145 lines
18 KiB
XML
145 lines
18 KiB
XML
<document id="915E17680E1ABC94EBEC93512B67C3A6" ENCODING="UTF8" ID-GBIF-Dataset="0ad0b492-3b2e-433f-9d58-3b4e05fb5d63" ModsDocID="22171" checkinTime="1249509882759" checkinUser="christiana" docAuthor="Eguchi, K." docDate="2008" docId="89E752138D40930C93108D0C8AB7A2CC" docLanguage="en" docName="22171.xml" docOrigin="Zootaxa 1902" docSource="http://hol.osu.edu/reference-full.html?id=22171" docTitle="Pheidole tumida Eguchi, 2008, sp.n." docType="treatment" docVersion="10" lastPageNumber="100" masterDocId="B03BC2DB73BB189B7C160C6A61E2E599" masterDocTitle="A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)." masterLastPageNumber="118" masterPageNumber="1" pageNumber="97" updateTime="1701311764087" updateUser="plazi">
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<mods:title id="C359002F67D8FD932BE659D3B704228A">A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae).</mods:title>
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<mods:namePart id="B3357DB9544858CDCC998C5E8DF7157B">Eguchi, K.</mods:namePart>
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<mods:identifier id="9A849F937E238D882CDBC052F08A4C7D" type="HNS-Pub">22171</mods:identifier>
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<treatment id="89E752138D40930C93108D0C8AB7A2CC" ID-DOI="http://doi.org/10.5281/zenodo.6235672" ID-GBIF-Taxon="100122337" ID-Zenodo-Dep="6235672" LSID="urn:lsid:plazi:treatment:89E752138D40930C93108D0C8AB7A2CC" httpUri="http://treatment.plazi.org/id/89E752138D40930C93108D0C8AB7A2CC" lastPageNumber="100" pageNumber="97">
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<subSubSection id="1768F6679BA8ACEA933F7FBE0D87022E" type="nomenclature">
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<paragraph id="44C487A724E5DC64A974BD47F8DA17CE" pageNumber="97">
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<taxonomicName id="080CF7D1EDFAB90A475A9DCC9AB88DD6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="tumida">Pheidole tumida</taxonomicName>
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<taxonomicNameLabel id="5F4A271F8034DA5426FBDC2B8BF0CFC6" rank="species">sp.n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="1FD33389C7EE7CD922BF172DF81D0302" type="description">
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<paragraph id="8230E8F525281F78487B55FB3E0881D6" pageNumber="97">Figs. 27a-g</paragraph>
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</subSubSection>
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<subSubSection id="B79CE7230AEDFD73149CC4BFA5595DEC" type="reference_group">
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<paragraph id="160E52488A8E8A83025C1E7A269DAF3B" pageNumber="97">
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<taxonomicName id="A7E2235E67B955DFDE11790B82A5B460" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33900" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodifera (Smith)" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="nodifera">Pheidole nodifera F. Smith</taxonomicName>
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. Eguchi, Bui et al. 2005: 90 (checklist).
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</paragraph>
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<paragraph id="2C0F6D289258F92F86926875262F0979" pageNumber="97">
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<taxonomicName id="D3F22AED234BF8800AA7AE4896EF7A65" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
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sp. (cf.
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<taxonomicName id="4BAF6DE138F9B8214F98EA5066DF3585" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33900" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodifera (Smith)" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="nodifera">nodifera</taxonomicName>
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). Eguchi, Bui & Janssen 2005 (ecological study).
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</paragraph>
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</subSubSection>
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<subSubSection id="DA25FE4550ABED7AADCA0167EAF063E6" type="materials_examined">
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<paragraph id="A575C540F9C32FD6306C2F8C32076BEB" pageNumber="97">
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Type material examined: Holotype: major, Cuc Phuong N.P., Ninh Binh, Vietnam (
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<geoCoordinate id="691E9463FFC68E1F6544FA1DF94C635E" direction="north" orientation="latitude" value="20.233334">20°14'N</geoCoordinate>
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,
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<geoCoordinate id="2AAD1FE568394E8FE98477549AD675FF" direction="east" orientation="longitude" value="105.6">105°36'E</geoCoordinate>
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, 370 m alt.) [K. Eguchi leg., colony: Eg01-VN-176] (IEBR); paratypes: 18 majors, 20 minors & 1 queen from the same colony as holotype (IEBR,
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<collectionCode id="B9FF4097E14F3D5720B89FED33A44582" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
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,
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<collectionCode id="0A1ED2847E712F36C2CBA0067DEA8801" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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& ACEG).
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</paragraph>
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<paragraph id="AFA7C54FDF959D0A2E1A5C311CB669E8" pageNumber="97">
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Other material examined: S. China: Guangxi: Gao Zhai (300 m alt.), Xing An County [Eg00-GNGX- 012]; Hong Kong: Victoria Park, Hong Kong I. [Eg99-HK-22], Taipo Kau, New Territory [Eg99-HK-07]. Vietnam: Bac Kan: Ba Be N.P., 22°24-25'N, 105°37-38'E, <260 m alt. [Eg04-VN-158, -164, -175, -177, - 180, -200, -206]; Quang Ninh: Ky Thuong
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<taxonomicNameLabel id="11CC3337206A7D4CAA9E9FBC55E6784A" rank="subSpecies">N.R.</taxonomicNameLabel>
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, 21°09-11'N, 107°06-07'E, 105-550 m alt. [Eg03-VN-165, - 191, -247; B&E03-77], Chua Yen Tu,
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<geoCoordinate id="1AE893E12C5DC08B0B2AA928777518B6" direction="north" orientation="latitude" value="21.15">21°09'N</geoCoordinate>
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,
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<geoCoordinate id="75A95FC2546615D159BF09D86513D375" direction="east" orientation="longitude" value="106.71667">106°43'E</geoCoordinate>
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, 520-845 m alt. [Eg04-VN-006, -010, -025]; Bac Giang: W. Yen Tu N.P. (=Tay Yen Tu N.P.), 21°10-11'N, 106°43-44'N, 170-415 m alt. [Eg03-VN-028, -039, -042, -056, -057, -073, -075, -099; Eg04-VN-086, -095; B&E03-13, -14, -19, -24]; Ha Tay (misspelled as "Ha Tai"): Ba Vi N.P.,
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<geoCoordinate id="A3C7B95343BBDC037BEE0DF80B22F546" direction="north" orientation="latitude" value="21.05">21°03'N</geoCoordinate>
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,
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<geoCoordinate id="7B57F0E1B0446530FC55218DB3F34D88" direction="east" orientation="longitude" value="105.36667">105°22'E</geoCoordinate>
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, 400-600 m alt. [Eg99-VN-081, -108; Eg02-VN-012]; Ninh Binh: Cuc Phuong N.P.,
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<geoCoordinate id="D7AEF82863F1F0D8662BE7C37C690DD7" direction="north" orientation="latitude" value="20.233334">20°14'N</geoCoordinate>
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,
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<geoCoordinate id="8ADB1A907BDC0226DFE3F96964E01022" direction="east" orientation="longitude" value="105.6">105°36'E</geoCoordinate>
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, 370 m alt. [Eg01-VN-162, -163, -170]; Nghe An: Pu Mat N.P. (Sang Le Forest Area,
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<geoCoordinate id="DCAF8F28FA43F71AF1E07FA5BB770831" direction="north" orientation="latitude" value="19.183332">19°11'N</geoCoordinate>
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, 104°37-38'E, <220 m alt.) [Eg01iv06-09]; Dong Nai: S. Cat Tien N.P., <160 m alt. [Eg04-VN-553]. Thailand: Chanthaburi: Khao Soi Dao [Eg01-TH-024]; Trang: Khao Chong Waterfall [Eg01- TH-750]; Narathiwat: Bala-Hala W.S. (Bala Area) [Eg01-TH-686]. Malaysia: Selangor: Ulu Gombak [FI98- 84]. Indonesia: Sungai Wain, E. Kalimantan [G. Fredriksson]; W. Java: Halimun [FI98-328]; E. Java: Purwodai[FI98-277]; Bali: Dusun PK Jelati, Mendaya [Eg98-BALI-727, -748], Mt. Kelatakan [Eg98-BALL-1111, - 1114; IKT. Ginarsa's colony: KT-163]. Eguchi's informal species code "
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<taxonomicName id="245EB954085605AFA6431FDCDCA265A0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
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sp. eg-100" applies to this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="49B42D2A84D503EB6EC2EDC98718EB50" type="description">
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<paragraph id="3ADFCC89BC879FEE7DA158CA400E4149" pageNumber="97">Worker measurements & indices: Holotype (major). - HL 1.98 mm; HW 1.99 mm; CI 101; SL 1.11 mm; SI 56; FL 1.58 mm; FI 79.</paragraph>
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<paragraph id="A3C491DF6982ECEA7EB1496984CD0611" pageNumber="97">Nontype major (n=4). - HL 1.63-1.90 mm; HW 1.63-1.98 mm; CI 100-105; SL 0.93-1.07 mm; SI 52- 60; FL 1.38-1.54 mm; FI 78-85.</paragraph>
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<paragraph id="73F5DDA856486A18DD71E40DC69A247A" pageNumber="97">Minor (n=5, including one paratype minor). - HL 0.71-0.78 mm; HW 0.57-0.66 mm; CI 79-87; SL 0.91-1.04 mm; SI 147-166; FL 1.03-1.18 mm; FI 166-192.</paragraph>
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<paragraph id="00335B9B41B9E89D0536A5E26DEB72DB" pageNumber="97">Worker description</paragraph>
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<paragraph id="CC84D492E0D4F55F486FD10161FCE0CC" lastPageNumber="99" pageNumber="97">
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Major. - Head in lateral view not or very weakly impressed on vertex; frons and anterior part of vertex rugose longitudinally; posterior part of vertex and dorsal and dorsolateral faces of vertexal lobe rugoso-reticulate obliquely, or reticulate; frontal carina weak, or present just as rugula(e); antennal scrobe inconspicuous or absent; median longitudinal carina of clypeus usually conspicuous, but sometimes weak or just present as weak rugula(e); median and submedian processes of hypostoma absent, or sometimes present but inconspicuous; lateral processes conspicuous but often small; antenna with a 3-segmented club; maximal diameter of eye longer than or almost as long as antennal segment X. Promesonotal dome in dorsal view rugoso-reticulate transversely or irregularly, in lateral view with a conspicuous prominence or at least low mound on its posterior slope; humerus (very) weakly produced laterad; the dome at the humeri narrower than at the bottom (or
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<pageStartToken id="4BE282201B55ABD95BFC5CB6D1AEB846" pageNumber="99">sometimes</pageStartToken>
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almost as broad as or a little broader than at the bottom); propodeal spine usually finger-shaped or spatulate. Petiole shorter than postpetiole (excluding helcium), in dorsal view with a well-developed flange laterally; subpetiolar process extremely developed, lobate; postpetiole massive. First gastral tergite rugoso-punctate or at least shagreened over the surface.
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</paragraph>
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<caption id="2278B8DCE01E16D79394DF113AC2B742">
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<paragraph id="B777FE662E7884304C492499EC5C06C8" pageNumber="98">
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FIGURE 27a-d,
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<taxonomicName id="237FE017312B706832126B3F81EB5471" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="98" phylum="Arthropoda" rank="species" species="tumida">Pheidole tumida</taxonomicName>
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<taxonomicNameLabel id="C8981F698864EF574D79757C1355935F" rank="species">sp.n.</taxonomicNameLabel>
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, holotype (major) [Eg01-VN-176] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
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</paragraph>
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</caption>
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<caption id="20C409181F71B6B1064FEAAC178329A9">
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<paragraph id="90240BEE2B562F93887AB99536368F3A" pageNumber="99">
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FIGURE 27e -g,
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<taxonomicName id="47AAE4482F2C33F637DB7226863CA322" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="99" phylum="Arthropoda" rank="species" species="tumida">Pheidole tumida</taxonomicName>
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<taxonomicNameLabel id="856CF151235F5D212041CB6AF4EE3C62" rank="species">sp.n.</taxonomicNameLabel>
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, paratype minor [Eg01-VN-176] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
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</paragraph>
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</caption>
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<paragraph id="BF8088E91CB808088E71CAAED408BB22" lastPageNumber="100" pageNumber="99">
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Minor. - Frons and vertex largely smooth; area between antennal insertion and eye often rugose or rugoso-punctate; preoccipital carina conspicuous dorsally and laterally; median part of clypeus smooth and shining; median longitudinal carina often present; antenna with a 3-segmented club; scape extending far beyond
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<pageStartToken id="86715F4C103F74A13F4740DCCAD79F66" pageNumber="100">posterolateral</pageStartToken>
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margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome largely smooth, but sometimes shagreened dorsolaterally and/or with several short rugulae on humerus, in lateral view with a conspicuous mound on its posterior slope; humerus of the dome in dorso-oblique view not or hardly produced (rarely produced weakly as a mound); mesopleuron, metapleuron and lateral face of propodeum dimly to weakly punctured at least partly; propodeal spine usually reduced to a short and thin spine or a small dent, or sometimes almost absent. Petiole shorter than postpetiole (excluding helcium); subpetiolar process absent, or present as a longitudinal carina; postpetiole massive.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C44576190EFA52FD1E99307A7B36A2C" type="discussion">
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<paragraph id="7D1950C5A43FE27E6A06032FCED54E05" pageNumber="100">Recognition: This species has the following combination of diagnostic characteristics: in the minor head and promesonotal dome largely smooth and shining; in the major hypostoma without median and submedian processes; in the major and minor promesonotal dome in lateral view with a conspicuous prominence or mound on its posterior slope; in the major subpetiolar process lobate; in the major and minor postpetiole massive.</paragraph>
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<paragraph id="1655C8850A7F0A16AF37EE39437D7423" pageNumber="100">
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<taxonomicName id="296C6192980A0B5E966BB416886BDAD6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="tumida">Pheidole tumida</taxonomicName>
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,
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<taxonomicName id="B732A9E27C4738B5A2663F23D4B3127D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33898" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole noda Smith" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="noda">Pheidole noda</taxonomicName>
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and
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<taxonomicName id="90CEB1499388518288595FBA59D95DF3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33900" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodifera (Smith)" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="nodifera">Pheidole nodifera</taxonomicName>
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are not distinguished from each other in minor's morphology. Thus, there is a certain possibility that
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<taxonomicName id="AB32470820AF6A35658D70FD984DFB66" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="29" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="tumida">P. tumida</taxonomicName>
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is conspecific with
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<taxonomicName id="9237F3078BB6173AF31DFBC8FF339AA5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33900" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="78" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodifera (Smith)" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="nodifera">P. nodifera</taxonomicName>
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which was described by F. Smith (1874) based on the minor alone from N. China.
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<taxonomicName id="083EFFD302017FE464CD4CE8548345EA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181984" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole humida Wilson" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="tumida">Pheidole tumida</taxonomicName>
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, characterised by a lobate subpetiolar process in the major, is easily distinguished from
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<taxonomicName id="1F702A3B15853E4ADFEA3378A27B420B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33898" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole noda Smith" order="Hymenoptera" pageNumber="100" phylum="Arthropoda" rank="species" species="noda">Pheidole noda</taxonomicName>
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and other Indo-Chinese species.
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</paragraph>
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</subSubSection>
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<subSubSection id="598CCBB5B7C73117B7495C78A768AB26" type="distribution">
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<paragraph id="9B4B946D2DADA453D952009804BED2C4" pageNumber="100">Distribution & bionomics: Known from the Indo-Chinese and Indo-Malayan subregions. Ranging from forests edges to well-develop forests. Nesting in the soil and rotting logs.</paragraph>
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</subSubSection>
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</treatment>
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</document> |