309 lines
37 KiB
XML
309 lines
37 KiB
XML
<document id="DE8AAEBF35C7F5EF8A48FA180961906C" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan & Kompier, Tom" docDate="2016" docId="8A1EB74F897EFF95C7F363AEFD299477" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta beaumonti Wilson 1997" docType="treatment" docVersion="7" lastPageNumber="531" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="530" updateTime="1698233463814" updateUser="plazi">
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<mods:titleInfo id="BC1902EEA56923E2939A1CD2BD5813F8">
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<mods:title id="4C83C9988A0BB32E6E6648A2BD004262">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="BD7D4636321C68F40FBBE609DFE77C92">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="2A8F8A01FAF64A30311DA343775D875B">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="7646FC449FEF982252434D9A5C43313C">text</mods:typeOfResource>
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<mods:title id="63230686BBFEAF4B0D0AC0646CAF5A5A">Zootaxa</mods:title>
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<mods:part id="52B7EEB4590821A5E62561173924BBF5">
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<mods:date id="701F5262EA1277608AC04F716CE40D3D">2016</mods:date>
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<mods:number id="24F50762AC3A86A1D8BDC99366177159">4098</mods:number>
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<treatment id="8A1EB74F897EFF95C7F363AEFD299477" ID-DOI="http://doi.org/10.5281/zenodo.6056371" ID-GBIF-Taxon="127620790" ID-Zenodo-Dep="6056371" LSID="urn:lsid:plazi:treatment:8A1EB74F897EFF95C7F363AEFD299477" httpUri="http://treatment.plazi.org/id/8A1EB74F897EFF95C7F363AEFD299477" lastPageId="2" lastPageNumber="531" pageId="1" pageNumber="530">
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<subSubSection id="4AAD55D2897EFF96C7F363AEFC9B9088" box="[151,770,473,499]" pageId="1" pageNumber="530" type="nomenclature">
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<paragraph id="02080659897EFF96C7F363AEFC9B9088" blockId="1.[151,770,473,533]" box="[151,770,473,499]" pageId="1" pageNumber="530">
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<heading id="5940B135897EFF96C7F363AEFC9B9088" bold="true" box="[151,770,473,499]" fontSize="11" level="1" pageId="1" pageNumber="530" reason="1">
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<emphasis id="30C3DA4B897EFF96C7F363AEFC9B9088" bold="true" box="[151,770,473,499]" pageId="1" pageNumber="530">
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1.
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<taxonomicName id="C5B77DDA897EFF96C7D063AEFDFF9088" ID-CoL="4N77Y" authority="Wilson, 1997" authorityName="Wilson" authorityYear="1997" box="[180,614,473,499]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="530" phylum="Arthropoda" rank="species" species="beaumonti">
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||
<emphasis id="30C3DA4B897EFF96C7D063AEFE279088" bold="true" box="[180,446,473,499]" italics="true" pageId="1" pageNumber="530">Protosticta beaumonti</emphasis>
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<bibRefCitation id="66267BA8897EFF96C6A163AEFDFF9088" author="Wilson" box="[453,614,473,499]" pageId="1" pageNumber="530" refString="Wilson, K. D. P. (1997) The Platystictidae of Hong Kong and Guangdong with descriptions of a new genus and two new species (Zygoptera). Odonatologica, 26 (1), 53 - 63." type="journal article" year="1997">Wilson, 1997</bibRefCitation>
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</taxonomicName>
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||
(dark form)
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="4AAD55D2897EFF96C7F3638BFBBA9321" pageId="1" pageNumber="530" type="description">
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<paragraph id="02080659897EFF96C7F3638BFEA0936F" blockId="1.[151,770,473,533]" box="[151,313,508,533]" pageId="1" pageNumber="530">
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(
|
||
<figureCitation id="9A8C1ADC897EFF96C7FA638BFF60936F" box="[158,249,508,533]" captionStart="FIGURE 1" captionStartId="1.[151,250,1915,1937]" captionTargetBox="[157,1432,706,1896]" captionTargetId="figure@1.[151,1435,701,1909]" captionTargetPageId="1" captionText="FIGURE 1. Protosticta beaumonti, Mau Son. [A – D] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus & paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E – F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." httpUri="https://zenodo.org/record/257724/files/figure.png" pageId="1" pageNumber="530">Figure 1</figureCitation>
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A–F)
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</paragraph>
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<paragraph id="02080659897EFF96C7F36032FBBA9321" blockId="1.[151,1436,580,663]" box="[151,1059,580,602]" pageId="1" pageNumber="530">
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||
<taxonomicName id="C5B77DDA897EFF96C7F36032FE9E9321" box="[151,263,581,602]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="30C3DA4B897EFF96C7F36032FE9E9321" box="[151,263,581,602]" italics="true" pageId="1" pageNumber="530">Protosticta</emphasis>
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||
</taxonomicName>
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sp.:
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<bibRefCitation id="66267BA8897EFF96C6536033FE529321" author="Wilson" box="[311,459,580,602]" pageId="1" pageNumber="530" refString="Wilson, K. D. P. (1995) Hong Kong dragonflies. Urban Council of Hong Kong, Hong Kong, 211 pp." type="book" year="1995">Wilson (1995)</bibRefCitation>
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, p. 21, p. 23, Figs. 78, 79, 81, 83, 85 (photos,
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<collectingCountry id="7AA046C9897EFF96C4F86032FB819321" box="[924,1048,581,602]" name="Hong Kong" pageId="1" pageNumber="530">Hong Kong</collectingCountry>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4AAD55D2897EFF95C7F36014FD9F95DB" lastPageId="2" lastPageNumber="531" pageId="1" pageNumber="530" type="reference_group">
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||
<paragraph id="02080659897EFF96C7F36014FB3E93EC" blockId="1.[151,1436,580,663]" pageId="1" pageNumber="530">
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||
<taxonomicName id="C5B77DDA897EFF96C7F36014FD879303" ID-CoL="4N77Y" authority="Wilson (1997)" authorityName="Wilson" authorityYear="1997" box="[151,542,610,632]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="530" phylum="Arthropoda" rank="species" species="beaumonti">
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<emphasis id="30C3DA4B897EFF96C7F36014FEE09303" box="[151,377,610,632]" italics="true" pageId="1" pageNumber="530">Protosticta beaumonti</emphasis>
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||
:
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<bibRefCitation id="66267BA8897EFF96C6EC6015FD879303" author="Wilson" box="[392,542,610,632]" pageId="1" pageNumber="530" refString="Wilson, K. D. P. (1997) The Platystictidae of Hong Kong and Guangdong with descriptions of a new genus and two new species (Zygoptera). Odonatologica, 26 (1), 53 - 63." type="journal article" year="1997">Wilson (1997)</bibRefCitation>
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||
</taxonomicName>
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||
, pp. 57–59, Figs. 11–16.—
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||
<bibRefCitation id="66267BA8897EFF96C4556015FBBE9303" author="Wilson" box="[817,1063,610,632]" pageId="1" pageNumber="530" refString="Wilson, K. D. P. & Reels G. T. (2003) Odonata of Guangxi Zhuang autonomous region, China, part I: Zygoptera. Odonatologica, 32 (3), 237 - 279." type="journal article" year="2003">Wilson & Reels (2003)</bibRefCitation>
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||
, pp. 273–275, Figs. 114–120 (
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||
<collectingCountry id="7AA046C9897EFF96C2066015FE9B93EC" name="Hong Kong" pageId="1" pageNumber="530">Hong Kong</collectingCountry>
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||
, Guangdong).—
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<bibRefCitation id="66267BA8897EFF96C6CC60F6FD0993EC" box="[424,656,641,663]" pageId="1" pageNumber="530" refString="Kitagawa, K. (1997) Records of the Odonata of Hong Kong, Part IV. Aeschna, 34, 11 - 21." type="journal article">
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||
Kitagawa
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<emphasis id="30C3DA4B897EFF96C57560F5FDDD93EC" box="[529,580,641,663]" italics="true" pageId="1" pageNumber="530">et al.</emphasis>
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||
(1997)
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||
</bibRefCitation>
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||
, p. 12,
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||
<figureCitation id="9A8C1ADC897EFF96C5BE60F6FCA193EC" box="[730,824,641,663]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1687,1709]" captionStartId-1="4.[151,250,1912,1934]" captionTargetBox-0="[159,1432,476,1674]" captionTargetBox-1="[169,1425,470,1883]" captionTargetId-0="figure@3.[151,1437,469,1675]" captionTargetId-1="figure@4.[150,1436,465,1891]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." captionText-1="FIGURE 3. Protosticta grandis, Cao Bang. [A – E] male— (A) head & thorax in lateral view; anal appendages in (B) dorsal & (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view & (F) lateral view; [G – H] female— (G) head & thorax in lateral view; (H) abdomen tip in lateral view." httpUri-0="https://zenodo.org/record/257725/files/figure.png" httpUri-1="https://zenodo.org/record/257726/files/figure.png" pageId="1" pageNumber="530">Figs. 2–3</figureCitation>
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||
(Sunset Peak, Lantau,
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||
<collectingCountry id="7AA046C9897EFF96C34560F5FB0493EC" box="[1057,1181,642,663]" name="Hong Kong" pageId="1" pageNumber="530">Hong Kong</collectingCountry>
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||
).
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||
</paragraph>
|
||
<caption id="56C856D1897EFF96C7F3650CFDAC9696" httpUri="https://zenodo.org/record/257724/files/figure.png" pageId="1" pageNumber="530" targetBox="[157,1432,706,1896]" targetPageId="1">
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||
<paragraph id="02080659897EFF96C7F3650CFDAC9696" blockId="1.[151,1436,1915,2029]" pageId="1" pageNumber="530">
|
||
<emphasis id="30C3DA4B897EFF96C7F3650CFE9796EB" bold="true" box="[151,270,1915,1937]" pageId="1" pageNumber="530">FIGURE 1</emphasis>
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||
.
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||
<taxonomicName id="C5B77DDA897EFF96C67F650BFDF696EA" ID-CoL="4N77Y" authority="Mau Son." authorityName="Mau Son." box="[283,623,1915,1937]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="530" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897EFF96C67F650BFD9A96EA" box="[283,515,1915,1937]" italics="true" pageId="1" pageNumber="530">Protosticta beaumonti,</emphasis>
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||
Mau Son.
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||
</taxonomicName>
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||
[A–D] male—(
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<emphasis id="30C3DA4B897EFF96C476650CFCBD96EB" bold="true" box="[786,804,1915,1936]" pageId="1" pageNumber="530">A</emphasis>
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) head & thorax in lateral view; (
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<emphasis id="30C3DA4B897EFF96C31D650CFB1396EB" bold="true" box="[1145,1162,1915,1936]" pageId="1" pageNumber="530">B</emphasis>
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) prothorax in dorsal view; (
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||
<emphasis id="30C3DA4B897EFF96C7FB65EDFF2896CB" bold="true" box="[159,177,1946,1968]" pageId="1" pageNumber="530">C</emphasis>
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||
) right cercus & paraproct in oblique-dorsal view; (
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||
<emphasis id="30C3DA4B897EFF96C5DF65EDFD5496D4" bold="true" box="[699,717,1946,1967]" pageId="1" pageNumber="530">D</emphasis>
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||
) penile organ in dorsal view; [E–F] female—(
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<emphasis id="30C3DA4B897EFF96C3CC65EDFB2096D4" bold="true" box="[1192,1209,1946,1967]" pageId="1" pageNumber="530">E</emphasis>
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||
) female in nature; (
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||
<emphasis id="30C3DA4B897EFF96C2E065EDFA0A96D4" bold="true" box="[1412,1427,1946,1967]" pageId="1" pageNumber="530">F</emphasis>
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) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07.VI.2014 by Tom Kompier, specimens were not collected).
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||
</paragraph>
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||
</caption>
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||
<paragraph id="02080659897DFF95C7F362E0FAA491AE" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
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||
<emphasis id="30C3DA4B897DFF95C7F362E0FE0D91CB" bold="true" box="[151,404,151,176]" pageId="2" pageNumber="531">Specimens examined</emphasis>
|
||
.
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||
<specimenCount id="14B1CDD0897DFF95C6CC62EFFE6291CB" box="[424,507,151,176]" pageId="2" pageNumber="531" type="male">1 male</specimenCount>
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||
, Mau Son Mountain, Loc Binh district, Lang Son Province,
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||
<date id="76092099897DFF95C3B262EFFAC191CB" box="[1238,1368,152,176]" pageId="2" pageNumber="531" value="2014-06-07">07.VI.2014</date>
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||
, Tom Kompier leg.;
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||
<specimenCount id="14B1CDD0897DFF95C65D62CBFE0B91AE" box="[313,402,188,213]" pageId="2" pageNumber="531" type="male">3 males</specimenCount>
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||
, Bhalee, Tay Giang district, Quang Nam Province,
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<date id="76092099897DFF95C4B662CBFBDE91AE" box="[978,1095,188,213]" pageId="2" pageNumber="531" value="2015-05-27">27.V.2015</date>
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, Quoc Toan Phan leg.
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||
</paragraph>
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||
<paragraph id="02080659897DFF95C7A362A8FAB990F3" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
|
||
<emphasis id="30C3DA4B897DFF95C7A362A8FE939183" bold="true" box="[199,266,223,248]" pageId="2" pageNumber="531">Notes</emphasis>
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||
. These are the first records of this species for
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<collectingCountry id="7AA046C9897DFF95C44F62A8FC179183" box="[811,910,223,248]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
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||
.
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||
<bibRefCitation id="66267BA8897DFF95C4FE62A8FBA79183" author="Wilson" box="[922,1086,223,248]" pageId="2" pageNumber="531" refString="Wilson, K. D. P. (1997) The Platystictidae of Hong Kong and Guangdong with descriptions of a new genus and two new species (Zygoptera). Odonatologica, 26 (1), 53 - 63." type="journal article" year="1997">Wilson (1997)</bibRefCitation>
|
||
described the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C2E36296FE949066" ID-CoL="4N77Y" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C2E36296FE949066" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
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||
from
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||
<collectingCountry id="7AA046C9897DFF95C6346372FE4F9066" box="[336,470,261,285]" name="Hong Kong" pageId="2" pageNumber="531">Hong Kong</collectingCountry>
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||
, and later described the dark form of this species from Guangdong in
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<collectingCountry id="7AA046C9897DFF95C3B96373FAB99066" box="[1245,1312,260,285]" name="China" pageId="2" pageNumber="531">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="66267BA8897DFF95C24A6373FE80903B" author="Wilson" pageId="2" pageNumber="531" refString="Wilson, K. D. P. & Reels G. T. (2003) Odonata of Guangxi Zhuang autonomous region, China, part I: Zygoptera. Odonatologica, 32 (3), 237 - 279." type="journal article" year="2003">Wilson & Reels 2003</bibRefCitation>
|
||
). The male specimens from northern and central
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<collectingCountry id="7AA046C9897DFF95C45E6350FC00903B" box="[826,921,295,320]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
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||
are consistent with the description of the dark form of
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||
<taxonomicName id="C5B77DDA897DFF95C792633AFD7A901F" ID-CoL="4N77Y" authority="Wilson & Reels (2003)" authorityName="Wilson & Reels" authorityYear="2003" box="[246,739,332,357]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
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||
<emphasis id="30C3DA4B897DFF95C792633AFE10901E" box="[246,393,332,357]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
sensu
|
||
<bibRefCitation id="66267BA8897DFF95C6BD633BFD7A901F" author="Wilson" box="[473,739,332,357]" pageId="2" pageNumber="531" refString="Wilson, K. D. P. & Reels G. T. (2003) Odonata of Guangxi Zhuang autonomous region, China, part I: Zygoptera. Odonatologica, 32 (3), 237 - 279." type="journal article" year="2003">Wilson & Reels (2003)</bibRefCitation>
|
||
</taxonomicName>
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||
, including the color pattern of the abdomen of the female.
|
||
<taxonomicName id="C5B77DDA897DFF95C2E3633AFE9490F3" ID-CoL="4N77Y" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C2E3633AFE9490F3" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="C5B77DDA897DFF95C6EF6306FE4590F3" ID-CoL="4N794" box="[395,476,369,392]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C6EF6306FE4590F3" box="[395,476,369,392]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
closely, but their dark forms can be separated by the following characters:
|
||
</paragraph>
|
||
<paragraph id="02080659897DFF95C7A363E2FCAD908F" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
|
||
In
|
||
<emphasis id="30C3DA4B897DFF95C78263E2FEAC90D7" box="[230,309,405,428]" italics="true" pageId="2" pageNumber="531">P. s a t o i</emphasis>
|
||
, the yellow marking between the posterior end of the metepisternum and the metepimeron consists of a very small stripe in the male (
|
||
<figureCitation id="9A8C1ADC897DFF95C57E63C0FDC590AB" box="[538,604,439,464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="2" pageNumber="531">Fig. 5</figureCitation>
|
||
A) and is absent in the female (
|
||
<figureCitation id="9A8C1ADC897DFF95C4DB63C0FB9890AB" box="[959,1025,439,464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="2" pageNumber="531">Fig. 5</figureCitation>
|
||
H) but consists of a very large stripe both in the male and female of
|
||
<taxonomicName id="C5B77DDA897DFF95C69063AAFD1C908E" ID-CoL="4N77Y" box="[500,645,476,501]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C69063AAFD1C908E" box="[500,645,476,501]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9A8C1ADC897DFF95C5F163ABFD47908F" box="[661,734,476,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1915,1937]" captionTargetBox="[157,1432,706,1896]" captionTargetId="figure@1.[151,1435,701,1909]" captionTargetPageId="1" captionText="FIGURE 1. Protosticta beaumonti, Mau Son. [A – D] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus & paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E – F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." httpUri="https://zenodo.org/record/257724/files/figure.png" pageId="2" pageNumber="531">Figs. 1</figureCitation>
|
||
A, 1E).
|
||
</paragraph>
|
||
<paragraph id="02080659897DFF95C7A36388FF599346" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
|
||
The dorsum of S9 of the female of
|
||
<taxonomicName id="C5B77DDA897DFF95C5366076FD7D9363" ID-CoL="4N77Y" box="[594,740,511,536]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C5366076FD7D9363" box="[594,740,511,536]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
is dark (
|
||
<figureCitation id="9A8C1ADC897DFF95C4236388FC1E9363" box="[839,903,511,536]" captionStart="FIGURE 1" captionStartId="1.[151,250,1915,1937]" captionTargetBox="[157,1432,706,1896]" captionTargetId="figure@1.[151,1435,701,1909]" captionTargetPageId="1" captionText="FIGURE 1. Protosticta beaumonti, Mau Son. [A – D] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus & paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E – F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." httpUri="https://zenodo.org/record/257724/files/figure.png" pageId="2" pageNumber="531">Fig. 1</figureCitation>
|
||
F), not whitish as in the female of
|
||
<taxonomicName id="C5B77DDA897DFF95C2696076FAC69363" ID-CoL="4N794" box="[1293,1375,513,536]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C2696076FAC69363" box="[1293,1375,513,536]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9A8C1ADC897DFF95C20A6388FF3F9346" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="2" pageNumber="531">Fig. 5</figureCitation>
|
||
J).
|
||
</paragraph>
|
||
<paragraph id="02080659897DFF95C7A36030FD9F95DB" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
|
||
The relationships among the light and dark forms of
|
||
<taxonomicName id="C5B77DDA897DFF95C471603EFCFF931B" ID-CoL="4N794" box="[789,870,585,608]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C471603EFCFF931B" box="[789,870,585,608]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C5B77DDA897DFF95C4F9603EFBB6931B" ID-CoL="4N77Y" box="[925,1071,583,608]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C4F9603EFBB6931B" box="[925,1071,583,608]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
are problematic. Van
|
||
<bibRefCitation id="66267BA8897DFF95C2416030FA05931B" author="Tol" box="[1317,1436,583,608]" pageId="2" pageNumber="531" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
|
||
made a convincing case that the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C5F2601AFD7193FF" ID-CoL="4N794" box="[662,744,621,644]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C5F2601AFD7193FF" box="[662,744,621,644]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
and the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C4B9601AFBB693FF" ID-CoL="4N794" box="[989,1071,621,644]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C4B9601AFBB693FF" box="[989,1071,621,644]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
are the same species. We follow this interpretation. Van Tol also suggested that
|
||
<emphasis id="30C3DA4B897DFF95C5AD60E6FC8493D3" box="[713,797,657,680]" italics="true" pageId="2" pageNumber="531">P. s a t o i</emphasis>
|
||
may be the same species as light-form
|
||
<taxonomicName id="C5B77DDA897DFF95C26160E6FA0593D3" ID-CoL="4N77Y" box="[1285,1436,655,680]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C26160E6FA0593D3" box="[1285,1436,655,680]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
described from
|
||
<collectingCountry id="7AA046C9897DFF95C62B60C2FE4E93B6" box="[335,471,693,717]" name="Hong Kong" pageId="2" pageNumber="531">Hong Kong</collectingCountry>
|
||
by
|
||
<bibRefCitation id="66267BA8897DFF95C56E60C3FD2A93B7" author="Wilson" box="[522,691,692,717]" pageId="2" pageNumber="531" refString="Wilson, K. D. P. (1997) The Platystictidae of Hong Kong and Guangdong with descriptions of a new genus and two new species (Zygoptera). Odonatologica, 26 (1), 53 - 63." type="journal article" year="1997">Wilson (1997)</bibRefCitation>
|
||
. At the same time we consider the dark form of
|
||
<emphasis id="30C3DA4B897DFF95C39A60C2FAC893B7" box="[1278,1361,693,716]" italics="true" pageId="2" pageNumber="531">P. s a t o i</emphasis>
|
||
to be different from the dark form specimens from Mau Son and Quang Nam based on coloration and minor morphological differences and consider these to be conspecific with the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C306608AFB6C926E" ID-CoL="4N77Y" box="[1122,1269,764,789]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C306608AFB6C926E" box="[1122,1269,764,789]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
from Guangxi (
|
||
<bibRefCitation id="66267BA8897DFF95C7FA6168FE339243" author="Wilson" box="[158,426,799,824]" pageId="2" pageNumber="531" refString="Wilson, K. D. P. & Reels G. T. (2003) Odonata of Guangxi Zhuang autonomous region, China, part I: Zygoptera. Odonatologica, 32 (3), 237 - 279." type="journal article" year="2003">Wilson and Reels 2003</bibRefCitation>
|
||
). Van Tol put forward the possibility that the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C35B6156FB179243" ID-CoL="4N794" box="[1087,1166,801,824]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C35B6156FB179243" box="[1087,1166,801,824]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
may be the same taxon as the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C6E56132FD8F9226" ID-CoL="4N77Y" box="[385,534,836,861]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C6E56132FD8F9226" box="[385,534,836,861]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
from Guangxi. We do not follow that interpretation here. However, from this follows that either the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C554611EFD5A92FB" ID-CoL="4N77Y" box="[560,707,871,896]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C554611EFD5A92FB" box="[560,707,871,896]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
is not the same species as the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C3D8611EFAD792FB" ID-CoL="4N77Y" box="[1212,1358,871,896]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C3D8611EFAD792FB" box="[1212,1358,871,896]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
(if the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C64A61FAFE2692DE" ID-CoL="4N77Y" box="[302,447,908,933]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C64A61FAFE2692DE" box="[302,447,908,933]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="C5B77DDA897DFF95C5AF61FAFC8492DF" ID-CoL="4N794" box="[715,797,909,932]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C5AF61FAFC8492DF" box="[715,797,909,932]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
) or that the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C35A61FAFB4992DE" ID-CoL="4N77Y" box="[1086,1232,908,933]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C35A61FAFB4992DE" box="[1086,1232,908,933]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
is not conspecific with
|
||
<taxonomicName id="C5B77DDA897DFF95C7B461C6FE8692B3" ID-CoL="4N794" box="[208,287,945,968]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="30C3DA4B897DFF95C7B461C6FE8692B3" box="[208,287,945,968]" italics="true" pageId="2" pageNumber="531">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
, in which case the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C5EB61C6FCB892B3" ID-CoL="4N77Y" box="[655,801,943,968]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C5EB61C6FCB892B3" box="[655,801,943,968]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
may or may not be the same species as the light form of
|
||
<taxonomicName id="C5B77DDA897DFF95C7F361A2FEB09296" ID-CoL="4N77Y" box="[151,297,980,1005]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C7F361A2FEB09296" box="[151,297,980,1005]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
. For the moment, we follow
|
||
<bibRefCitation id="66267BA8897DFF95C51561A3FCEC9297" author="Wilson" box="[625,885,980,1005]" pageId="2" pageNumber="531" refString="Wilson, K. D. P. & Reels G. T. (2003) Odonata of Guangxi Zhuang autonomous region, China, part I: Zygoptera. Odonatologica, 32 (3), 237 - 279." type="journal article" year="2003">Wilson & Reels (2003)</bibRefCitation>
|
||
that the light and dark forms of
|
||
<taxonomicName id="C5B77DDA897DFF95C38561A2FAEB9296" ID-CoL="4N77Y" box="[1249,1394,980,1005]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C38561A2FAEB9296" box="[1249,1394,980,1005]" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
are conspecific, as we have not studied the material. For the same reason of not having studied the Chinese specimens, it remains possible that the specimens from Mau Son and Quang Nam here identified as the dark form of
|
||
<taxonomicName id="C5B77DDA897DFF95C2E3666AFE949523" ID-CoL="4N77Y" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="30C3DA4B897DFF95C2E3666AFE949523" italics="true" pageId="2" pageNumber="531">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
are in fact an additional and as yet undescribed species. Detailed molecular studies are required to verify the total number of species involved and their relationships. The occurrence of at least two species of this complex in
|
||
<collectingCountry id="7AA046C9897DFF95C7D066F0FE8A95DB" box="[180,275,1159,1184]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
|
||
seems beyond doubt.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AAD55D2897DFF95C7A366DBFBD495BF" box="[199,1101,1196,1221]" pageId="2" pageNumber="531" type="description">
|
||
<paragraph id="02080659897DFF95C7A366DBFBD495BF" blockId="2.[151,1436,151,1293]" box="[199,1101,1196,1221]" pageId="2" pageNumber="531">
|
||
<emphasis id="30C3DA4B897DFF95C7A366DBFEEF95BE" bold="true" box="[199,374,1196,1221]" pageId="2" pageNumber="531">Measurements</emphasis>
|
||
. Male. Hindwing 24–25; abdomen including appendages 43–44.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AAD55D2897DFF95C7A366B8FD299477" pageId="2" pageNumber="531" type="distribution">
|
||
<paragraph id="02080659897DFF95C7A366B8FD299477" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="531">
|
||
<emphasis id="30C3DA4B897DFF95C7A366B8FEC09593" bold="true" box="[199,345,1231,1256]" pageId="2" pageNumber="531">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="7AA046C9897DFF95C60E66B8FE509593" box="[362,457,1231,1256]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
|
||
(Mau Son mountain, Lang Son Province; Tay Giang district, Quang Nam Province),
|
||
<collectingCountry id="7AA046C9897DFF95C7F36683FF439476" box="[151,218,1268,1293]" name="China" pageId="2" pageNumber="531">China</collectingCountry>
|
||
(Diding, Guangxi),
|
||
<collectingCountry id="7AA046C9897DFF95C6DA6682FDD89476" box="[446,577,1269,1293]" name="Hong Kong" pageId="2" pageNumber="531">Hong Kong</collectingCountry>
|
||
(Lantau).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |