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<document ID-CLB-Dataset="21020" ID-DOI="http://dx.doi.org/10.3897/zse.96.52860" ID-GBIF-Dataset="e27c18de-98bd-41e1-9354-11d49edd5d33" ID-Pensoft-Pub="1860-0743-2-577" ID-Pensoft-UUID="D4882E78D81B5FA394A6A156886835E4" ModsDocID="1860-0743-2-577" checkinTime="1599019521532" checkinUser="pensoft" docAuthor="Vinarski, Maxim V., Aksenova, Olga V. &amp; Bolotov, Ivan N." docDate="2020" docId="894E1A94D5C051EC9473A1FB446FB83E" docLanguage="en" docName="ZoosystEvol 96(2): 577-608" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.52860" docTitle="Ampullaceana relicta subsp. pinteri" docType="treatment" docVersion="6" id="D4882E78D81B5FA394A6A156886835E4" lastPageNumber="577" masterDocId="D4882E78D81B5FA394A6A156886835E4" masterDocTitle="Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)" masterLastPageNumber="608" masterPageNumber="577" pageNumber="577" updateTime="1732655424229" updateUser="ExternalLinkService">
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<mods:title id="F94A21A6AD3479A050E56120817F90FA">Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)</mods:title>
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<mods:namePart id="5765D72F70D7052B16E120B0F9633926">Vinarski, Maxim V.</mods:namePart>
<mods:affiliation id="AAD7B19CB21D610A9EAED243A9D1525D">Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; Omsk State University, 28 Adrianova Str., 644077, Omsk, Russia</mods:affiliation>
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<mods:nameIdentifier id="FE075720A62230A7EA36CFE89EB49BBE" type="email">radix.vinarski@gmail.com</mods:nameIdentifier>
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<mods:namePart id="164C945E47E76D38C16CDEA11D707EAD">Aksenova, Olga V.</mods:namePart>
<mods:affiliation id="979580C8B4A2A77028D86D3A76A302A6">Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia</mods:affiliation>
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<mods:namePart id="13D34621F79E54502425411F32016F88">Bolotov, Ivan N.</mods:namePart>
<mods:affiliation id="827BF54D380313ECD06CD8AA482F6756">N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia &amp; Northern (Arctic) Federal University, 17 Severnaya Dvina Emb., 163002, Arkhangelsk, Russia</mods:affiliation>
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<mods:title id="6F44B06A199F4C8D538A97F1AC0525C6">Zoosystematics and Evolution</mods:title>
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<mods:date id="87719C30FA92222A4A164A2FF9B26D55">2020</mods:date>
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<mods:start id="F151230AD1636D67928240CF31726C08">577</mods:start>
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<treatment id="894E1A94D5C051EC9473A1FB446FB83E" ID-GBIF-Taxon="167407617" LSID="urn:lsid:plazi:treatment:894E1A94D5C051EC9473A1FB446FB83E" httpUri="http://treatment.plazi.org/id/894E1A94D5C051EC9473A1FB446FB83E" lastPageNumber="577" pageId="0" pageNumber="577" scope_class="Gastropoda" scope_family="Lymnaeidae" scope_phylum="Mollusca">
<subSubSection id="0C8AA2917EE43FFBAC107FF8468CB224" pageId="0" pageNumber="577" type="nomenclature">
<paragraph id="F3B6ACC1E95887278F12D1F8D76F17BD" pageId="0" pageNumber="577">
16b.
<taxonomicName id="3049CAA7C4F4B4D5DEB52177AF05F39C" ID-CoL="5DGYM" LSID="894E1A94-D5C0-51EC-9473-A1FB446FB83E" authority="(Schuett, 1974)" baseAuthorityName="Schuett" baseAuthorityYear="1974" family="Lymnaeidae" genus="Ampullaceana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ampullaceana relicta subsp. pinteri" order="Hygrophila" pageId="0" pageNumber="577" rank="subSpecies" species="relicta" subSpecies="pinteri">
Ampullaceana relicta pinteri (
<normalizedToken id="AEB82B8DB1214F90B9BEFDAE2B02DF3A" originalValue="Schütt">Schuett</normalizedToken>
, 1974)
</taxonomicName>
<figureCitation id="5CEE195F5160F3F11F3E9FA2FBE561A6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells of species of the genera Ampullaceana, Kamtschaticana, Myxas, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana relicta relicta (05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW). B. A. r. pinteri, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW). C, D. Kamtschaticana kamtschatica (Russia, Kamchatka Peninsula, Valley of Geysers; RMBH). E. Myxas glutinosa (14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP). F, G. Tibetoradix hookeri, two syntypes (NHMUK). H. T. kozlovi, the holotype (ZISP). I. Racesina luteola (India, Bengal, Paksay, Pabna; NHMUK). J. R. oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI). K. R. siamensis (&quot; Thailand &quot;; ZMB). L. Orientogalba viridis, a syntype (Guam Island; NMNH). M. O. ollula (1883, Japan, Yokohama; NHMUK). N. Limnaea bowelli, a type (after Preston 1909). O. Orientogalba cf. bowelli (30.06.1924, China, Tibet, Lingka, 14,500 feet a. s. l.; NHMUK). P. Bullastra cumingiana (Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (C, D, F-H, L, M, O), 5 mm (A, B, E, J-K, N, P)." figureDoi="10.3897/zse.96.52860.figure9" httpUri="https://binary.pensoft.net/fig/448273" pageId="0" pageNumber="577">Fig. 9B.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3F61D4C445339BC7334B4C8965D3D486" pageId="0" pageNumber="577" type="reference_group">
<paragraph id="48C98B0559E7CF227C751F5D7C793FED" pageId="0" pageNumber="577">
<taxonomicName id="0919B21B8CB6F41686C1175E917387FE" family="Lymnaeidae" genus="Radix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Radix pinteri" order="Hygrophila" pageId="0" pageNumber="577" rank="species" species="pinteri">Radix pinteri</taxonomicName>
<bibRefCitation id="151E34B91186A6315C66DBC5D7E78222" author="Schuett, H" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="0" pageNumber="577" pagination="473 - 480" refId="B113" refString="Schuett, H, 1974. Zwei neue reliktaere Suesswassermollusken der Dinariden. Annalen des Naturhistorischen Museums in Wien 78: 473 - 480" title="Zwei neue reliktaere Suesswassermollusken der Dinariden." volume="78" year="1974">
<normalizedToken id="91EB4C049B17F75EAE38BC88D095065A" originalValue="Schütt">Schuett</normalizedToken>
1974
</bibRefCitation>
: 471, pl. 1.
</paragraph>
<paragraph id="64191339A205A91A5FFCCCFA1F61EDAB" pageId="0" pageNumber="577">
<taxonomicName id="A20F821CA59D1C890A75A12B6F1BB9EC" family="Lymnaeidae" genus="Radix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Radix pinteri" order="Hygrophila" pageId="0" pageNumber="577" rank="species" species="pinteri">Radix pinteri</taxonomicName>
-
<bibRefCitation id="6150C1A0AF3712A785C5894AC0A22F57" DOI="https://doi.org/10.1007/s10750-008-9555-1" author="Albrecht, C" journalOrPublisher="Hydrobiologia" pageId="0" pageNumber="577" pagination="157 - 167" refId="B6" refString="Albrecht, C, Wolff, C, Gloeer, P, Wilke, T, 2008. Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa. Hydrobiologia 615: 157 - 167, DOI: https://doi.org/10.1007/s10750-008-9555-1" title="Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa." url="https://doi.org/10.1007/s10750-008-9555-1" volume="615" year="2008">Albrecht et al. 2008</bibRefCitation>
: 160, fig. 2;
<bibRefCitation id="92DBB5D54B7FEBD26DD8AECDB9ECE981" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="577" refId="B147" refString="Welter-Schultes, FW, 2012. European Non-Marine Molluscs, a Guide for Species Identification. Planet Poster Editions, Goettingen" title="European Non-Marine Molluscs, a Guide for Species Identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 54, textfig.;
<bibRefCitation id="FCDEB1F3C79A519378BEDC961C92C7E0" author="Gloeer, P" journalOrPublisher="Conchbooks, Hackenheim" pageId="0" pageNumber="577" refId="B42" refString="Gloeer, P, 2019. The Freshwater Gastropoda of the West-Palaearctis. V. I. Fresh- and brackish waters except spring and subterranean snails. The author, 400 pp." title="The Freshwater Gastropoda of the West-Palaearctis. V. I. Fresh- and brackish waters except spring and subterranean snails. The author, 400 pp." year="2019">
<normalizedToken id="B1C044B975AB3FFB0C81B249A1CD2F2B" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 244, fig. 304.
</paragraph>
<paragraph id="C625ADA559956784571496776FB6EC95" pageId="0" pageNumber="577">
<taxonomicName id="16E50D2154EE4006DDCEDC72F6FF62D7" baseAuthorityName="Schuett" baseAuthorityYear="1974" family="Lymnaeidae" genus="Ampullaceana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ampullaceana relicta subsp. pinteri" order="Hygrophila" pageId="0" pageNumber="577" rank="subSpecies" species="relicta" subSpecies="pinteri">Ampullaceana relicta pinteri</taxonomicName>
-
<bibRefCitation id="B2158E6D304E505FA22106C2A42E110F" DOI="https://doi.org/10.1038/s41598-018-29451-1" author="Aksenova, OV" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="577" pagination="1 - 17" refId="B1" refString="Aksenova, OV, Bolotov, IN, Gofarov, MY, Kondakov, AV, Vinarski, MV, Bespalaya, YV, Kolosova, YS, Palatov, DM, Sokolova, SE, Spitsyn, VM, Tomilova, AA, Travina, OV, Vikhrev, IV, 2018a. Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World. Scientific Reports 8 (11199): 1 - 17, DOI: https://doi.org/10.1038/s41598-018-29451-1" title="Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World." url="https://doi.org/10.1038/s41598-018-29451-1" volume="8" year="2018 a">Aksenova et al. 2018a</bibRefCitation>
: 4.
</paragraph>
</subSubSection>
<subSubSection id="E6C57E930339FE0E0D175D458DA7F26B" pageId="0" pageNumber="577" type="tl">
<paragraph id="71887CCF5CBF41DF20CC441E79BD9BEB" pageId="0" pageNumber="577">TL.</paragraph>
<paragraph id="8AFB7AF364DF294285F8EE32065072F9" pageId="0" pageNumber="577">North Macedonia, Lake Prespa near village of Perovo.</paragraph>
</subSubSection>
<subSubSection id="1DFE73AED496E1E071FB34F0CA7F2DB9" pageId="0" pageNumber="577" type="types">
<paragraph id="83AB927A084F9AF34B3771DC6E6311AA" pageId="0" pageNumber="577">Types.</paragraph>
<paragraph id="39FB79CC6247C4A281E70DD146BE9ACB" pageId="0" pageNumber="577">
Holotype - SMF; paratypes were placed in NHMW and in a series of private collections in Germany and Hungary (
<bibRefCitation id="2D4CC7B003F2DC0C37DA8DAA9AC6BA65" author="Schuett, H" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="0" pageNumber="577" pagination="473 - 480" refId="B113" refString="Schuett, H, 1974. Zwei neue reliktaere Suesswassermollusken der Dinariden. Annalen des Naturhistorischen Museums in Wien 78: 473 - 480" title="Zwei neue reliktaere Suesswassermollusken der Dinariden." volume="78" year="1974">
<normalizedToken id="B98FB9183F49C4D701543C023A6EAA08" originalValue="Schütt">Schuett</normalizedToken>
1974
</bibRefCitation>
); currently, specimens from these private collections are kept in the Hungarian Natural History Museum (Budapest) and in the Natural History Museum of Bern (Switzerland). We examined the paratypes kept in NHMW and in the Budapest museum (see Fig.
<figureCitation id="3358221EF5AE5DED7BB4614F8E92BC30" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells of species of the genera Ampullaceana, Kamtschaticana, Myxas, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana relicta relicta (05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW). B. A. r. pinteri, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW). C, D. Kamtschaticana kamtschatica (Russia, Kamchatka Peninsula, Valley of Geysers; RMBH). E. Myxas glutinosa (14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP). F, G. Tibetoradix hookeri, two syntypes (NHMUK). H. T. kozlovi, the holotype (ZISP). I. Racesina luteola (India, Bengal, Paksay, Pabna; NHMUK). J. R. oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI). K. R. siamensis (&quot; Thailand &quot;; ZMB). L. Orientogalba viridis, a syntype (Guam Island; NMNH). M. O. ollula (1883, Japan, Yokohama; NHMUK). N. Limnaea bowelli, a type (after Preston 1909). O. Orientogalba cf. bowelli (30.06.1924, China, Tibet, Lingka, 14,500 feet a. s. l.; NHMUK). P. Bullastra cumingiana (Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (C, D, F-H, L, M, O), 5 mm (A, B, E, J-K, N, P)." figureDoi="10.3897/zse.96.52860.figure9" httpUri="https://binary.pensoft.net/fig/448273" pageId="0" pageNumber="577">9B</figureCitation>
).
</paragraph>
<caption id="153B8A7B7D08748FE52124FC4E7D2150" doi="10.3897/zse.96.52860.figure9" httpUri="https://binary.pensoft.net/fig/448273" pageId="0" pageNumber="577" start="Figure 9" startId="F9">
<paragraph id="E45225AEAC6D0D000E8D9F61C8C66774" pageId="0" pageNumber="577">
<emphasis id="D49CB457F56672DCD7E24654BC9DAC5C" bold="true" pageId="0" pageNumber="577">Figure 9.</emphasis>
Shells of species of the genera
<taxonomicName id="C086356ECC87AB4D8E77EEB9BA6BE2E7" authorityName="Servain" authorityYear="1882" class="Gastropoda" family="Lymnaeidae" genus="Ampullaceana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ampullaceana" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="F13CACA443BDA2433F9A5A10FA09A13C" italics="true" pageId="0" pageNumber="577">Ampullaceana</emphasis>
</taxonomicName>
,
<taxonomicName id="3EE59F6D77B787256BDC6C72FF3821D9" authorityName="Kruglov &amp; Starobogatov" authorityYear="1984" class="Gastropoda" family="Lymnaeidae" genus="Kamtschaticana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kamtschaticana" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="F0A3877787D02C9959FA0C0702C3EF63" italics="true" pageId="0" pageNumber="577">Kamtschaticana</emphasis>
</taxonomicName>
,
<taxonomicName id="6178AA603129995B423B67656F4B73D8" authorityName="G. B. Sowerby I" authorityYear="1822" class="Gastropoda" family="Lymnaeidae" genus="Myxas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myxas" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="BE068CFA2755E21173E72A4FCAF2F2A6" italics="true" pageId="0" pageNumber="577">Myxas</emphasis>
</taxonomicName>
,
<taxonomicName id="32C1CE4C3F2F719922C294E2AE858BA3" authorityName="Bolotov, Vinarski &amp; Aksenova" authorityYear="2018" class="Gastropoda" family="Lymnaeidae" genus="Tibetoradix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tibetoradix" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="0CB457B67747F7754760953988B97804" italics="true" pageId="0" pageNumber="577">Tibetoradix</emphasis>
</taxonomicName>
,
<taxonomicName id="BA34FF6DE9201A271B565A616B46FEC5" authorityName="Vinarski &amp; Bolotov" authorityYear="2018" class="Gastropoda" family="Lymnaeidae" genus="Racesina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Racesina" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="F8AA657943ACA83F71BC14DD08B49E83" italics="true" pageId="0" pageNumber="577">Racesina</emphasis>
</taxonomicName>
,
<taxonomicName id="09B87DDD98BA4716A5F2CBD42C69F0AD" authorityName="Kruglov &amp; Starobogatov" authorityYear="1985" class="Gastropoda" family="Lymnaeidae" genus="Orientogalba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientogalba" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="3DBA81B63C6BECA885478E5EB50EA9D8" italics="true" pageId="0" pageNumber="577">Orientogalba</emphasis>
</taxonomicName>
and
<taxonomicName id="B9ACE6B68D1AEE7DEED81BE5A40DEC1D" authorityName="Bergh" authorityYear="1901" class="Gastropoda" family="Lymnaeidae" genus="Bullastra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bullastra" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis id="54B6ECD588FED82C9428A7F40C901327" italics="true" pageId="0" pageNumber="577">Bullastra</emphasis>
</taxonomicName>
.
<emphasis id="ECD94BC2CE05F60B36037EFFBF10383E" bold="true" pageId="0" pageNumber="577">A.</emphasis>
<taxonomicName id="037D0E5A13EB24C23062BBE21CDA3556" authority="relicta" authorityName="relicta (Polinski" authorityYear="1929" class="Gastropoda" family="Lymnaeidae" genus="Ampullaceana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ampullaceana relicta subsp. relicta" pageId="0" pageNumber="577" phylum="Mollusca" rank="subSpecies" species="relicta" subSpecies="relicta">
<emphasis id="B64E5FC5851C83046BCE0B27C35BAB0E" italics="true" pageId="0" pageNumber="577">Ampullaceana relicta relicta</emphasis>
</taxonomicName>
(05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW).
<emphasis id="53AB75010E98FB28D160927FDAD6121D" bold="true" pageId="0" pageNumber="577">B.</emphasis>
<taxonomicName id="1E3E00561BB10480FA16CD32EF94AEB8" lsidName="A. r. subsp. pinteri" pageId="0" pageNumber="577" rank="subSpecies" species="r." subSpecies="pinteri">
<emphasis id="AAA3099C247C945E4178B9ED35E15EE8" italics="true" pageId="0" pageNumber="577">A. r. pinteri</emphasis>
</taxonomicName>
, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW).
<emphasis id="34C848F8D87D42F68E8C24CBE4FAFA9E" bold="true" pageId="0" pageNumber="577">C, D.</emphasis>
<taxonomicName id="1E93EA7F1053EA52E223C05BC146069A" baseAuthorityName="Middendorff" baseAuthorityYear="1850" class="Gastropoda" family="Lymnaeidae" genus="Kamtschaticana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kamtschaticana kamtschatica" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="kamtschatica">
<emphasis id="C6EB0635806FBE61941694BE609752F9" italics="true" pageId="0" pageNumber="577">Kamtschaticana kamtschatica</emphasis>
</taxonomicName>
(Russia, Kamchatka Peninsula, Valley of Geysers; RMBH).
<emphasis id="5AF7AFB86B01C0AA8EC21684BB1DD972" bold="true" pageId="0" pageNumber="577">E.</emphasis>
<taxonomicName id="CB787E186F28058F71D6EC72E39AC6B7" baseAuthorityName="O. F. Mueller" baseAuthorityYear="1774" class="Gastropoda" family="Lymnaeidae" genus="Myxas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myxas glutinosa" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="glutinosa">
<emphasis id="BB55E7CF32AF7B71E1459B029D40D101" italics="true" pageId="0" pageNumber="577">Myxas glutinosa</emphasis>
</taxonomicName>
(14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP).
<emphasis id="9483A4C86B26CA39ECD75E0827573A6D" bold="true" pageId="0" pageNumber="577">F, G.</emphasis>
<taxonomicName id="9C91423DC13A7DF9D54FF70AB31F0E85" baseAuthorityName="Reeve" baseAuthorityYear="1850" class="Gastropoda" family="Lymnaeidae" genus="Tibetoradix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tibetoradix hookeri" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="hookeri">
<emphasis id="EC7FBA89B9EE7341A8A6E1CE17B7636C" italics="true" pageId="0" pageNumber="577">Tibetoradix hookeri</emphasis>
</taxonomicName>
, two syntypes (NHMUK).
<emphasis id="0D4EFC494672EAF47DFBFD631717FB03" bold="true" pageId="0" pageNumber="577">H.</emphasis>
<taxonomicName id="124AF6DABF109D4ADBD0385B90BAF191" lsidName="T. kozlovi" pageId="0" pageNumber="577" rank="species" species="kozlovi">
<emphasis id="EADDD749E32614B899C72713C0A19EE0" italics="true" pageId="0" pageNumber="577">T. kozlovi</emphasis>
</taxonomicName>
, the holotype (ZISP).
<emphasis id="06FFF18B6E011105801F8153AC316504" bold="true" pageId="0" pageNumber="577">I.</emphasis>
<taxonomicName id="BAAED542830477145F5DD6EFCB06F159" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Lymnaeidae" genus="Racesina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Racesina luteola" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="luteola">
<emphasis id="1B7F467131772E2A5C27E899A21F4898" italics="true" pageId="0" pageNumber="577">Racesina luteola</emphasis>
</taxonomicName>
(India, Bengal, Paksay, Pabna; NHMUK).
<emphasis id="38199449D5D6D035C642B02114A2B657" bold="true" pageId="0" pageNumber="577">J.</emphasis>
<taxonomicName id="1361BA530456EC11D129812272951791" lsidName="R. oxiana" pageId="0" pageNumber="577" rank="species" species="oxiana">
<emphasis id="0C617AFD87991ADF31142F8B8335735F" italics="true" pageId="0" pageNumber="577">R. oxiana</emphasis>
</taxonomicName>
(04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI).
<emphasis id="91EFC531EE1B8A88735F1AFEB4496E09" bold="true" pageId="0" pageNumber="577">K.</emphasis>
<taxonomicName id="A7EF93837A1B02F09C58E1EB4A760CA1" lsidName="R. siamensis" pageId="0" pageNumber="577" rank="species" species="siamensis">
<emphasis id="9A3A291BB514F7B865DD2347C3441CAE" italics="true" pageId="0" pageNumber="577">R. siamensis</emphasis>
</taxonomicName>
(
<normalizedToken id="A6AC4166CBA595FA7C338504063B9B07" originalValue="“Thailand”">&quot;Thailand&quot;</normalizedToken>
; ZMB).
<emphasis id="54EB1F137E11F95A070B324A5D3FA16D" bold="true" pageId="0" pageNumber="577">L.</emphasis>
<taxonomicName id="DE4664A1DC75FDDFD48671EEC80CA1B9" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1832" class="Gastropoda" family="Lymnaeidae" genus="Orientogalba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientogalba viridis" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="viridis">
<emphasis id="A0662DD5A25E4BC5FE9FE02E715862F7" italics="true" pageId="0" pageNumber="577">Orientogalba viridis</emphasis>
</taxonomicName>
, a syntype (Guam Island; NMNH).
<emphasis id="80B0F33DF9029CE1C8F79A7C477E1B52" bold="true" pageId="0" pageNumber="577">M.</emphasis>
<taxonomicName id="196FD55A7AC5EF29FFE032FD998A14E7" lsidName="O. ollula" pageId="0" pageNumber="577" rank="species" species="ollula">
<emphasis id="99C02E8DCF49C4FD542A9452DF419569" italics="true" pageId="0" pageNumber="577">O. ollula</emphasis>
</taxonomicName>
(1883, Japan, Yokohama; NHMUK).
<emphasis id="7BF9D0E4FD826B2A12C6A4A0D4E8CE96" bold="true" pageId="0" pageNumber="577">N.</emphasis>
<taxonomicName id="B8EF56E145AEB2874EB2D98716886CF4" authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Lymnaeidae" genus="Limnaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnaea bowelli" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="bowelli">
<emphasis id="EF8DD078B76EA46BD9B5715294E0CC9B" italics="true" pageId="0" pageNumber="577">Limnaea bowelli</emphasis>
</taxonomicName>
, a type (after
<bibRefCitation id="464FE89CAFB8B2FC9201FA96F30A3484" author="Preston, HB" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="577" pagination="115 - 116" publicationUrl="https://biodiversitylibrary.org/page/11129971" refId="B104" refString="Preston, HB, 1909. Report on a small collection of freshwater Mollusca (Limnaea and Pisidium) from Tibet. Records of the Indian Museum 3: 115 - 116, https://biodiversitylibrary.org/page/11129971" title="Report on a small collection of freshwater Mollusca (Limnaea and Pisidium) from Tibet." url="https://biodiversitylibrary.org/page/11129971" volume="3" year="1909">Preston 1909</bibRefCitation>
).
<emphasis id="B1C8294C78783880E15397B2941E72F0" bold="true" pageId="0" pageNumber="577">O.</emphasis>
<taxonomicName id="015B9B3BFC2D1C083CB1E49A5295AA1B" baseAuthorityName="Preston" baseAuthorityYear="1909" class="Gastropoda" family="Lymnaeidae" genus="Orientogalba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientogalba bowelli" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="bowelli">Orientogalba cf. bowelli</taxonomicName>
(30.06.1924, China, Tibet, Lingka, 14,500 feet a.s.l.; NHMUK).
<emphasis id="1A7853CEC68C4F5E0070BD83525522AD" bold="true" pageId="0" pageNumber="577">P.</emphasis>
<taxonomicName id="933D192E0D82ACDAE6A6812763B9E1E8" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1845" class="Gastropoda" family="Lymnaeidae" genus="Bullastra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bullastra cumingiana" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="cumingiana">
<emphasis id="29004909E31AE0BA6A822A7145BD2FC0" italics="true" pageId="0" pageNumber="577">Bullastra cumingiana</emphasis>
</taxonomicName>
(Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (
<emphasis id="8BD6E3FDD31BDE49A7CBF737A8458709" bold="true" pageId="0" pageNumber="577">C, D, F-H, L, M, O</emphasis>
), 5 mm (
<emphasis id="99B78FE90EC8386A78236CC1A354BAC6" bold="true" pageId="0" pageNumber="577">A, B, E, J-K, N, P</emphasis>
).
</paragraph>
</caption>
<paragraph id="7411F59C747EDAC14EEB148286BC9DDC" pageId="0" pageNumber="577">
This subspecies has been characterised both morphologically and genetically in a series of papers (
<bibRefCitation id="CE753DD5867A5E7C1EF6036C4E001C08" author="Schuett, H" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="0" pageNumber="577" pagination="473 - 480" refId="B113" refString="Schuett, H, 1974. Zwei neue reliktaere Suesswassermollusken der Dinariden. Annalen des Naturhistorischen Museums in Wien 78: 473 - 480" title="Zwei neue reliktaere Suesswassermollusken der Dinariden." volume="78" year="1974">
<normalizedToken id="C9D36025B2664A1E8C16242C27A0AB2F" originalValue="Schütt">Schuett</normalizedToken>
1974
</bibRefCitation>
;
<bibRefCitation id="02E2F17F2D009443E07D58BBB06456D9" DOI="https://doi.org/10.1007/s10750-008-9555-1" author="Albrecht, C" journalOrPublisher="Hydrobiologia" pageId="0" pageNumber="577" pagination="157 - 167" refId="B6" refString="Albrecht, C, Wolff, C, Gloeer, P, Wilke, T, 2008. Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa. Hydrobiologia 615: 157 - 167, DOI: https://doi.org/10.1007/s10750-008-9555-1" title="Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa." url="https://doi.org/10.1007/s10750-008-9555-1" volume="615" year="2008">Albrecht et al. 2008</bibRefCitation>
;
<bibRefCitation id="AF734FF94ADF36A19CA0B6D93DCF7CB5" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="577" refId="B147" refString="Welter-Schultes, FW, 2012. European Non-Marine Molluscs, a Guide for Species Identification. Planet Poster Editions, Goettingen" title="European Non-Marine Molluscs, a Guide for Species Identification." year="2012">Welter-Schultes 2012</bibRefCitation>
). Most authors regard this taxon as a separate species. For example,
<bibRefCitation id="F74682AD8EAA7B21CB439D3183926F30" DOI="https://doi.org/10.1007/s10750-008-9555-1" author="Albrecht, C" journalOrPublisher="Hydrobiologia" pageId="0" pageNumber="577" pagination="157 - 167" refId="B6" refString="Albrecht, C, Wolff, C, Gloeer, P, Wilke, T, 2008. Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa. Hydrobiologia 615: 157 - 167, DOI: https://doi.org/10.1007/s10750-008-9555-1" title="Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa." url="https://doi.org/10.1007/s10750-008-9555-1" volume="615" year="2008">Albrecht et al. (2008)</bibRefCitation>
considered
<taxonomicName id="8686FD8CF97699547E0A82119E2BFF16" lsidName="A. r. subsp. relicta" pageId="0" pageNumber="577" rank="subSpecies" species="r." subSpecies="relicta">
<emphasis id="FD5E2A99EA17A13304180F3669C5E07E" italics="true" pageId="0" pageNumber="577">A. r. relicta</emphasis>
</taxonomicName>
and
<taxonomicName id="D2C09B5D02ABC568267707A5F166303D" lsidName="A. r. subsp. pinteri" pageId="0" pageNumber="577" rank="subSpecies" species="r." subSpecies="pinteri">
<emphasis id="0CF91C9C2E67E23FF5A491017A7BED5B" italics="true" pageId="0" pageNumber="577">A. r. pinteri</emphasis>
</taxonomicName>
as two sister species from two
<normalizedToken id="B5216BDBAC4A050D9CB3122EE50F679C" originalValue="‘sister’">'sister'</normalizedToken>
lakes, Ohrid and Prespa (but see
<bibRefCitation id="AAF93C8D18C6BD2831E1DC5C0C199961" DOI="https://doi.org/10.1038/s41598-018-29451-1" author="Aksenova, OV" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="577" pagination="1 - 17" refId="B1" refString="Aksenova, OV, Bolotov, IN, Gofarov, MY, Kondakov, AV, Vinarski, MV, Bespalaya, YV, Kolosova, YS, Palatov, DM, Sokolova, SE, Spitsyn, VM, Tomilova, AA, Travina, OV, Vikhrev, IV, 2018a. Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World. Scientific Reports 8 (11199): 1 - 17, DOI: https://doi.org/10.1038/s41598-018-29451-1" title="Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World." url="https://doi.org/10.1038/s41598-018-29451-1" volume="8" year="2018 a">Aksenova et al. 2018a</bibRefCitation>
). According to the anatomical data presented in
<bibRefCitation id="6226DD610644A7F863211DEA567FA8E0" DOI="https://doi.org/10.1007/s10750-008-9555-1" author="Albrecht, C" journalOrPublisher="Hydrobiologia" pageId="0" pageNumber="577" pagination="157 - 167" refId="B6" refString="Albrecht, C, Wolff, C, Gloeer, P, Wilke, T, 2008. Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa. Hydrobiologia 615: 157 - 167, DOI: https://doi.org/10.1007/s10750-008-9555-1" title="Concurrent evolution of ancient sister lakes and sister species: the freshwater gastropod genus Radix in lakes Ohrid and Prespa." url="https://doi.org/10.1007/s10750-008-9555-1" volume="615" year="2008">Albrecht et al. (2008)</bibRefCitation>
, the morphology of the copulatory apparatus in
<taxonomicName id="FADCD5B616E08E0829D0EFDFA4660FA9" lsidName="A. r. subsp. pinteri" pageId="0" pageNumber="577" rank="subSpecies" species="r." subSpecies="pinteri">
<emphasis id="1298FC6E013CDF72AF5B3CBB61901243" italics="true" pageId="0" pageNumber="577">A. r. pinteri</emphasis>
</taxonomicName>
is virtually identical with that of the nominative subspecies.
</paragraph>
<paragraph id="D335F50D54FACDCEFDF3B791D4881607" pageId="0" pageNumber="577">
<taxonomicName id="368FA63B7F4C80072B0F6D7DFA3CF283" lsidName="A. r. subsp. pinteri" pageId="0" pageNumber="577" rank="subSpecies" species="r." subSpecies="pinteri">
<emphasis id="C776405B13B54C698EE3551687BA84DE" italics="true" pageId="0" pageNumber="577">A. r. pinteri</emphasis>
</taxonomicName>
is endemic to Prespa Lake (Albania, Greece and Macedonia).
</paragraph>
</subSubSection>
</treatment>
</document>
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