.46), hand 0.61– 0.64/0.24–0.25 (2.44–2.67), movable chelal finger length 0.96–0.98. Chelicera 0.62–0.64/0.27–0.30 (2.13–2.30), movable finger length 0.34–0.35.Carapace 0.56–0.57/0.58–0.60 (0.95–0.97).Leg I: trochanter0.20–0.21/0.12 (1.67– 1.75), femur 0.69–0.71/0.07–0.08 (8.63–10.14), patella 0.35–0.37/0.07 (5.00–5.29), tibia 0.34/0.06–0.07 (4.86– 5.67), tarsus 0.72/0.05–0.06 (12.00–14.40). Leg IV: trochanter 0.24–0.25/0.14–0.16 (1.50–1.79), femoropatella 0.99–1.07/0.22–0.24 (4.46–4.50), tibia 0.65–0.69/0.10 (6.50–6.90), basitarsus 0.34/0.08 (4.25), telotarsus 0.83– 0.90/0.05–0.06 (15.00–16.60).</paragraph>
|
||
</subSubSection>
|
||
<caption id="56B2663BFFC6A923FF11F899FF33F818" ID-DOI="http://doi.org/10.5281/zenodo.7251612" ID-Zenodo-Dep="7251612" httpUri="https://zenodo.org/record/7251612/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1907,1931]" targetBox="[151,1436,202,1883]" targetPageId="8">
|
||
<paragraph id="027236B3FFC6A923FF11F899FF33F818" blockId="8.[151,1436,1907,2003]" pageId="8" pageNumber="9">
|
||
<emphasis id="30B9EAA1FFC6A923FF11F899FEFEF840" bold="true" box="[151,277,1907,1931]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFC6A923FE99F899FDC9F840" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[287,546,1907,1931]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC6A923FE99F899FDC9F840" box="[287,546,1907,1931]" italics="true" pageId="8" pageNumber="9">Lagynochthonius crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC6A923FDADF899FD95F840" bold="true" box="[555,638,1907,1931]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="2B8A57DAFFC6A923FDADF899FD95F840" box="[555,638,1907,1931]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="56B2663BFFC7A922FF11F8CFFED4F839" ID-DOI="http://doi.org/10.5281/zenodo.7251616" ID-Zenodo-Dep="7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1829,1853]" targetBox="[186,1403,183,1797]" targetPageId="9">
|
||
<paragraph id="027236B3FFC7A922FF11F8CFFED4F839" blockId="9.[151,1437,1829,2034]" pageId="9" pageNumber="10">
|
||
<emphasis id="30B9EAA1FFC7A922FF11F8CFFEFEF8F5" bold="true" box="[151,277,1829,1854]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFC7A922FE98F8CFFDCAF8F6" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[286,545,1829,1853]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC7A922FE98F8CFFDCAF8F6" box="[286,545,1829,1853]" italics="true" pageId="9" pageNumber="10">Lagynochthonius crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC7A922FDACF8CCFD96F8F5" bold="true" box="[554,637,1830,1854]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="2B8A57DAFFC7A922FDACF8CCFD96F8F5" box="[554,637,1830,1854]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4AD76538FFC4A921FF41FEA1FEBDFD62" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="027236B3FFC4A921FF41FEA1FD93FE1A" blockId="10.[151,1437,150,861]" pageId="10" pageNumber="11">
|
||
<emphasis id="30B9EAA1FFC4A921FF41FEA1FED0FEAE" bold="true" box="[199,315,331,357]" pageId="10" pageNumber="11">Remarks.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFC4A921FEB9FEA1FDB3FEAF" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[319,600,331,357]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFC4A921FEB9FEA1FDB3FEAF" box="[319,600,331,357]" italics="true" pageId="10" pageNumber="11">Lagynochthonius crassus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFC4A921FDDAFEA1FD59FEAE" bold="true" box="[604,690,331,357]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="2B8A57DAFFC4A921FDDAFEA1FD59FEAE" box="[604,690,331,357]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="C5CD4D30FFC4A921FC55FEA1FB74FEAE" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[979,1183,331,357]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis id="30B9EAA1FFC4A921FC55FEA1FB74FEAE" box="[979,1183,331,357]" italics="true" pageId="10" pageNumber="11">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C5CD4D30FFC4A921FB56FEA1FA7EFEAE" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[1232,1429,331,357]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis id="30B9EAA1FFC4A921FB56FEA1FA7EFEAE" box="[1232,1429,331,357]" italics="true" pageId="10" pageNumber="11">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
, as well as the 10 new troglomorphic
|
||
<taxonomicName id="C5CD4D30FFC4A921FDABFE85FCAEFE42" box="[557,837,367,393]" pageId="10" pageNumber="11">
|
||
<emphasis id="30B9EAA1FFC4A921FDABFE85FD05FE42" box="[557,750,367,393]" italics="true" pageId="10" pageNumber="11">Lagynochthonius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described below, by the presence of a pair of eyespots (not corneate eyes) and a pair of thickened chelal hands (e.g. chela 7.00–7.14 times longer than broad, hand 2.73– 2.82 times longer than broad, all in males).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFC4A921FF41FE31FEBDFD62" blockId="10.[151,1437,150,861]" pageId="10" pageNumber="11">
|
||
The new species is similar to another troglomorphic species from continental Asia:
|
||
<taxonomicName id="C5CD4D30FFC4A921FB0AFE31FEE7FDD2" authority="Judson, 2007" authorityName="Judson" authorityYear="2007" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fragilis">
|
||
<emphasis id="30B9EAA1FFC4A921FB0AFE31FB16FE3E" box="[1164,1277,475,501]" italics="true" pageId="10" pageNumber="11">L. fragilis</emphasis>
|
||
<bibRefCitation id="665C4B42FFC4A921FA80FE31FA77FE3E" author="Judson, M. L. I." box="[1286,1436,475,501]" pageId="10" pageNumber="11" pagination="53 - 68" refId="ref38953" refString="Judson, M. L. I. (2007) A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae). Zootaxa, 1627, 53 - 68. https: // doi. org / 10.11646 / zootaxa. 1627.1.4" type="journal article" year="2007">Judson, 2007</bibRefCitation>
|
||
(
|
||
<collectingCountry id="7ADA7623FFC4A921FF19FE15FEEFFDD2" box="[159,260,511,537]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
, by having a pair of eyespots, the same chaetotaxy of carapace (s4s-4-4-2-2) and the presence of same number of setae on tergites I–V, but differs in having more slender pedipalps (e.g. femur 7.50–7.87 times longer than broad
|
||
<emphasis id="30B9EAA1FFC4A921FE92FDADFED8FDAB" box="[276,307,583,608]" italics="true" pageId="10" pageNumber="11">vs.</emphasis>
|
||
6.3–6.6 times in females, chela length 1.55–1.60
|
||
<emphasis id="30B9EAA1FFC4A921FCDAFDADFC90FDAB" box="[860,891,583,608]" italics="true" pageId="10" pageNumber="11">vs.</emphasis>
|
||
<quantity id="C5359B56FFC4A921FC07FDADFBC8FDAA" box="[897,1059,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.255" metricValueMax="1.32" metricValueMin="1.19" pageId="10" pageNumber="11" unit="mm" value="1.255" valueMax="1.32" valueMin="1.19">1.19–1.32 mm</quantity>
|
||
in females), the presence of more chelal teeth, and a larger body size (body length 1.95–2.01
|
||
<emphasis id="30B9EAA1FFC4A921FCBFFD81FCB3FD4F" box="[825,856,619,644]" italics="true" pageId="10" pageNumber="11">vs.</emphasis>
|
||
<quantity id="C5359B56FFC4A921FCE6FD81FC2EFD4E" box="[864,965,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="11" unit="mm" value="1.6">1.60 mm</quantity>
|
||
in females) (
|
||
<bibRefCitation id="665C4B42FFC4A921FBDDFD81FB06FD4E" author="Judson, M. L. I." box="[1115,1261,619,645]" pageId="10" pageNumber="11" pagination="53 - 68" refId="ref38953" refString="Judson, M. L. I. (2007) A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae). Zootaxa, 1627, 53 - 68. https: // doi. org / 10.11646 / zootaxa. 1627.1.4" type="journal article" year="2007">Judson 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="665C4B42FFC4A921FB7CFD81FA7CFD4E" author="Li, Y. C. & Liu, H. & Shi, A. M." box="[1274,1431,619,645]" pageId="10" pageNumber="11" pagination="28 - 34" refId="ref39131" refString="Li, Y. C., Liu, H. & Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">
|
||
Li
|
||
<emphasis id="30B9EAA1FFC4A921FA9DFD81FABEFD4E" box="[1307,1365,619,645]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="665C4B42FFC4A921FF11FD65FEA1FD62" author="Hou, Y. M. & Gao, Z. Z. & Zhang, F." box="[151,330,655,681]" pageId="10" pageNumber="11" pagination="65 - 83" refId="ref38684" refString="Hou, Y. M., Gao, Z. Z. & Zhang, F. (2022) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys, 1097, 65 - 83. https: // doi. org / 10.3897 / zookeys. 1097.82527" type="journal article" year="2022">
|
||
Hou
|
||
<emphasis id="30B9EAA1FFC4A921FF48FD65FEECFD62" box="[206,263,655,681]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFC4A921FF41FD58FD05FC96" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="027236B3FFC4A921FF41FD58FD05FC96" blockId="10.[151,1437,150,861]" pageId="10" pageNumber="11">
|
||
<emphasis id="30B9EAA1FFC4A921FF41FD58FE12FD07" bold="true" box="[199,505,690,717]" pageId="10" pageNumber="11">Distribution and habitat.</emphasis>
|
||
This species is only known from the
|
||
<typeStatus id="DD768811FFC4A921FC3BFD59FC06FD06" box="[957,1005,691,717]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality, Unnamed Cave 1 (
|
||
<figureCitation id="9AF62A36FFC4A921FABFFD59FA7CFD06" box="[1337,1431,691,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); M–N. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="10" pageNumber="11">Figs 1L</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFC4A921FF11FD3DFF35FD3A" box="[151,222,727,753]" captionStart="FIGURE 2" captionStartId="6.[151,250,1944,1968]" captionTargetBox="[151,1436,273,1921]" captionTargetId="figure-80@6.[151,1436,273,1921]" captionTargetPageId="6" captionText="FIGURE 2. Unnamed Cave 1, type locality of Lagynochthonius crassus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where L. crassus sp. nov. specimens were collected; D–E. Live individuals of L. crassus sp. nov. (D: male, E: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251608" httpUri="https://zenodo.org/record/7251608/files/figure.png" pageId="10" pageNumber="11">2A–C</figureCitation>
|
||
), which is located about
|
||
<quantity id="C5359B56FFC4A921FE71FD3DFDDAFD3A" box="[503,561,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="km" value="1.0">1 km</quantity>
|
||
northwest of Laomenzhai Village (Maguan County). This limestone cave has one triangular entrance (about 1.5 meters high and
|
||
<quantity id="C5359B56FFC4A921FD54FD11FCCCFCDE" box="[722,807,763,789]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="1.0">1 meter</quantity>
|
||
wide), approximately 200 meters in length. The interior space is large and extends horizontally. All of the specimens were collected under stones and detritus in the deep zone (temperature around 14°C, humidity over 85%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |