treatments-xml/data/8A/64/87/8A6487A5FFC4A924FF11FC44FC9CFB22.xml
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<document id="71CFF4178BFA6418124AF3C6D4868CE7" ID-CLB-Dataset="172790" ID-DOI="10.11646/zootaxa.5198.1.1" ID-GBIF-Dataset="17babbe7-1645-47c8-ac5f-2299f1ad436c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251602" ID-ZooBank="019FE488-4263-4BC2-8606-446E599E226A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666769542164" checkinUser="plazi" docAuthor="Hou, Yanmeng, Gao, Zhizhong &amp; Zhang, Feng" docDate="2022" docId="8A6487A5FFC4A924FF11FC44FC9CFB22" docLanguage="en" docName="zootaxa.5198.1.1.pdf" docOrigin="Zootaxa 5198 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5198.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Lagynochthonius fengi Hou &amp; Gao &amp; Zhang 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="765DFFDDFFCEA92BFF86FFEAFFEBFFCB" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="11" updateTime="1701997268366" updateUser="plazi">
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<mods:title id="33C7B30EC97E196E93E4FFDB9D2F82C7">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="B86554D820F328B20927AAA37F9B0087">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="AE3694C5645E6B064A1157B81B0D16FE">Gao, Zhizhong</mods:namePart>
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<mods:namePart id="478070D6CD35E9843E6ACCB779A55136">Zhang, Feng</mods:namePart>
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<treatment id="8A6487A5FFC4A924FF11FC44FC9CFB22" ID-DOI="http://doi.org/10.5281/zenodo.7255275" ID-GBIF-Taxon="203411091" ID-Zenodo-Dep="7255275" LSID="urn:lsid:plazi:treatment:8A6487A5FFC4A924FF11FC44FC9CFB22" httpUri="http://treatment.plazi.org/id/8A6487A5FFC4A924FF11FC44FC9CFB22" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11">
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<taxonomicName id="C5CD4D30FFC4A921FF11FC44FE5BFC03" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[151,432,942,969]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC4A921FF11FC44FE5BFC03" bold="true" box="[151,432,942,969]" italics="true" pageId="10" pageNumber="11">Lagynochthonius fengi</emphasis>
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<taxonomicNameLabel id="2B8A57DAFFC4A921FE3EFC45FDFEFC02" box="[440,533,943,969]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(Ã氏IJ伪oi)
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<figureCitation id="9AF62A36FFC4A921FF11FC39FF13FC26" box="[151,248,979,1005]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="11.[151,250,1543,1567]" captionStartId-1="12.[151,250,1493,1517]" captionStartId-2="13.[151,250,1927,1951]" captionStartId-3="14.[151,250,1874,1898]" captionTargetBox-0="[151,1437,381,1520]" captionTargetBox-1="[151,1417,649,1470]" captionTargetBox-2="[151,1435,181,1903]" captionTargetBox-3="[179,1409,181,1850]" captionTargetId-0="figure-114@11.[151,1437,381,1520]" captionTargetId-1="figure-252@12.[151,1436,633,1470]" captionTargetId-2="figure-19@13.[151,1436,181,1903]" captionTargetId-3="figure-20@14.[179,1409,181,1850]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 6. Unnamed Cave 2, type locality of Lagynochthonius fengi sp. nov., A. Entrance; B. Inside the cave entrance; CD. Areas where L. fengi sp. nov. specimens were collected; EF. Live individuals of L. fengi sp. nov. (E: male, F: female) in their natural environment." captionText-1="FIGURE 7. Lagynochthonius fengi sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." captionText-3="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.7251620" figureDoi-1="http://doi.org/10.5281/zenodo.7251622" figureDoi-2="http://doi.org/10.5281/zenodo.7251626" figureDoi-3="http://doi.org/10.5281/zenodo.7251630" httpUri-0="https://zenodo.org/record/7251620/files/figure.png" httpUri-1="https://zenodo.org/record/7251622/files/figure.png" httpUri-2="https://zenodo.org/record/7251626/files/figure.png" httpUri-3="https://zenodo.org/record/7251630/files/figure.png" pageId="10" pageNumber="11">Figs 69</figureCitation>
</paragraph>
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<paragraph id="027236B3FFC4A921FF11FBF1FC8DFB0E" blockId="10.[151,1437,1050,2013]" pageId="10" pageNumber="11">
<emphasis id="30B9EAA1FFC4A921FF11FBF1FEA6FBFF" bold="true" box="[151,333,1050,1077]" pageId="10" pageNumber="11">Type material.</emphasis>
<materialsCitation id="B2A53CEEFFC4A921FEDBFBF1FAC9FBB6" ID-GBIF-Occurrence="3946861327" collectingDate="2021-09-30" collectionCode="MHBU-HBUARA" collectorName="Zegang Feng &amp; Yanmeng Hou &amp; Lu Zhang &amp; Liu Fu" country="China" county="Yanjin County" elevation="594" latitude="27.91409" location="Maliu Village" longLatPrecision="1" longitude="104.39159" municipality="Miaoba Town" pageId="10" pageNumber="11" specimenCode="MHBU-HBUARA#2021-41701" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="DD768811FFC4A921FEDBFBF1FE2EFBFE" box="[349,453,1051,1077]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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(Ps.-
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):
<collectingCountry id="7ADA7623FFC4A921FC30FBF1FC17FBFE" box="[950,1020,1051,1077]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
,
<collectingRegion id="C009F851FFC4A921FB88FBF1FB32FBFE" box="[1038,1241,1051,1077]" country="China" name="Yunnan" pageId="10" pageNumber="11">Yunnan Province</collectingRegion>
,
<collectingCounty id="EB134E3FFFC4A921FB6AFBF1FA72FBFE" box="[1260,1433,1051,1077]" pageId="10" pageNumber="11">Yanjin County</collectingCounty>
,
<collectingMunicipality id="E216ACC9FFC4A921FF11FBD5FEDFFB92" box="[151,308,1087,1113]" pageId="10" pageNumber="11">Miaoba Town</collectingMunicipality>
,
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, cave without name (
<location id="07126068FFC4A921FD44FBD5FC9BFB92" LSID="urn:lsid:plazi:treatment:8A6487A5FFC4A924FF11FC44FC9CFB22:07126068FFC4A921FD44FBD5FC9BFB92" box="[706,880,1087,1113]" country="China" county="Yanjin County" latitude="27.91409" longLatPrecision="1" longitude="104.39159" municipality="Miaoba Town" name="Unnamed Cave" pageId="10" pageNumber="11" stateProvince="Yunnan">Unnamed Cave</location>
2), under the stones in deep zone [
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,
<geoCoordinate id="67F95074FFC4A921FF11FB8EFEBBFBB4" box="[151,336,1124,1151]" degrees="104" direction="east" minutes="23" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="29.72" value="104.39159">104°2329.72″E</geoCoordinate>
],
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<quantity id="C5359B56FFC4A921FEE6FB88FE4DFBB6" box="[352,422,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="10" pageNumber="11" unit="m" value="594.0">594 m</quantity>
a.s.l.
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,
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<collectingDate id="6637E99BFFC4A921FE6BFB89FD2CFBB6" box="[493,711,1123,1149]" pageId="10" pageNumber="11" value="2021-09-30">30 September 2021</collectingDate>
</date>
,
<collectorName id="AF385365FFC4A921FD57FB89FC8EFBB6" box="[721,869,1123,1149]" pageId="10" pageNumber="11">Zegang Feng</collectorName>
,
<collectorName id="AF385365FFC4A921FCE8FB89FBFAFBB6" box="[878,1041,1123,1149]" pageId="10" pageNumber="11">Yanmeng Hou</collectorName>
,
<collectorName id="AF385365FFC4A921FB9DFB89FB63FBB6" box="[1051,1160,1123,1149]" pageId="10" pageNumber="11">Lu Zhang</collectorName>
&amp;
<collectorName id="AF385365FFC4A921FB2FFB89FB18FBB6" box="[1193,1267,1123,1149]" pageId="10" pageNumber="11">Liu Fu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="B2A53CEEFFC4A921FAAEFB89FC71FB6A" ID-GBIF-Occurrence="3946861305" collectionCode="MHBU-HBUARA" country="China" pageId="10" pageNumber="11" specimenCode="MHBU-HBUARA#2021-41702" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="DD768811FFC4A921FAAEFB89FA72FBB6" box="[1320,1433,1123,1149]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="14CBFD3AFFC4A921FF11FB6DFF20FB6A" box="[151,203,1159,1185]" pageId="10" pageNumber="11" type="male">3 ♂</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFC4A921FE8EFB6DFD91FB6A" box="[264,634,1159,1185]" collectionCode="MHBU-HBUARA" pageId="10" pageNumber="11">MHBU-HBUARA#2021-41702</specimenCode>
-HBUARA#2021-41704)
</materialsCitation>
,
<materialsCitation id="B2A53CEEFFC4A921FC28FB6DFC88FB0E" ID-GBIF-Occurrence="3946861323" collectingDate="2021-09-30" collectionCode="MSWU-HBUARA" collectorName="Zegang Feng &amp; Yanmeng Hou &amp; Lu Zhang &amp; Liu Fu" country="China" county="Yanjin County" elevation="594" latitude="27.91409" location="Maliu Village" longLatPrecision="1" longitude="104.39159" municipality="Miaoba Town" pageId="10" pageNumber="11" specimenCode="MSWU-HBUARA#2021-41705" specimenCount="9" specimenCount-female="9" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="14CBFD3AFFC4A921FC28FB6DFC09FB6A" box="[942,994,1159,1185]" pageId="10" pageNumber="11" type="female">9 ♀</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFC4A921FB99FB6CFA7EFB6A" box="[1055,1429,1158,1185]" collectionCode="MSWU-HBUARA" pageId="10" pageNumber="11">MSWU-HBUARA#2021-41705</specimenCode>
- HBUARA#2021-41713), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFC4A921FF41FB25FEFFFAC6" pageId="10" pageNumber="11" type="etymology">
<paragraph id="027236B3FFC4A921FF41FB25FEFFFAC6" blockId="10.[151,1437,1050,2013]" pageId="10" pageNumber="11">
<emphasis id="30B9EAA1FFC4A921FF41FB25FEA5FB22" bold="true" box="[199,334,1231,1257]" pageId="10" pageNumber="11">Etymology.</emphasis>
This species is named for Zegang Feng, who participated in field work and collected some of the specimens.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFC4A921FF41FAFCFCD5FA2E" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="027236B3FFC4A921FF41FAFCFCD5FA2E" blockId="10.[151,1437,1050,2013]" pageId="10" pageNumber="11">
<emphasis id="30B9EAA1FFC4A921FF41FAFCFE6DFAFF" bold="true" box="[199,390,1302,1332]" pageId="10" pageNumber="11">
Diagnosis (
<specimenCount id="14CBFD3AFFC4A921FECBFAF2FE89FAFF" box="[333,354,1304,1332]" pageId="10" pageNumber="11" type="male"></specimenCount>
<specimenCount id="14CBFD3AFFC4A921FEE4FAF2FE9CFAFF" box="[354,375,1304,1332]" pageId="10" pageNumber="11" type="female"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with 2 setae; tergites IIV each with 2 setae. Pedipalps slender, femur 7.577.64 (
<specimenCount id="14CBFD3AFFC4A921FA86FAB5FAFCFAB2" box="[1280,1303,1375,1401]" pageId="10" pageNumber="11" type="male"></specimenCount>
), 7.718.21 (
<specimenCount id="14CBFD3AFFC4A921FF24FA69FF5FFA56" box="[162,180,1411,1437]" pageId="10" pageNumber="11" type="female"></specimenCount>
) times longer than broad; chela 7.007.29 (
<specimenCount id="14CBFD3AFFC4A921FD22FA69FD50FA56" box="[676,699,1411,1437]" pageId="10" pageNumber="11" type="male"></specimenCount>
), 6.957.05 (
<specimenCount id="14CBFD3AFFC4A921FCD5FA69FC8EFA56" box="[851,869,1411,1437]" pageId="10" pageNumber="11" type="female"></specimenCount>
) times longer than broad; both chelal fingers with intercalary teeth and fixed chelal finger with a modified accessory tooth (
<emphasis id="30B9EAA1FFC4A921FC4EFA4DFC35FA0A" box="[968,990,1447,1473]" italics="true" pageId="10" pageNumber="11">td</emphasis>
) on dorso-antiaxial face; chemosensory setae (
<emphasis id="30B9EAA1FFC4A921FF59FA21FF1DFA2F" box="[223,246,1483,1508]" italics="true" pageId="10" pageNumber="11">sc</emphasis>
) present on dorsum of chelal hand; sensilla present.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFC4A924FF41FA04FABDFD62" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11" type="description">
<paragraph id="027236B3FFC4A921FF41FA04FC3CF9C2" blockId="10.[151,1437,1050,2013]" box="[199,983,1518,1545]" pageId="10" pageNumber="11">
<emphasis id="30B9EAA1FFC4A921FF41FA04FEB2F9C3" bold="true" box="[199,345,1518,1544]" pageId="10" pageNumber="11">Description.</emphasis>
Males (
<typeStatus id="DD768811FFC4A921FE35FA05FDFCF9C2" box="[435,535,1519,1545]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
and
<typeStatus id="DD768811FFC4A921FDCBFA05FD51F9C2" box="[589,698,1519,1545]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="9AF62A36FFC4A921FD56FA05FCC1F9C2" box="[720,810,1519,1545]" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[151,1437,381,1520]" captionTargetId="figure-114@11.[151,1437,381,1520]" captionTargetPageId="11" captionText="FIGURE 6. Unnamed Cave 2, type locality of Lagynochthonius fengi sp. nov., A. Entrance; B. Inside the cave entrance; CD. Areas where L. fengi sp. nov. specimens were collected; EF. Live individuals of L. fengi sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251620" httpUri="https://zenodo.org/record/7251620/files/figure.png" pageId="10" pageNumber="11">Figs 6E</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FCB0FA05FCB0F9C2" box="[822,859,1519,1545]" captionStart="FIGURE 7" captionStartId="12.[151,250,1493,1517]" captionTargetBox="[151,1417,649,1470]" captionTargetId="figure-252@12.[151,1436,633,1470]" captionTargetPageId="12" captionText="FIGURE 7. Lagynochthonius fengi sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251622" httpUri="https://zenodo.org/record/7251622/files/figure.png" pageId="10" pageNumber="11">7A</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FCE0FA05FC46F9C2" box="[870,941,1519,1545]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="10" pageNumber="11">8AD</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FC3FFA05FC22F9C2" box="[953,969,1519,1545]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="10" pageNumber="11">9</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFC4A921FF41F9F9FF26F99A" blockId="10.[151,1437,1050,2013]" pageId="10" pageNumber="11">
Colour (
<figureCitation id="9AF62A36FFC4A921FEA5F9F9FE96F9E6" box="[291,381,1555,1581]" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[151,1437,381,1520]" captionTargetId="figure-114@11.[151,1437,381,1520]" captionTargetPageId="11" captionText="FIGURE 6. Unnamed Cave 2, type locality of Lagynochthonius fengi sp. nov., A. Entrance; B. Inside the cave entrance; CD. Areas where L. fengi sp. nov. specimens were collected; EF. Live individuals of L. fengi sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251620" httpUri="https://zenodo.org/record/7251620/files/figure.png" pageId="10" pageNumber="11">Figs 6E</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FE0EF9F9FE46F9E6" box="[392,429,1555,1581]" captionStart="FIGURE 7" captionStartId="12.[151,250,1493,1517]" captionTargetBox="[151,1417,649,1470]" captionTargetId="figure-252@12.[151,1436,633,1470]" captionTargetPageId="12" captionText="FIGURE 7. Lagynochthonius fengi sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251622" httpUri="https://zenodo.org/record/7251622/files/figure.png" pageId="10" pageNumber="11">7A</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FE3EF9F9FDEAF9E6" box="[440,513,1555,1581]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="10" pageNumber="11">8AD</figureCitation>
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
</paragraph>
<paragraph id="027236B3FFC4A921FF41F9B1FC0CF85E" blockId="10.[151,1437,1050,2013]" pageId="10" pageNumber="11">
Cephalothorax (
<figureCitation id="9AF62A36FFC4A921FEFCF9B1FE3FF9BE" box="[378,468,1627,1653]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="10" pageNumber="11">Figs 8B</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FE59F9B1FDEDF9BE" box="[479,518,1627,1653]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="10" pageNumber="11">9A</figureCitation>
): carapace 1.001.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 1213 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="9AF62A36FFC4A921FEDFF891FE59F85E" box="[345,434,1915,1941]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="10" pageNumber="11">Fig. 9C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="027236B3FFC4A920FF41F875FDB5FEAE" blockId="10.[151,1437,1050,2013]" lastBlockId="11.[151,1436,151,357]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Chelicera (
<figureCitation id="9AF62A36FFC4A921FEC2F875FE4AF872" box="[324,417,1951,1977]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="10" pageNumber="11">Figs 8C</figureCitation>
,
<figureCitation id="9AF62A36FFC4A921FE36F875FE3FF872" box="[432,468,1951,1977]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="10" pageNumber="11">9B</figureCitation>
): large, about as long as carapace, 2.332.43 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 1415 teeth, distal one largest; movable finger with 1213 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="9AF62A36FFC5A920FBFAFF35FB3DFF32" box="[1148,1238,223,249]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="11" pageNumber="12">Fig. 9B</figureCitation>
). Serrula exterior with 2123 and serrula interior with 1314 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="9AF62A36FFC5A920FE70FEA1FDBAFEAE" box="[502,593,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="11" pageNumber="12">Fig. 9D</figureCitation>
).
</paragraph>
<caption id="56B2663BFFC5A920FF11F9EDFE80F9AC" ID-DOI="http://doi.org/10.5281/zenodo.7251620" ID-Zenodo-Dep="7251620" httpUri="https://zenodo.org/record/7251620/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1543,1567]" targetBox="[151,1437,381,1520]" targetPageId="11">
<paragraph id="027236B3FFC5A920FF11F9EDFE80F9AC" blockId="11.[151,1437,1543,1639]" pageId="11" pageNumber="12">
<emphasis id="30B9EAA1FFC5A920FF11F9EDFEF8F9D4" bold="true" box="[151,275,1543,1567]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
Unnamed Cave 2, type locality of
<taxonomicName id="C5CD4D30FFC5A920FDF0F9EDFCB5F9D4" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[630,862,1543,1567]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC5A920FDF0F9EDFCB5F9D4" box="[630,862,1543,1567]" italics="true" pageId="11" pageNumber="12">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC5A920FCE2F9EDFC5EF9D4" bold="true" box="[868,949,1543,1567]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="2B8A57DAFFC5A920FCE2F9EDFC5EF9D4" box="[868,949,1543,1567]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; CD. Areas where
<taxonomicName id="C5CD4D30FFC5A920FE9BF9C6FE80F988" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[285,363,1579,1603]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC5A920FE9BF9C6FE80F988" box="[285,363,1579,1603]" italics="true" pageId="11" pageNumber="12">L. fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC5A920FEF7F9C1FE29F988" bold="true" box="[369,450,1579,1603]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="2B8A57DAFFC5A920FEF7F9C1FE29F988" box="[369,450,1579,1603]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; EF. Live individuals of
<taxonomicName id="C5CD4D30FFC5A920FC53F9C6FBC8F988" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[981,1059,1579,1603]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC5A920FC53F9C6FBC8F988" box="[981,1059,1579,1603]" italics="true" pageId="11" pageNumber="12">L. fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC5A920FBAFF9C1FB91F988" bold="true" box="[1065,1146,1579,1603]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="2B8A57DAFFC5A920FBAFF9C1FB91F988" box="[1065,1146,1579,1603]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(E: male, F: female) in their natural environment.
</paragraph>
</caption>
<paragraph id="027236B3FFC5A927FF41F97FFD21FE1A" blockId="11.[151,1437,1685,2036]" lastBlockId="12.[151,1437,151,609]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Pedipalp (
<figureCitation id="9AF62A36FFC5A920FEBFF97FFE73F97B" box="[313,408,1685,1712]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="11" pageNumber="12">Figs 8A</figureCitation>
,
<figureCitation id="9AF62A36FFC5A920FE23F97CFE07F97B" box="[421,492,1686,1712]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="11" pageNumber="12">9EG</figureCitation>
): long and slender, trochanter 1.691.73, femur 7.577.64, patella 2.182.44, chela 7.007.29, hand 2.863.05 times longer than broad; femur 2.742.86 times longer than patella; movable chelal finger 1.381.44 times longer than hand and 0.580.59 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="9AF62A36FFC5A920FDB3F8EBFD61F8D7" box="[565,650,1793,1820]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="11" pageNumber="12">Fig. 9E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="30B9EAA1FFC5A920FB0EF8A3FB75F8A8" box="[1160,1182,1865,1891]" italics="true" pageId="11" pageNumber="12">ib</emphasis>
and
<emphasis id="30B9EAA1FFC5A920FB5FF8A3FB11F8A8" box="[1241,1274,1865,1891]" italics="true" pageId="11" pageNumber="12">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="30B9EAA1FFC5A920FD35F887FD26F84C" box="[691,717,1901,1927]" italics="true" pageId="11" pageNumber="12">eb</emphasis>
,
<emphasis id="30B9EAA1FFC5A920FD5CF887FD14F84C" box="[730,767,1901,1927]" italics="true" pageId="11" pageNumber="12">esb</emphasis>
and
<emphasis id="30B9EAA1FFC5A920FCB2F884FCA5F84C" box="[820,846,1902,1927]" italics="true" pageId="11" pageNumber="12">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis id="30B9EAA1FFC5A920FF6CF878FF11F860" box="[234,250,1938,1963]" italics="true" pageId="11" pageNumber="12">it</emphasis>
slightly distal to
<emphasis id="30B9EAA1FFC5A920FE38F878FE36F860" box="[446,477,1938,1963]" italics="true" pageId="11" pageNumber="12">est</emphasis>
, situated subdistally;
<emphasis id="30B9EAA1FFC5A920FD52F878FD03F860" box="[724,744,1938,1963]" italics="true" pageId="11" pageNumber="12">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="30B9EAA1FFC5A920FF5DF85FFF1EF804" box="[219,245,1973,1999]" italics="true" pageId="11" pageNumber="12">dx</emphasis>
situated distal to
<emphasis id="30B9EAA1FFC5A920FE3EF85CFE27F804" box="[440,460,1974,1999]" italics="true" pageId="11" pageNumber="12">et</emphasis>
;
<emphasis id="30B9EAA1FFC5A920FE5CF85FFE18F804" box="[474,499,1973,1999]" italics="true" pageId="11" pageNumber="12">sb</emphasis>
situated midway between
<emphasis id="30B9EAA1FFC5A920FC9FF85FFCCCF804" box="[793,807,1973,1999]" italics="true" pageId="11" pageNumber="12">b</emphasis>
and
<emphasis id="30B9EAA1FFC5A920FCDAF85CFC84F804" box="[860,879,1974,1999]" italics="true" pageId="11" pageNumber="12">st</emphasis>
;
<emphasis id="30B9EAA1FFC5A920FCFBF85FFC60F804" box="[893,907,1973,1999]" italics="true" pageId="11" pageNumber="12">b</emphasis>
and
<emphasis id="30B9EAA1FFC5A920FC39F85CFC2CF804" box="[959,967,1974,1999]" italics="true" pageId="11" pageNumber="12">t</emphasis>
situated subdistally,
<emphasis id="30B9EAA1FFC5A920FB29F85CFB5CF804" box="[1199,1207,1974,1999]" italics="true" pageId="11" pageNumber="12">t</emphasis>
situated between
<emphasis id="30B9EAA1FFC5A920FAFBF85CFA77F804" box="[1405,1436,1974,1999]" italics="true" pageId="11" pageNumber="12">est</emphasis>
and
<emphasis id="30B9EAA1FFC5A920FF43F830FF3EF838" box="[197,213,2010,2035]" italics="true" pageId="11" pageNumber="12">it</emphasis>
and distal to
<emphasis id="30B9EAA1FFC5A920FEE1F833FE9EF838" box="[359,373,2009,2035]" italics="true" pageId="11" pageNumber="12">b</emphasis>
;
<emphasis id="30B9EAA1FFC5A920FE05F830FE49F838" box="[387,418,2010,2035]" italics="true" pageId="11" pageNumber="12">est</emphasis>
situated distal to
<emphasis id="30B9EAA1FFC5A920FDE5F833FD9AF838" box="[611,625,2009,2035]" italics="true" pageId="11" pageNumber="12">b</emphasis>
(
<figureCitation id="9AF62A36FFC5A920FDF9F833FD38F83F" box="[639,723,2009,2036]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="11" pageNumber="12">Fig. 9F</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="30B9EAA1FFC2A927FEDFFF7DFE98FF7B" box="[345,371,151,176]" italics="true" pageId="12" pageNumber="13">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 2325 macrodenticles, slightly retrorse and pointed, plus 14
<specimenCount id="14CBFD3AFFC2A927FCBAFF34FC3EFF32" box="[828,981,222,249]" pageId="12" pageNumber="13" type="intercalary">15 intercalary</specimenCount>
microdenticles and a modified accessory tooth on dorso-antiaxial face (
<emphasis id="30B9EAA1FFC2A927FE65FEE9FE12FED6" box="[483,505,259,285]" italics="true" pageId="12" pageNumber="13">td</emphasis>
, near tip),
<quantity id="C5359B56FFC2A927FDE9FEE9FD3AFED6" box="[623,721,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.0032999999999999" metricValueMax="1.0413999999999999" metricValueMin="0.9652" pageId="12" pageNumber="13" unit="in" value="39.5" valueMax="41.0" valueMin="38.0">3841 in</quantity>
total; movable chelal finger with 1719 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 11
<specimenCount id="14CBFD3AFFC2A927FBA5FECDFB54FE8A" box="[1059,1215,295,321]" pageId="12" pageNumber="13" type="intercalary">12 intercalary</specimenCount>
microdenticles and 1113 vestigial, rounded and contiguous basal teeth,
<quantity id="C5359B56FFC2A927FD6CFEA1FCA6FEAE" box="[746,845,331,357]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.0668" metricValueMin="1.0413999999999999" pageId="12" pageNumber="13" unit="in" value="41.5" valueMax="42.0" valueMin="41.0">4142 in</quantity>
total (
<figureCitation id="9AF62A36FFC2A927FC13FEA1FC00FEAE" box="[917,1003,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="12" pageNumber="13">Fig. 9F</figureCitation>
). Fixed chelal finger with sensilla
<emphasis id="30B9EAA1FFC2A927FAE8FEA1FA6FFEAE" box="[1390,1412,331,357]" italics="true" pageId="12" pageNumber="13">af</emphasis>
<subScript id="9E4934F6FFC2A927FA02FEB7FA77FEA0" attach="left" box="[1412,1436,349,363]" fontSize="6" pageId="12" pageNumber="13">12</subScript>
close together, near tip; movable chelal finger with 4 sensilla:
<emphasis id="30B9EAA1FFC2A927FCD5FE85FC9EFE43" box="[851,885,367,392]" italics="true" pageId="12" pageNumber="13">am</emphasis>
<subScript id="9E4934F6FFC2A927FCF3FE6BFC66FE44" attach="left" box="[885,909,385,399]" fontSize="6" pageId="12" pageNumber="13">12</subScript>
near tip,
<emphasis id="30B9EAA1FFC2A927FC7EFE85FBEDFE43" box="[1016,1030,367,392]" italics="true" pageId="12" pageNumber="13">p</emphasis>
<subScript id="9E4934F6FFC2A927FB80FE6BFBE5FE44" attach="left" box="[1030,1038,385,399]" fontSize="6" pageId="12" pageNumber="13">1</subScript>
slightly distad of
<emphasis id="30B9EAA1FFC2A927FB5CFE85FB18FE42" box="[1242,1267,367,393]" italics="true" pageId="12" pageNumber="13">sb</emphasis>
and very close to chelal teeth,
<emphasis id="30B9EAA1FFC2A927FEB9FE79FEA6FE67" box="[319,333,403,428]" italics="true" pageId="12" pageNumber="13">p</emphasis>
<subScript id="9E4934F6FFC2A927FECBFE4FFEBEFE78" attach="left" box="[333,341,421,435]" fontSize="6" pageId="12" pageNumber="13">2</subScript>
proximad of
<emphasis id="30B9EAA1FFC2A927FE6DFE79FDEFFE66" box="[491,516,403,429]" italics="true" pageId="12" pageNumber="13">sb</emphasis>
(
<figureCitation id="9AF62A36FFC2A927FD94FE79FD8DFE66" box="[530,614,403,429]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="12" pageNumber="13">Fig. 9F</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="9AF62A36FFC2A927FDE5FE5DFD56FE1A" box="[611,701,439,465]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="12" pageNumber="13">Fig. 9G</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFC2A927FF41FE31FA77FDAA" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="13">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy IXII: 2: 2: 2: 2: 4: 4: 4: 5: 5: 2: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IIIXII: 1314: 1213: 78: 79: 79: 89: 78: 67: 0: 2. Anterior genital operculum with 9 setae, genital opening slit-like, with 1112 marginal setae on each side,
<quantity id="C5359B56FFC2A927FB2DFDADFB0DFDAA" box="[1195,1254,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="12" pageNumber="13" unit="in" value="32.0">32 in</quantity>
total (
<figureCitation id="9AF62A36FFC2A927FAB5FDADFA7BFDAA" box="[1331,1424,583,609]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="12" pageNumber="13">Fig. 8D</figureCitation>
).
</paragraph>
<caption id="56B2663BFFC2A927FF11FA3FFA76FA26" ID-DOI="http://doi.org/10.5281/zenodo.7251622" ID-Zenodo-Dep="7251622" box="[151,1437,1493,1518]" httpUri="https://zenodo.org/record/7251622/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1493,1517]" targetBox="[151,1417,649,1470]" targetPageId="12">
<paragraph id="027236B3FFC2A927FF11FA3FFA76FA26" blockId="12.[151,1437,1493,1518]" box="[151,1437,1493,1518]" pageId="12" pageNumber="13">
<emphasis id="30B9EAA1FFC2A927FF11FA3FFEF9FA26" bold="true" box="[151,274,1493,1517]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
<taxonomicName id="C5CD4D30FFC2A927FE91FA3FFE16FA26" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[279,509,1493,1517]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC2A927FE91FA3FFE16FA26" box="[279,509,1493,1517]" italics="true" pageId="12" pageNumber="13">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC2A927FD84FA3FFDBAFA26" bold="true" box="[514,593,1493,1517]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="2B8A57DAFFC2A927FD84FA3FFDBAFA26" box="[514,593,1493,1517]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="027236B3FFC2A927FF41F9F6FC2DF8C6" blockId="12.[151,1437,1563,2022]" pageId="12" pageNumber="13">
Legs (
<figureCitation id="9AF62A36FFC2A927FE88F9F1FE6BF9FD" box="[270,384,1563,1590]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[179,1409,181,1850]" captionTargetId="figure-20@14.[179,1409,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius fengi sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="12" pageNumber="13">Fig. 9H, I</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur I, IV and patella IV. Femur of leg I 1.94 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.372.39 times longer than tibia. Femoropatella of leg IV 4.484.60 times longer than deep; tibia 6.206.67 times longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.295.00 times longer than deep (TS= 0.37), telotarsus 14.8015.00 times longer than deep and 2.472.50 times longer than basitarsus (TS= 0.420.44). Setae of leg I (trochanter to tibia) 23: 911: 9: 1213, setae of leg IV (trochanter to basitarsus) 23: 3: 56: 89: 810. Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="027236B3FFC2A927FF41F8FDFEF3F82D" blockId="12.[151,1437,1563,2022]" pageId="12" pageNumber="13">
Adult females (
<typeStatus id="DD768811FFC2A927FEF5F8F2FE09F8F9" box="[371,482,1816,1842]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="9AF62A36FFC2A927FE71F8FDFDA5F8FA" box="[503,590,1815,1841]" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[151,1437,381,1520]" captionTargetId="figure-114@11.[151,1437,381,1520]" captionTargetPageId="11" captionText="FIGURE 6. Unnamed Cave 2, type locality of Lagynochthonius fengi sp. nov., A. Entrance; B. Inside the cave entrance; CD. Areas where L. fengi sp. nov. specimens were collected; EF. Live individuals of L. fengi sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251620" httpUri="https://zenodo.org/record/7251620/files/figure.png" pageId="12" pageNumber="13">Figs 6F</figureCitation>
,
<figureCitation id="9AF62A36FFC2A927FDDEF8F2FD90F8F9" box="[600,635,1816,1842]" captionStart="FIGURE 7" captionStartId="12.[151,250,1493,1517]" captionTargetBox="[151,1417,649,1470]" captionTargetId="figure-252@12.[151,1436,633,1470]" captionTargetPageId="12" captionText="FIGURE 7. Lagynochthonius fengi sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251622" httpUri="https://zenodo.org/record/7251622/files/figure.png" pageId="12" pageNumber="13">7B</figureCitation>
,
<figureCitation id="9AF62A36FFC2A927FD01F8F2FD42F8F9" box="[647,681,1816,1842]" captionStart="FIGURE 8" captionStartId="13.[151,250,1927,1951]" captionTargetBox="[151,1435,181,1903]" captionTargetId="figure-19@13.[151,1436,181,1903]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius fengi sp. nov., holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="12" pageNumber="13">8E</figureCitation>
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 34,
<specimenCode id="526B9EC8FFC2A927FE92F8D6FEADF89D" box="[276,326,1851,1878]" collectionCode="III" pageId="12" pageNumber="13">III 5</specimenCode>
,
<specimenCode id="526B9EC8FFC2A927FED4F8D6FE6DF89D" box="[338,390,1851,1878]" collectionCode="IV" pageId="12" pageNumber="13">IV 5</specimenCode>
; tergal chaetotaxy IXII: 2: 2: 2: 2: 4: 4: 5: 5: 5: 2:
<specimenCode id="526B9EC8FFC2A927FC45F8D6FC08F89D" box="[963,995,1852,1878]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T2</specimenCode>
T: 0; sternal chaetotaxy IVXII: 1213: 8: 89: 78: 8: 7: 67: 0: 2; anterior genital operculum with 9 setae, posterior margin with 1415 marginal setae,
<quantity id="C5359B56FFC2A927FF11F86EFF10F856" box="[151,251,1923,1950]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.096" metricValueMin="5.842" pageId="12" pageNumber="13" unit="in" value="23.5" valueMax="24.0" valueMin="23.0">2324 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: basitarsus 4.004.29 times longer than deep (TS= 0.370.43), telotarsus 15.2015.40 times longer than deep and 2.572.71 times longer than basitarsus (TS= 0.410.44).
</paragraph>
<caption id="56B2663BFFC3A926FF11F86DFAB4F808" ID-DOI="http://doi.org/10.5281/zenodo.7251626" ID-Zenodo-Dep="7251626" httpUri="https://zenodo.org/record/7251626/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1927,1951]" targetBox="[151,1435,181,1903]" targetPageId="13">
<paragraph id="027236B3FFC3A926FF11F86DFAB4F808" blockId="13.[151,1436,1927,1987]" pageId="13" pageNumber="14">
<emphasis id="30B9EAA1FFC3A926FF11F86DFEFAF854" bold="true" box="[151,273,1927,1951]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
<taxonomicName id="C5CD4D30FFC3A926FE90F86DFE17F854" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[278,508,1927,1951]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC3A926FE90F86DFE17F854" box="[278,508,1927,1951]" italics="true" pageId="13" pageNumber="14">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC3A926FD86F86DFDA4F854" bold="true" box="[512,591,1927,1951]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="2B8A57DAFFC3A926FD86F86DFDA4F854" box="[512,591,1927,1951]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (AD), paratype female (E):A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view).
</paragraph>
</caption>
<caption id="56B2663BFFC0A925FF11F8B8FB97F831" ID-DOI="http://doi.org/10.5281/zenodo.7251630" ID-Zenodo-Dep="7251630" httpUri="https://zenodo.org/record/7251630/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1874,1898]" targetBox="[179,1409,181,1850]" targetPageId="14">
<paragraph id="027236B3FFC0A925FF11F8B8FB97F831" blockId="14.[151,1436,1874,2043]" pageId="14" pageNumber="15">
<emphasis id="30B9EAA1FFC0A925FF11F8B8FEF9F8A1" bold="true" box="[151,274,1874,1898]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
<taxonomicName id="C5CD4D30FFC0A925FE9EF8B8FE14F8A1" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[280,511,1874,1898]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC0A925FE9EF8B8FE14F8A1" box="[280,511,1874,1898]" italics="true" pageId="14" pageNumber="15">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC0A925FD83F8B8FDBEF8A1" bold="true" box="[517,597,1874,1898]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="2B8A57DAFFC0A925FD83F8B8FDBEF8A1" box="[517,597,1874,1898]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD).
</paragraph>
</caption>
<paragraph id="027236B3FFC1A924FF41FF7DFA7EFE66" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.072.10. Pedipalps: trochanter 0.260.27/0.150.16 (1.691.73), femur 1.061.07/0.14 (7.577.64), patella 0.370.39/0.160.17 (2.182.44), chela 1.531.54/0.210.22 (7.007.29), hand 0.630.64/0.210.22 (2.863.05), movable chelal finger length 0.880.91. Chelicera 0.56/0.230.24 (2.332.43), movable finger length 0.300.31. Carapace 0.53/0.520.53 (1.001.02). Leg I: trochanter 0.170.18/0.120.13 (1.311.50), femur 0.600.62/0.08 (7.507.75), patella 0.310.32/0.07 (4.434.57), tibia 0.270.28/0.06 (4.504.67), tarsus 0.640.67/0.050.06 (11.1712.80). Leg IV: trochanter 0.230.24/0.14 (1.641.71), femoropatella 0.920.94/0.200.21 (4.484.60), tibia 0.600.62/0.090.10 (6.206.67), basitarsus 0.30/0.060.07 (4.295.00), telotarsus 0.740.75/0.05 (14.8015.00).</paragraph>
<paragraph id="027236B3FFC1A924FF41FE5DFABDFD62" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">Females: body length 2.252.32. Pedipalps: trochanter 0.260.29/0.150.16 (1.731.81), femur 1.081.15/0.14 (7.718.21), patella 0.390.41/0.17 (2.292.41), chela 1.531.55/0.22 (6.957.05), hand 0.64/0.22 (2.91), movable chelal finger length 0.890.90. Chelicera 0.570.61/0.250.27 (2.262.28), movable finger length 0.310.32. Carapace 0.560.58/0.550.60 (0.971.02). Leg I: trochanter 0.170.18/0.13 (1.311.38), femur 0.600.64/0.08 (7.508.00), patella 0.310.32/0.07 (4.434.57), tibia 0.290.31/0.05 (5.806.20), tarsus 0.620.66/0.06 (10.33 11.00). Leg IV: trochanter 0.250.27/0.15 (1.671.80), femoropatella 0.910.95/0.200.22 (4.324.55), tibia 0.61 0.65/0.090.10 (6.506.78), basitarsus 0.280.30/0.07 (4.004.29), telotarsus 0.760.77/0.05 (15.2015.40).</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFC1A924FF41FD59FB36FBDA" pageId="15" pageNumber="16" type="discussion">
<paragraph id="027236B3FFC1A924FF41FD59FE5AFCF2" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">
<emphasis id="30B9EAA1FFC1A924FF41FD59FED0FD06" bold="true" box="[199,315,691,717]" pageId="15" pageNumber="16">Remarks.</emphasis>
<taxonomicName id="C5CD4D30FFC1A924FEC2FD59FDAEFD06" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[324,581,691,717]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC1A924FEC2FD59FDAEFD06" box="[324,581,691,717]" italics="true" pageId="15" pageNumber="16">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC1A924FDC8FD59FD42FD06" bold="true" box="[590,681,691,717]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="2B8A57DAFFC1A924FDC8FD59FD42FD06" box="[590,681,691,717]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="C5CD4D30FFC1A924FCC4FD59FBF1FD07" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[834,1050,691,716]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFC1A924FCC4FD59FBF1FD07" box="[834,1050,691,716]" italics="true" pageId="15" pageNumber="16">L. yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC1A924FBA5FD59FB95FD06" bold="true" box="[1059,1150,691,717]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="2B8A57DAFFC1A924FBA5FD59FB95FD06" box="[1059,1150,691,717]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having the intercalary teeth on both chelal fingers and a pointed and triangular epistome, but differs by the shorter chela (e.g. chela length 1.531.54
<emphasis id="30B9EAA1FFC1A924FE88FD11FEC6FCDF" box="[270,301,763,788]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
<quantity id="C5359B56FFC1A924FEB2FD11FE71FCDE" box="[308,410,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="15" pageNumber="16" unit="mm" value="1.84">1.84 mm</quantity>
, movable chelal finger length 0.880.91
<emphasis id="30B9EAA1FFC1A924FCE0FD11FC6FFCDF" box="[870,900,763,788]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
<quantity id="C5359B56FFC1A924FC0DFD10FC18FCDE" box="[907,1011,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="16" unit="mm" value="1.15">1.15 mm</quantity>
; all in males) and the number of setae on tergites IIV (2
<emphasis id="30B9EAA1FFC1A924FEEBFCF5FE67FCF3" box="[365,396,799,824]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
4).
</paragraph>
<paragraph id="027236B3FFC1A924FF41FCA9FB2EFC6E" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">
<taxonomicName id="C5CD4D30FFC1A924FF41FCA9FE20FC96" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,459,835,861]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC1A924FF41FCA9FE20FC96" box="[199,459,835,861]" italics="true" pageId="15" pageNumber="16">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC1A924FE50FCA9FDDFFC96" bold="true" box="[470,564,835,861]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="2B8A57DAFFC1A924FE50FCA9FDDFFC96" box="[470,564,835,861]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="C5CD4D30FFC1A924FD52FCA9FC4CFC96" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[724,935,835,861]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bailongtanensis">
<emphasis id="30B9EAA1FFC1A924FD52FCA9FC4CFC96" box="[724,935,835,861]" italics="true" pageId="15" pageNumber="16">L. bailongtanensis</emphasis>
</taxonomicName>
in the shape of the epistome (pointed and triangular
<emphasis id="30B9EAA1FFC1A924FE88FC8DFECEFC4B" box="[270,293,871,896]" italics="true" pageId="15" pageNumber="16">vs</emphasis>
. rounded and obtuse), the pattern of dentation on the chelal fingers (with intercalary teeth on both chelal fingers
<emphasis id="30B9EAA1FFC1A924FEB0FC61FEA6FC6F" box="[310,333,907,932]" italics="true" pageId="15" pageNumber="16">vs</emphasis>
. without intercalary teeth), and the number of setae on tergites IIV (2
<emphasis id="30B9EAA1FFC1A924FBEBFC61FB60FC6F" box="[1133,1163,907,932]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
34)
</paragraph>
<paragraph id="027236B3FFC1A924FF41FC45FB36FBDA" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">
<taxonomicName id="C5CD4D30FFC1A924FF41FC45FE2DFC02" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,454,943,969]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFC1A924FF41FC45FE2DFC02" box="[199,454,943,969]" italics="true" pageId="15" pageNumber="16">Lagynochthonius fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFC1A924FE4DFC45FDCFFC02" bold="true" box="[459,548,943,969]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="2B8A57DAFFC1A924FE4DFC45FDCFFC02" box="[459,548,943,969]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="C5CD4D30FFC1A924FCCBFC45FBFFFC02" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[845,1044,943,969]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="laoxueyanensis">
<emphasis id="30B9EAA1FFC1A924FCCBFC45FBFFFC02" box="[845,1044,943,969]" italics="true" pageId="15" pageNumber="16">L. laoxueyanensis</emphasis>
</taxonomicName>
by the presence of intercalary teeth on both chelal fingers, the number of setae on tergites III and IV (2 and 2, respectively
<emphasis id="30B9EAA1FFC1A924FBD2FC39FB98FC27" box="[1108,1139,979,1004]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
4 and 4, respectively), and a larger body size (body length 2.072.10
<emphasis id="30B9EAA1FFC1A924FDF6FC1DFD64FBDB" box="[624,655,1015,1040]" italics="true" pageId="15" pageNumber="16">vs.</emphasis>
<quantity id="C5359B56FFC1A924FD10FC1DFD11FBDA" box="[662,762,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="15" pageNumber="16" unit="mm" value="1.78">1.78 mm</quantity>
in males) (
<bibRefCitation id="665C4B42FFC1A924FCF0FC1DFBF9FBDA" author="Li, Y. C. &amp; Liu, H. &amp; Shi, A. M." box="[886,1042,1015,1041]" pageId="15" pageNumber="16" pagination="28 - 34" refId="ref39131" refString="Li, Y. C., Liu, H. &amp; Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">
Li
<emphasis id="30B9EAA1FFC1A924FC1EFC1DFC22FBDA" box="[920,969,1015,1041]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2019
</bibRefCitation>
;
<bibRefCitation id="665C4B42FFC1A924FB98FC1DFB3AFBDA" author="Hou, Y. M. &amp; Gao, Z. Z. &amp; Zhang, F." box="[1054,1233,1015,1041]" pageId="15" pageNumber="16" pagination="65 - 83" refId="ref38684" refString="Hou, Y. M., Gao, Z. Z. &amp; Zhang, F. (2022) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys, 1097, 65 - 83. https: // doi. org / 10.3897 / zookeys. 1097.82527" type="journal article" year="2022">
Hou
<emphasis id="30B9EAA1FFC1A924FBD3FC1DFB6DFBDA" box="[1109,1158,1015,1041]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFC1A924FF41FBF0FC9CFB22" pageId="15" pageNumber="16" type="distribution">
<paragraph id="027236B3FFC1A924FF41FBF0FC9CFB22" blockId="15.[151,1437,151,1257]" pageId="15" pageNumber="16">
<emphasis id="30B9EAA1FFC1A924FF41FBF0FE19FBFF" bold="true" box="[199,498,1050,1077]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
This species is only known from the
<typeStatus id="DD768811FFC1A924FC1FFBF1FC22FBFE" box="[921,969,1051,1077]" pageId="15" pageNumber="16">type</typeStatus>
locality, Unnamed Cave 2 (
<figureCitation id="9AF62A36FFC1A924FA82FBF1FAB4FBFE" box="[1284,1375,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); MN. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="15" pageNumber="16">Figs 1B</figureCitation>
,
<figureCitation id="9AF62A36FFC1A924FAEAFBF1FF45FB92" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[151,1437,381,1520]" captionTargetId="figure-114@11.[151,1437,381,1520]" captionTargetPageId="11" captionText="FIGURE 6. Unnamed Cave 2, type locality of Lagynochthonius fengi sp. nov., A. Entrance; B. Inside the cave entrance; CD. Areas where L. fengi sp. nov. specimens were collected; EF. Live individuals of L. fengi sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251620" httpUri="https://zenodo.org/record/7251620/files/figure.png" pageId="15" pageNumber="16">6A D</figureCitation>
), which is located about
<quantity id="C5359B56FFC1A924FE47FBD5FDE4FB92" box="[449,527,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="km" value="0.6">0.6 km</quantity>
west of Maliu Village (Yanjin County) near the top of a mountain. This limestone cave has a small and slit-shaped entrance (about
<quantity id="C5359B56FFC1A924FD3AFB89FD0CFBB6" box="[700,743,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1.0">1 m</quantity>
high and
<quantity id="C5359B56FFC1A924FCD1FB89FC69FBB6" box="[855,898,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="16" unit="m" value="3.0">3 m</quantity>
wide) and only a narrow tunnel (approximately
<quantity id="C5359B56FFC1A924FF11FB6DFF3AFB6A" box="[151,209,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="80.0">80 m</quantity>
in length) leading to the deepest part of the cave, which is a slightly wider, low-temperature, high-humidity and completely lightless environment (temperature around 11°C, humidity over 90%). All of the specimens were collected under stones in the deepest part of the cave (deep zone).
</paragraph>
</subSubSection>
</treatment>
</document>