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<document ID-CLB-Dataset="7916" ID-DOI="http://dx.doi.org/10.3897/subtbiol.38.61733" ID-GBIF-Dataset="eb517ca7-0640-4667-840f-30e4fcf67a61" ID-Pensoft-Pub="1314-2615-38-1" ID-Pensoft-UUID="91E16135CFC6599C890A767C46346BB1" ID-ZooBank="CBC7B4A64CEE41D1A6679094CE8ACB3F" ModsDocID="1314-2615-38-1" checkinTime="1614718792233" checkinUser="pensoft" docAuthor="Garcia 1, Rafael, Andujarx, Carmelo, Oromi, Pedro, Emerson, Brent &amp; Lopez, Heriberto" docDate="2021" docId="89B050D79E6C57B4AE1E7660F59019AC" docLanguage="en" docName="SubterBiol 38: 1-18" docOrigin="Subterranean Biology 38" docPubDate="2021-03-02" docSource="http://dx.doi.org/10.3897/subtbiol.38.61733" docTitle="Baezia madai Garcia &amp; Oromi 2021, sp. nov." docType="treatment" docUuid="C1BDDC6F-6737-407B-88FC-D76E45875C59" docUuidSource="ZooBank" docVersion="6" id="91E16135CFC6599C890A767C46346BB1" lastPageNumber="1" masterDocId="91E16135CFC6599C890A767C46346BB1" masterDocTitle="Three new subterranean species of Baezia (Curculionidae, Molytinae) for the Canary Islands" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1732823915817" updateUser="ExternalLinkService">
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<mods:title id="006DB500BE02F40178A5C4F7EA7C89F3">Three new subterranean species of Baezia (Curculionidae, Molytinae) for the Canary Islands</mods:title>
</mods:titleInfo>
<mods:name id="AEB37269498B2C6B1A6A31BBE396AE79" type="personal">
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<mods:namePart id="8A4C7F9CA85DB1C1A4E1010830BC3977">Garcia 1, Rafael</mods:namePart>
<mods:affiliation id="55589E2304475D0647B4C2CB26A54833">C /. San Miguel 9, 38700 Santa Cruz de La Palma, La Palma, Canary Islands, Spain</mods:affiliation>
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<mods:namePart id="271D68688644B2D2B72EAA3D50D21449">Andujarx, Carmelo</mods:namePart>
<mods:affiliation id="225B7C8A47512DF35277F9C94158B643">Island Ecology and Evolution Research Group, Instituto de Productos Naturales y Agrobiologia (IPNA-CSIC), 38206, La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
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<mods:namePart id="009FA2EF6ADAA19C8F2BD703272D44BD">Oromi, Pedro</mods:namePart>
<mods:affiliation id="5BB0E619AF49445B069A84C4CBD81309">Depto. Biologia Animal, Edafologia y Geologia, Universidad de La Laguna, 38206, La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
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<mods:namePart id="CACF1AF5BC37F09056E8F99B714E1579">Emerson, Brent</mods:namePart>
<mods:affiliation id="AA4CFEB771C9E73FC47539DEE89259A1">Island Ecology and Evolution Research Group, Instituto de Productos Naturales y Agrobiologia (IPNA-CSIC), 38206, La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
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<mods:namePart id="4FA0AE043E23C3A713156C5B0CD37697">Lopez, Heriberto</mods:namePart>
<mods:nameIdentifier id="0FCD9D8028C6ECD86ED221C493634A88" type="ORCID">https://orcid.org/0000-0001-6988-5204</mods:nameIdentifier>
<mods:affiliation id="207CF3583EA59365C80C78AFEF93EDC0">Island Ecology and Evolution Research Group, Instituto de Productos Naturales y Agrobiologia (IPNA-CSIC), 38206, La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
<mods:nameIdentifier id="91745AE23CDF1F0D7D75166C085FDCFC" type="email">herilope@ipna.csic.es</mods:nameIdentifier>
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<mods:typeOfResource id="372FCBD8FDA48137EC2E7B8237E30487">text</mods:typeOfResource>
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<mods:title id="3CD9918839FB93F0DC49BED0AD560D07">Subterranean Biology</mods:title>
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<mods:part id="9C49CA8BEEFAC778A51D60849DCA419B">
<mods:date id="B730E8BBE2E9E940DB803E02CB714364">2021</mods:date>
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<mods:number id="5CEB2D3845CAA6031E9B5C216EDB2C1C">2021-03-02</mods:number>
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<mods:number id="269B71264707EDAAA8A11D72E7CAFF45">38</mods:number>
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<mods:url id="7B1C34D2881AC585263C24DF61CD8400">http://dx.doi.org/10.3897/subtbiol.38.61733</mods:url>
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<mods:classification id="CBFC1EF8A6E6549D68C4228130FF3A0A">journal article</mods:classification>
<mods:identifier id="24DCE7843067239F1CEA21C67968FA55" type="DOI">http://dx.doi.org/10.3897/subtbiol.38.61733</mods:identifier>
<mods:identifier id="D0F78A66B980365CCE0B9B93EC14502F" type="Pensoft-Pub">1314-2615-38-1</mods:identifier>
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<treatment id="89B050D79E6C57B4AE1E7660F59019AC" ID-GBIF-Taxon="179985092" LSID="urn:lsid:zoobank.org:act:C1BDDC6F-6737-407B-88FC-D76E45875C59" httpUri="http://treatment.plazi.org/id/89B050D79E6C57B4AE1E7660F59019AC" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Insecta" scope_family="Curculionidae" scope_order="Coleoptera" scope_subFamily="Molytinae" scope_superFamily="Curculionoidea" scope_tribe="Typoderini">
<subSubSection id="05621C53FDB59C203A2C3C2588E99E81" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="ACA4D5506BFFF83A6929442E6BA7BC02" pageId="0" pageNumber="1">
<taxonomicName id="87F9751DD1F32E0B959FFA3653F0F32A" LSID="http://zoobank.org/C1BDDC6F-6737-407B-88FC-D76E45875C59" authority="García &amp; Oromí" authorityName="Garcia &amp; Oromi" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Baezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baezia madai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madai" status="sp. nov.">
Baezia madai
<normalizedToken id="3AB3743060E7C881226EC55B33C7631C" originalValue="García">Garcia</normalizedToken>
&amp;
<normalizedToken id="EF4A7626C3EFF665574559117A6FA10F" originalValue="Oromí">Oromi</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="0DDA636C9AC4E8C0D8ACC95BCCA0CE1E" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="1FD5A3419AF39A7D78D206567FFEA250" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">Figs 3A-E</figureCitation>
<figureCitation id="FF0F857BE4C9ABC74DEAE02E0273D0F5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hind leg and pronotum of A, B Baezia litoralis C, D B. vulcania E, F B. madai sp. nov. G, H B. tizziri sp. nov. I, J B. aranfaybo sp. nov. K, L B. martini M, N B. bimbache." figureDoi="10.3897/subtbiol.38.61733.figure5" httpUri="https://binary.pensoft.net/fig/515630" pageId="0" pageNumber="1">, 5E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DCE0B9D6353E3D36B6548E207D0F9371" pageId="0" pageNumber="1" type="type locality">
<paragraph id="EACA3C005EEBB5444E590F84C3192511" pageId="0" pageNumber="1">
<typeStatus id="EE54330ED4F2814BF98CA0BC9FCDB5D6">Type</typeStatus>
locality.
</paragraph>
<paragraph id="20E41AF52202F757634494CA58DAAFF5" pageId="0" pageNumber="1">
<materialsCitation id="19BF97C8F32F70D6DDAECCDBD3B47A65" ID-GBIF-Occurrence="3063649303" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="La Palma &amp; El Paso &amp; de Barros" country="Spain" elevation="536" latitude="28.662191" location="Cueva de Barros" longLatPrecision="1" longitude="-17.889992" municipality="Cueva de Las Tijaraferas" specimenCount="1" stateProvince="Canarias" typeStatus="holotype">
<collectingCountry id="065A02C597E1F6726E568BAE50FFFA82" name="Spain">Spain</collectingCountry>
,
<collectingRegion id="A3EA35CD094F5B6F00BAAC9319488DD7" country="Spain" name="Canarias">Canary Islands</collectingRegion>
,
<collectorName id="32E971A5CE14ACFC09E411D2FC8714B0">La Palma</collectorName>
,
<collectorName id="0F66A55587A9AED3E58E803CA9C62281">El Paso</collectorName>
:
<collectingMunicipality id="AB60FBDCADEDC8D231224F4FB541CDAF">Cueva de Las Tijaraferas</collectingMunicipality>
, also named
<location id="D73A9485A329DCFCF1E5873C312A7E78" LSID="urn:lsid:plazi:treatment:89B050D79E6C57B4AE1E7660F59019AC:D73A9485A329DCFCF1E5873C312A7E78" country="Spain" latitude="28.662191" longLatPrecision="1" longitude="-17.889992" municipality="Cueva de Las Tijaraferas" name="Cueva de Barros" stateProvince="Canarias">
Cueva
<collectorName id="13680A18C77F0DE383584FFE559B780F">de Barros</collectorName>
</location>
(
<geoCoordinate id="D7568A00C28E0C170EA33EE2262256D1" degrees="28" direction="north" minutes="39" orientation="latitude" precision="1" seconds="43.89" value="28.662191">28°39'43.89&quot;N</geoCoordinate>
,
<geoCoordinate id="F09BD30609D34D1F37DEDBB4107C806A" degrees="17" direction="west" minutes="53" orientation="longitude" precision="1" seconds="23.97" value="-17.889992">17°53'23.97&quot;W</geoCoordinate>
,
<elevation id="4A3C48157D12E84223423C1D02A490F7" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">
<quantity id="9AB9612C830DBECE23127299DEDF9D8A" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">536 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D776CDF6CD792A1EB516F69E632E2199" pageId="0" pageNumber="1" type="type material">
<paragraph id="212964814809299178ED1B312DC124EB" pageId="0" pageNumber="1">
<typeStatus id="A2F751A395210777FBB9CD0C0C3991E0">Type</typeStatus>
material.
</paragraph>
<paragraph id="6D764A4E1A0BE9D78FF105A88CA94866" pageId="0" pageNumber="1">
<emphasis id="A98FD6BB3D2BDFB26F5EA00BDE75E867" bold="true" pageId="0" pageNumber="1">
<emphasis id="01B17A41E0AC4E94B03BDDA8189F6C2B" italics="true" pageId="0" pageNumber="1">
<typeStatus id="61AF3CA38C264215DF586FAF3FA67666">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation id="E3C47EC317F8D57C7B2E58963B96C057" ID-GBIF-Occurrence="3063649302" collectingDate="1986-07-10" collectionCode="DZUL" collectorName="La Palma &amp; El Paso &amp; de Barros &amp; J. L. Martin" county="Cueva de Barros" elevation="536" latitude="28.662191" location="La Palma" longLatPrecision="1" longitude="-17.889992" municipality="Cueva de Las Tijaraferas" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="C304E540D43ED4CBD4AEDC96629B4A17" type="male">1♂</specimenCount>
,
<collectorName id="7F93FD375BBDABF477C48A9152ACFE99">
<location id="F17B699E89FECE080FF5EB2957C8F281" LSID="urn:lsid:plazi:treatment:89B050D79E6C57B4AE1E7660F59019AC:F17B699E89FECE080FF5EB2957C8F281" county="Cueva de Barros" latitude="28.662191" longLatPrecision="1" longitude="-17.889992" municipality="Cueva de Las Tijaraferas" name="La Palma">La Palma</location>
</collectorName>
,
<collectorName id="6D2CCCA3ED487AA2B8E70A01884A4F66">
<location id="443C45383DFB3372E14F5AA1EE96C633" LSID="urn:lsid:plazi:treatment:89B050D79E6C57B4AE1E7660F59019AC:443C45383DFB3372E14F5AA1EE96C633" county="Cueva de Barros" latitude="28.662191" longLatPrecision="1" longitude="-17.889992" municipality="Cueva de Las Tijaraferas" name="El Paso">El Paso</location>
</collectorName>
,
<collectingMunicipality id="98D8F807E27E2BF56E95E1DEF02A9E36">Cueva de Las Tijaraferas</collectingMunicipality>
, also named
<collectingCounty id="5A47F7A4467E141ECA9AEAD3BF284229">
Cueva
<collectorName id="DA9F2B3F908070C3A6B3B4AEC7907AA6">de Barros</collectorName>
</collectingCounty>
(
<geoCoordinate id="D4A593182B85F928CAA0332374670C5C" degrees="28" direction="north" minutes="39" orientation="latitude" precision="1" seconds="43.89" value="28.662191">28°39'43.89&quot;N</geoCoordinate>
,
<geoCoordinate id="D32845F60B992044194E478B78AB48AE" degrees="17" direction="west" minutes="53" orientation="longitude" precision="1" seconds="23.97" value="-17.889992">17°53'23.97&quot;W</geoCoordinate>
,
<elevation id="023953F307778729DD8CB08CE71D3989" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">
<quantity id="7FB233D38E40089FA276015DD2AE107A" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">536 m</quantity>
a.s.l.
</elevation>
),
<date id="1740B788CF7CB1581FB9A2BA0C9E9379" value="1986-07-10">
<collectingDate id="417AE08D0E6B25205826A68032ECDE03" value="1986-07-10">10 July 1986</collectingDate>
</date>
,
<collectorName id="1DE511E008F0C199D740767FFB0AC46E">
J.L.
<normalizedToken id="ADC35C064EAC2324C295D731ED859F71" originalValue="Martín">Martin</normalizedToken>
</collectorName>
leg. (
<collectionCode id="23C9CD7A0587B1B1861DEA7231E61567" country="Spain" httpUri="http://grbio.org/cool/x2t0-vyyc" name="Departamento de Zoologia, Universidad de La Laguna" type="">DZUL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4E921C537BD6A4BB9971CDF0DFEEF707" pageId="0" pageNumber="1" type="description">
<paragraph id="F30899E92A555439318F7829EF3AD547" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="95CF98644E2F1A67233D3E3C25ACA600" pageId="0" pageNumber="1">
<emphasis id="3A10FBC2CAB3F02ED457A4839D5B19F6" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
Total length (including rostrum)
<quantity id="9925F78D8A1EDCC1D47C1210492E4CCA" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
,
<quantity id="E487B41199A98B7037EFA41BF0C5E556" metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
without rostrum and head, and maximum width
<quantity id="0E3B3E74B2B9854836380FAE49032F4C" metricMagnitude="-4" metricUnit="m" metricValue="9.5" unit="mm" value="0.95">0.95 mm</quantity>
. Body matte reddish-brown (Fig.
<figureCitation id="171D9D8A75CBC69DBDD3B3793EBE5CBE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">3A</figureCitation>
); apex of rostrum, antennae and legs covered with scattered yellow-testaceous erect setae, pronotum and elytra with short fine testaceous and claviform pilosity denser and aligned on whole surface. Apterous.
</paragraph>
<paragraph id="0C85D6BB68E230E9BD49B1BF9FEB9E39" pageId="0" pageNumber="1">
<emphasis id="DAE81D961D182753C693D9858D5A894B" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
partially retracted into pronotum, microreticulated with abundant irregular punctation, lacking eyes.
</paragraph>
<caption id="CE446A563824187C3FE459B8E3713A89" doi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="DA0D7B4D1A6F734D85ECB6DD915D44ED" pageId="0" pageNumber="1">
<emphasis id="CB9B0061CF3F5CD2AF876F450E959F18" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="2197F00B77EA222A7DAE3A6089E4BF6A" authorityName="Garcia &amp; Oromi" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Baezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baezia madai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madai">
<emphasis id="8BBA60AC924BD38A9A558E0F85465247" italics="true" pageId="0" pageNumber="1">Baezia madai</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="78E83943BCFBFFCB95BB501CB39D5E94" bold="true" pageId="0" pageNumber="1">A</emphasis>
dorsal habitus
<emphasis id="27679C979886A17B7D74E5DEEAE1BF7A" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
aedeagus in dorsal and lateral view
<emphasis id="B505B803F95A97EBEEF21F577F34B05B" bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="4B265D4B5B6A6E63ED6BBFA09A95A07F" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="F7227F9AA4E752ACC927DC28189042D0" authorityName="Brauckmann &amp; Herd" authorityYear="2005" class="Insecta" family="Hapalopteridae" genus="Spiculum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Spiculum" order="Caloneurodea" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Spiculum</taxonomicName>
gastrale
</emphasis>
<emphasis id="9A5BD085D288CCC8DB925025917E5007" bold="true" pageId="0" pageNumber="1">E</emphasis>
tegmen.
</paragraph>
</caption>
<paragraph id="BB2A553BC7F046E3C965C46A36CEF1B2" pageId="0" pageNumber="1">
<emphasis id="EB079A54A3D10079FCF1F2A860933845" bold="true" italics="true" pageId="0" pageNumber="1">Rostrum</emphasis>
robust, widest at antennal insertion, 2
<normalizedToken id="DEB364132BC351626F4D04D98BBA95EF" originalValue="×">x</normalizedToken>
as long as wide at scrobes level, 0.63
<normalizedToken id="8AF49D4E03E94C644CCA229780C8C630" originalValue="×">x</normalizedToken>
as long as pronotum. In lateral view lower margin concave, upper margin slightly convex, more declivous near apex; apex punctated, shiny, with erect setae. Scrobes deep, their apical third visible from above. Mandibles smooth and black. Rostrum slightly more depressed than forehead, with dorsal surface irregular with longitudinal sulci separated by five fine broken keels; ventral surface rough.
</paragraph>
<paragraph id="248094C4C3626EBB15959AFAE77C46DB" pageId="0" pageNumber="1">
<emphasis id="2D87ACF801B8BB88EFB2B3DEA0C0328F" bold="true" pageId="0" pageNumber="1">
<emphasis id="995C0B09776D67E8E02853F36FB6B7A4" italics="true" pageId="0" pageNumber="1">Antennae</emphasis>
.
</emphasis>
Scapes straight, increasingly widened from middle, 5.6
<normalizedToken id="D2E1EC861560A3DCCFEFE67DDEA5D70E" originalValue="×">x</normalizedToken>
as long as its maximum width. The specimen lacks the rest of the antennae.
</paragraph>
<paragraph id="EF01F81D40D03B157A85D8BA18564BC8" pageId="0" pageNumber="1">
<emphasis id="AF374E499D5EF0A8B23C960DFEA14A0F" bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
slightly elongated with fine median keel, maximum width towards middle and sides slightly convex, constricted behind apex, with a slight sinuation at middle; anterior margin as wide as posterior one (Fig.
<figureCitation id="96FC04521FA40031C5A16CEBA204EEBE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hind leg and pronotum of A, B Baezia litoralis C, D B. vulcania E, F B. madai sp. nov. G, H B. tizziri sp. nov. I, J B. aranfaybo sp. nov. K, L B. martini M, N B. bimbache." figureDoi="10.3897/subtbiol.38.61733.figure5" httpUri="https://binary.pensoft.net/fig/515630" pageId="0" pageNumber="1">5F</figureCitation>
). Surface matte chagrinated with microreticulation; punctures obvious, almost coalescent, setae lying down and scattered, little more erect towards margins.
</paragraph>
<paragraph id="CA7058E89169E29DC49BA35E1E0E7BA8" pageId="0" pageNumber="1">
<emphasis id="EAB1C77EB6A514E1BB764D5615EA5CBB" bold="true" italics="true" pageId="0" pageNumber="1">Scutellum</emphasis>
very small, triangular.
</paragraph>
<paragraph id="99CCCCD105D15C28520AA0A8E4E99B3F" pageId="0" pageNumber="1">
<emphasis id="C116987F66746D03770DC8050E0E5F1F" bold="true" italics="true" pageId="0" pageNumber="1">Pterothorax</emphasis>
with elytra elongate, lacking humeral calli; 2.6
<normalizedToken id="29111D201383DE7E1F05255C090256CA" originalValue="×">x</normalizedToken>
as long as pronotum, 1.84
<normalizedToken id="6332DF3D6D54631714361F7F2026C7C0" originalValue="×">x</normalizedToken>
as long as wide, base wider than base of pronotum; maximum width towards middle, basal margin 0.68
<normalizedToken id="9803445DB86CF9634AABC8AF7AA1D043" originalValue="×">x</normalizedToken>
that width. Surface matte, chagrinated, strongly microreticulated; interstriae smooth; striae very fine, slightly defined by aligned punctures coinciding with small, erect setae.
</paragraph>
<paragraph id="19248B319EC4011C93C7AB1C6ED06D3D" pageId="0" pageNumber="1">
<emphasis id="AAAD6F3EED08C25A9AFFAF212991CA65" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
with integument surface slightly shiny, microreticulated; with fine, short setose pilosity; well-defined punctation. First and second ventrites with wide median depression; remaining ones slightly convex, a little but increasingly elevated towards apex, suddenly cut down at end, giving stair-like appearance. Ventrite 5 2.4
<normalizedToken id="ED94A2CEA3AC0CDB1AB4EE420762C3B0" originalValue="×">x</normalizedToken>
as wide as long, strongly chagrinated, with deep punctation.
</paragraph>
<paragraph id="24988D6586CDA154BB42FB24E4FD5685" pageId="0" pageNumber="1">
<emphasis id="C6A813C70FBFAADCC695D324F57CED49" bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
elongate, with matte surface, microreticulate with abundant semierect setae. Procoxae separated by distance of 0.16
<normalizedToken id="EF4FC75903920D2DE172D1844418BFE0" originalValue="×">x</normalizedToken>
of their diameter. Mesocoxae separated by distance of 0.71
<normalizedToken id="BF6304134018612E30ECA7CC4E0B077B" originalValue="×">x</normalizedToken>
of their diameter, and metacoxae 1.63
<normalizedToken id="7B9DFA45AA76625F8D9074EE75C016EA" originalValue="×">x</normalizedToken>
of their diameter. Femora slightly dilated at middle, from middle they gradually narrow until they strangle near apex (Fig.
<figureCitation id="FECAC04BF0B33D55B9249A968A2FB296" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hind leg and pronotum of A, B Baezia litoralis C, D B. vulcania E, F B. madai sp. nov. G, H B. tizziri sp. nov. I, J B. aranfaybo sp. nov. K, L B. martini M, N B. bimbache." figureDoi="10.3897/subtbiol.38.61733.figure5" httpUri="https://binary.pensoft.net/fig/515630" pageId="0" pageNumber="1">5E</figureCitation>
); pro-, meso- and metafemora 3.6
<normalizedToken id="79CDF4C70736C0652E7FDFFBA8122ED9" originalValue="×">x</normalizedToken>
, 4
<normalizedToken id="94384746E7687D04CE2CF56AF1A4B486" originalValue="×">x</normalizedToken>
and 4.8
<normalizedToken id="9ABC6E5641F86A84F92345295BC6D55F" originalValue="×">x</normalizedToken>
respectively as long as their maximum width. Tibiae straight, external edge slightly convex; internally with weak apical sinuosity and small bump towards middle (Fig.
<figureCitation id="4B9B3A428856AECE8D49F0D4447EE92D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hind leg and pronotum of A, B Baezia litoralis C, D B. vulcania E, F B. madai sp. nov. G, H B. tizziri sp. nov. I, J B. aranfaybo sp. nov. K, L B. martini M, N B. bimbache." figureDoi="10.3897/subtbiol.38.61733.figure5" httpUri="https://binary.pensoft.net/fig/515630" pageId="0" pageNumber="1">5E</figureCitation>
); tibiae uncinate, with uncus provided of a sharp tip; pro-, meso- and metatibiae 4.75
<normalizedToken id="21A29B401C60B79371116E246434509B" originalValue="×">x</normalizedToken>
, 4.75
<normalizedToken id="5349EDE8F83C9CF82A53F44B5B5626D3" originalValue="×">x</normalizedToken>
and 6.66
<normalizedToken id="1E20DE562986B2BB22A8312806D534D0" originalValue="×">x</normalizedToken>
respectively as long as their maximum width (excluding uncus). Protarsi with tarsomeres I 1.67
<normalizedToken id="14A43E982F79CB9DCFD04A7E00B76C0F" originalValue="×">x</normalizedToken>
, II 0.76
<normalizedToken id="994BCD22A85676ED2D345D906D5970A2" originalValue="×">x</normalizedToken>
, III 0.87
<normalizedToken id="7C9386E27E9C72227FDC7909CCEAE185" originalValue="×">x</normalizedToken>
and
<collectionCode id="FC6A8CBA13037092534F175E15BF1767" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
2.5
<normalizedToken id="07ABC7997C6864CCC052929DDF577585" originalValue="×">x</normalizedToken>
as long as wide respectively, third one clearly bilobed, fifth bearing two free simple acute claws; tarsal sole brushes with long sparse hyaline hairs.
</paragraph>
<paragraph id="9E277EB3CF9D224170A8834484B349D8" pageId="0" pageNumber="1">
<emphasis id="8BB97C5AABFDEDCDBEFC5F77ACEE34E4" bold="true" pageId="0" pageNumber="1">
<emphasis id="D97418684D0F4A035457427CA6D1171E" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
.
</emphasis>
Median lobe almost symmetrical in dorsal view, sides slightly convex, apex rounded (Fig.
<figureCitation id="7C769B3B82FFCD4AA4F082625C5795D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">3B</figureCitation>
); clearly curved in lateral view, with acute apex (Fig.
<figureCitation id="C9D55A28BD261836399177E7AC89479B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">3C</figureCitation>
). Internal sac with abundant teeth and spicules arranged in two elongated groups, with large acute teeth mixed with others smaller and with asperities; three basal sclerotized pieces.
<emphasis id="1C0E37343AC741A53F748398C4345C2B" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="77F41EBB7632D2C322A20D2FE501AB40" authorityName="Brauckmann &amp; Herd" authorityYear="2005" class="Insecta" family="Hapalopteridae" genus="Spiculum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Spiculum" order="Caloneurodea" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Spiculum</taxonomicName>
gastrale
</emphasis>
robust and bowed with highly asymmetric arms (Fig.
<figureCitation id="1317AF5B54FA3FC865B9129D37695659" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">3D</figureCitation>
). Tegmen with short manubrium; wide, hairy parameroids forming lobes, separated by a deep wide notch slightly more than half its length (Fig.
<figureCitation id="45AFE766983400161C7D1F4FE222A06A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baezia madai sp. nov. A dorsal habitus B, C aedeagus in dorsal and lateral view D Spiculum gastrale E tegmen." figureDoi="10.3897/subtbiol.38.61733.figure3" httpUri="https://binary.pensoft.net/fig/515628" pageId="0" pageNumber="1">3E</figureCitation>
).
</paragraph>
<paragraph id="5D3A4ED227B5CF54D5373DAA54B3384D" pageId="0" pageNumber="1">
<emphasis id="A8155E93DE21377EDFFFEF8F3CFEB09F" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9531044BF090EDFCC2306E8517391467" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="E09143A0D6C83AC7E55ABDE4E96927AC" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="67CCFAD130ED5D3CF23CE18A514689BF" pageId="0" pageNumber="1">
This new species is morphologically close to
<taxonomicName id="B0FB0A8EAE39AEB0E183F1B58B7D1727" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. vulcania" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vulcania">
<emphasis id="3EC383D50B31D207812FFCEA8B443845" italics="true" pageId="0" pageNumber="1">B. vulcania</emphasis>
</taxonomicName>
. However,
<taxonomicName id="C1927B1A82234E6C298BDBA70075787A" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. madai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madai">
<emphasis id="27007F0E3715E76A0DB6D2EECD98E869" italics="true" pageId="0" pageNumber="1">B. madai</emphasis>
</taxonomicName>
can be differentiated by its smaller size, matte body surface and lesser and shorter pilosity; scape longer and increasingly widened towards middle; elongated pronotum, with sides almost straight (slightly convex in
<taxonomicName id="12079F0BF05BD944323F50A22744A944" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. vulcania" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vulcania">
<emphasis id="F1FBD7FA5B6470FC5961A8B841B25C1D" italics="true" pageId="0" pageNumber="1">B. vulcania</emphasis>
</taxonomicName>
) and with a weak median keel (absent in
<taxonomicName id="94180575EFCD0EABF58A6223C70F08DE" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. vulcania" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vulcania">
<emphasis id="40C79F9D60FD1088AB65D5C6E6AEE193" italics="true" pageId="0" pageNumber="1">B. vulcania</emphasis>
</taxonomicName>
). In addition, its elytra are proportionally longer, the femora and tibiae less dilated on inner side, the tibiae proportionally longer; the median lobe dorsally parallel-sided (sides gently rounded in
<taxonomicName id="D7DAE6267649CA5550169448C71EC194" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. vulcania" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vulcania">
<emphasis id="7FCCABCD75C7DFE67EEAFC262CEB6957" italics="true" pageId="0" pageNumber="1">B. vulcania</emphasis>
</taxonomicName>
) and acute apex (rounded and slightly prominent at middle in
<taxonomicName id="C471B1762D82E20A53266039F2026A1B" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. vulcania" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vulcania">
<emphasis id="433D5FBF6B781597F4A38F9F3EED9A8E" italics="true" pageId="0" pageNumber="1">B. vulcania</emphasis>
</taxonomicName>
), with straighter profile, and the temones proportionally longer.
</paragraph>
</subSubSection>
<subSubSection id="C486818E9F51FE5C755D39D863438C28" pageId="0" pageNumber="1" type="etymology">
<paragraph id="91BE4D0CD112F882E7623394F1B3EC12" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="5C5B4F674E1AACA243B1D34E1B0D04B7" pageId="0" pageNumber="1">
Specific name in apposition of Madai, a Guanche (Tenerife aboriginal) word meaning
<normalizedToken id="8A64CE4688013AB93A3D60D9CBAB6448" originalValue="“deep”">&quot;deep&quot;</normalizedToken>
(
<bibRefCitation id="CF2193168F924C233BF21C00BA45019C" author="Alvarez, JA" journalOrPublisher="Centro de la Cultura Popular Canaria, Santa Cruz de Tenerife" pageId="0" pageNumber="1" refId="B3" refString="Alvarez, JA, 1991. Lenguaje de los Antiguos Islenos. Centro de la Cultura Popular Canaria, Santa Cruz de Tenerife" title="Lenguaje de los Antiguos Islenos." year="1991">
<normalizedToken id="E7546AA96ADE516859C0E4CBFE6F4821" originalValue="Álvarez">Alvarez</normalizedToken>
1991
</bibRefCitation>
), alluding to the habitat of this species.
</paragraph>
</subSubSection>
<subSubSection id="00BD9CC5EAE4CBD305919B86F762A321" pageId="0" pageNumber="1" type="habitat">
<paragraph id="83167DABB57BC54040AA6CD504B52F76" pageId="0" pageNumber="1">Habitat and distribution.</paragraph>
<paragraph id="88EB145EAFE70ADDD03E27435A6966E0" pageId="0" pageNumber="1">
<taxonomicName id="F81A71EA1DD6B812C9F6EA94839015B9" authorityName="Garcia &amp; Oromi" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Baezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baezia madai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madai">
<emphasis id="F5BC261D1AC3112684B2BB9074BB64F1" italics="true" pageId="0" pageNumber="1">Baezia madai</emphasis>
</taxonomicName>
has only been collected in Cueva de Las Tijaraferas lava tube, despite systematic biospeleological surveys conducted in other caves of the same area for many years (e.g.
<bibRefCitation id="6CAB6A4F88FF7C4B058C3BA790CAAB0A" author="Fernandez, O" journalOrPublisher="Vulcania" pageId="0" pageNumber="1" pagination="6 - 45" refId="B6" refString="Fernandez, O, Garcia, R, Dumpierrez, D, Gomez, D, Mata, M, Govantes, E, Gonzalez, AJ, Martin, M, Rodriguez, Y, 2007. Las cavidades volcanicas de El Paso, III: Sectores del Barranco de los Cardos-Barranco de El Riachuelo y de Torres (La Palma, Islas Canarias). Vulcania 8: 6 - 45" title="Las cavidades volcanicas de El Paso, III: Sectores del Barranco de los Cardos-Barranco de El Riachuelo y de Torres (La Palma, Islas Canarias)." volume="8" year="2007">
<normalizedToken id="457070222A94771FCF037A677C82C2B3" originalValue="Fernández">Fernandez</normalizedToken>
et al. 2007
</bibRefCitation>
). The cave is located in the wall of a small ravine, in a place where the potential vegetation is thermo-sclerophyllous, but partially degraded and nowadays mainly replaced by
<taxonomicName id="BF8DC25CD87AA9B0F7EF14CD933AFA31" class="Dicotyledoneae" family="Rosaceae" genus="Amygdalus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Amygdalus communis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="communis">
<emphasis id="8E63F30B809DFE3D3D16ED4C67BD93AB" italics="true" pageId="0" pageNumber="1">Amygdalus communis</emphasis>
</taxonomicName>
,
<taxonomicName id="E8C9E998B9AF8507B62DAA91EEAA16AB" class="Dicotyledoneae" family="Cactaceae" genus="Opuntia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Opuntia" order="Cactales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis id="8C604D42E049282395DBD284A16C936A" italics="true" pageId="0" pageNumber="1">Opuntia</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="424F7F41A33E30D5DF2466C42A90C45C" authorityName="Sweet" authorityYear="1818" class="Dicotyledoneae" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Euphorbia lamarckii" order="Geraniales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="lamarckii">
<emphasis id="4566F0A1A0E8B9431AE3933E52A2753D" italics="true" pageId="0" pageNumber="1">Euphorbia lamarckii</emphasis>
</taxonomicName>
and
<taxonomicName id="4B54FCE066FCFC454057267471722611" class="Dicotyledoneae" family="Polygonaceae" genus="Rumex" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rumex lunaria" order="Polygonales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="lunaria">
<emphasis id="F532899E42068FAF7587404F1AF8D0F5" italics="true" pageId="0" pageNumber="1">Rumex lunaria</emphasis>
</taxonomicName>
. In addition to its biological interest, this cave (only
<quantity id="D487C1B9B34E23F4B1D8BE6FE993031A" metricMagnitude="1" metricUnit="m" metricValue="6.3" unit="m" value="63.0">63 m</quantity>
in length) holds many archaeological remains (pottery, bones, shells, etc.) that attest to its use as home by the prehispanic inhabitants of the island, the Auaritas. Inside this cave, there are several sections with high environmental humidity, roots hanging from the ceiling and walls, and soil covered with fine sediments. These points in the cave are the most suitable for underground fauna. Besides
<taxonomicName id="F1E8DFA2A6DEA3DCF6F3AD046990447E" authorityName="Garcia &amp; Oromi" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Baezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baezia madai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madai">
<emphasis id="B0B7B7167D13D726C638F3DB4E4367A2" italics="true" pageId="0" pageNumber="1">Baezia madai</emphasis>
</taxonomicName>
sp. nov., the presence of other troglobitic species has been confirmed, such as the sandhopper
<taxonomicName id="B5532443EBDE1815811F0115B5811937" authorityName="Stock &amp; Martin" authorityYear="1988" class="Malacostraca" family="Talitridae" genus="Palmorchestia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Palmorchestia hypogaea" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hypogaea">
<emphasis id="98695D6855D92D01ADF8726EFB3B56C0" italics="true" pageId="0" pageNumber="1">Palmorchestia hypogaea</emphasis>
</taxonomicName>
Stock &amp;
<normalizedToken id="027F0989A97C6B4C286179D6BA6CB39D" originalValue="Martín">Martin</normalizedToken>
, 1988, the cockroach
<taxonomicName id="7017753C0555971779433DA669D411AB" authorityName="Izquierdo &amp; Martin" authorityYear="1999" class="Insecta" family="Blattellidae" genus="Loboptera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Loboptera teneguia" order="Blattaria" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="teneguia">
<emphasis id="1041CBDF507A87A95B8B0EED547C1F98" italics="true" pageId="0" pageNumber="1">Loboptera teneguia</emphasis>
</taxonomicName>
Izquierdo &amp;
<normalizedToken id="3E1A30B57B1628F5C417CF1B6238BD9F" originalValue="Martín">Martin</normalizedToken>
, 1999, and the ground beetles
<taxonomicName id="9F630255CABD784EA291183AE8F7BA5C" authorityName="Machado" authorityYear="1987" class="Insecta" family="Carabidae" genus="Licinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Licinopsis angustula" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="angustula">
<emphasis id="92F494F104F0D0EC1FC47D81BA334D46" italics="true" pageId="0" pageNumber="1">Licinopsis angustula</emphasis>
</taxonomicName>
Machado, 1987 and
<taxonomicName id="4B89E529E677CD08A69FC40758148F18" class="Insecta" family="Carabidae" genus="Thalassophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Thalassophilus subterraneus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis id="062153939B90B55B92DD637F3CB69FA9" italics="true" pageId="0" pageNumber="1">Thalassophilus subterraneus</emphasis>
</taxonomicName>
Machado, 1990. During the last decade, the authors have sampled the cave several times with pitfall traps and collected dead roots to remove weevil individuals from them, but without obtaining additional material beyond the only known specimen.
</paragraph>
</subSubSection>
</treatment>
</document>