238 lines
20 KiB
XML
238 lines
20 KiB
XML
<document ID-DOI="10.11646/zootaxa.3683.4.8" ID-GBIF-Dataset="790ed03f-43eb-409c-8fad-3bbbffa0910f" ID-ISSN="1175-5326" ID-Zenodo-Dep="247516" ID-ZooBank="A7A91D16-85AD-40DB-9627-FD02101289A4" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1460645395031" checkinUser="plazi" docAuthor="Rakotonirina, Jean Claude & Fisher, Brian L." docDate="2013" docId="8931A65BFFB84242FF54440EFB9CFAD4" docLanguage="en" docName="zt03683p485.pdf" docOrigin="Zootaxa 3683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachycondyla agnivo Rakotonirina and Fisher, sp. n." docType="treatment" docVersion="9" lastPageNumber="459" masterDocId="7508DE23FFB3424EFFC34533FFA5FF9A" masterDocTitle="Revision of the Pachycondyla sikorae species-group (Hymenoptera: Formicidae) in Madagascar" masterLastPageNumber="485" masterPageNumber="447" pageNumber="458" updateTime="1683651725442" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Pachycondyla sikorae species-group (Hymenoptera: Formicidae) in Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rakotonirina, Jean Claude</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-07-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>3683</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>447</mods:start>
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<mods:end>485</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.3683.4.8</mods:identifier>
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<mods:identifier type="GBIF-Dataset">790ed03f-43eb-409c-8fad-3bbbffa0910f</mods:identifier>
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||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">247516</mods:identifier>
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<mods:identifier type="ZooBank">A7A91D16-85AD-40DB-9627-FD02101289A4</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6162579" ID-GBIF-Taxon="119571904" ID-Zenodo-Dep="6162579" LSID="urn:lsid:plazi:treatment:8931A65BFFB84242FF54440EFB9CFAD4" httpUri="http://treatment.plazi.org/id/8931A65BFFB84242FF54440EFB9CFAD4" lastPageId="12" lastPageNumber="459" pageId="11" pageNumber="458">
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||
<subSubSection box="[151,803,317,343]" pageId="11" pageNumber="458" type="nomenclature">
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||
<paragraph blockId="11.[151,803,317,375]" box="[151,803,317,343]" pageId="11" pageNumber="458">
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<heading bold="true" box="[151,803,317,343]" fontSize="11" level="1" pageId="11" pageNumber="458" reason="1">
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<emphasis bold="true" box="[151,803,317,343]" pageId="11" pageNumber="458">
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<taxonomicName authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[151,725,317,343]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="458" phylum="Arthropoda" rank="species" species="agnivo" status="sp. n.">
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<emphasis bold="true" box="[151,409,317,343]" italics="true" pageId="11" pageNumber="458">Pachycondyla agnivo</emphasis>
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Rakotonirina and Fisher
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</taxonomicName>
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,
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<taxonomicNameLabel box="[737,803,317,343]" pageId="11" pageNumber="458" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,456,351,375]" pageId="11" pageNumber="458" type="description">
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<paragraph blockId="11.[151,803,317,375]" box="[151,456,351,375]" pageId="11" pageNumber="458">
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(
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<figureCitation box="[158,281,351,375]" captionStart="FIGURES 16 – 17" captionStartId="8.[151,264,1509,1532]" captionTargetBox="[151,1436,889,1470]" captionTargetId="figure@8.[151,1436,889,1480]" captionTargetPageId="8" captionText="FIGURES 16 – 17. First gastral segment in dorsal view; Fig. 16: anterior face forming shallow cavity (CASENT 0317589: holotype); Fig. 17: anterior face without shallow cavity (CASENT 0247289)." httpUri="https://zenodo.org/record/247525/files/figure.png" pageId="11" pageNumber="458">Figures 16</figureCitation>
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,
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<figureCitation box="[293,408,351,375]" captionStart="FIGURES 28 – 30" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[151,1433,679,1900]" captionTargetId="figure@11.[151,1436,679,1917]" captionTargetPageId="11" captionText="FIGURES 28 – 30. Worker caste of Pachycondyla agnivo: lateral and dorsal view of body and full-face view of head, holotype CASENT 0317589." httpUri="https://zenodo.org/record/247531/files/figure.png" pageId="11" pageNumber="458">28, 29, 30</figureCitation>
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,
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<figureCitation box="[419,448,351,375]" captionStart="FIGURES 70 – 75" captionStartId="35.[151,264,1874,1897]" captionTargetBox="[174,1415,210,1820]" captionTargetId="figure@35.[161,1425,195,1853]" captionTargetPageId="35" captionText="FIGURES 70 – 75. Distribution maps of the Pachycondyla sikorae species-group in Madagascar. Fig. 70: P. agnivo; Fig. 71: P. antsiraka; Fig. 72: P. daraina; Fig. 73: P. g o ro g o t a; Fig. 74: P. haratsingy (circle) and P. i v o l o (triangle); Fig. 75: P. maeva." httpUri="https://zenodo.org/record/247545/files/figure.png" pageId="11" pageNumber="458">
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<number box="[419,448,351,375]" pageId="11" pageNumber="458" value="70.0">70</number>
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="458" type="materials_examined">
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<paragraph blockId="11.[151,1436,424,483]" pageId="11" pageNumber="458">
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<materialsCitation ID-GBIF-Occurrence="3744209312" collectingDate="1998-12-07" collectingDateMax="1998-12-10" collectingDateMin="1998-12-07" collectionCode="CASC" collectorName="H. J. Ratsirarson" country="Madagascar" elevation="530" latitude="-18.695" location="FC Andriantantely" longLatPrecision="55" longitude="48.8133" pageId="11" pageNumber="458" specimenCode="HJR122, CASENT0317589" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="holotype">
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<emphasis box="[151,321,428,450]" italics="true" pageId="11" pageNumber="458">
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<typeStatus box="[151,244,428,450]" pageId="11" pageNumber="458">Holotype</typeStatus>
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<specimenCount box="[249,321,428,450]" pageId="11" pageNumber="458" type="worker">worker</specimenCount>
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</emphasis>
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:
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<collectingCountry box="[337,461,428,450]" name="Madagascar" pageId="11" pageNumber="458">Madagascar</collectingCountry>
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,
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<collectingRegion box="[471,583,428,450]" country="Madagascar" name="Toamasina" pageId="11" pageNumber="458">Toamasina</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:8931A65BFFB84242FF54440EFB9CFAD4:04474196FFB84245FD93449EFCABFE58" box="[592,782,428,450]" country="Madagascar" latitude="-18.695" longLatPrecision="55" longitude="48.8133" name="FC Andriantantely" pageId="11" pageNumber="458" stateProvince="Toamasina">FC Andriantantely</location>
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,
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<geoCoordinate box="[791,871,428,450]" degrees="18.695" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="55" value="-18.695">
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-
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<number box="[798,871,428,450]" pageId="11" pageNumber="458" value="18.695">18.695</number>
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</geoCoordinate>
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,
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<number box="[882,967,429,450]" pageId="11" pageNumber="458" value="48.8133">
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<geoCoordinate box="[882,967,429,450]" degrees="48.8133" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="5" value="48.8133">48.8133</geoCoordinate>
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</number>
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,
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<quantity box="[976,1042,428,450]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="458" unit="m" value="530.0">
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<elevation box="[976,1042,428,450]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="458" unit="m" value="530.0">530 m</elevation>
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</quantity>
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, sifted litter, rainforest,
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<date box="[1282,1436,429,450]" pageId="11" pageNumber="458" value="1998-12-07" valueMax="1998-12-10" valueMin="1998-12-07">
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<collectingDate box="[1282,1436,429,450]" pageId="11" pageNumber="458" value="1998-12-07" valueMax="1998-12-10" valueMin="1998-12-07">
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<number box="[1282,1332,429,450]" pageId="11" pageNumber="458" value="8.5" valueMax="10.0" valueMin="7.0">7–10</number>
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Dec
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<number box="[1385,1436,429,450]" pageId="11" pageNumber="458" value="1998.0">1998</number>
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</collectingDate>
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</date>
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(
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<collectorName box="[207,373,461,483]" pageId="11" pageNumber="458">H.J. Ratsirarson</collectorName>
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), collection code:
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<specimenCode box="[558,644,461,482]" pageId="11" pageNumber="458">
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HJR
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<number box="[606,644,461,482]" pageId="11" pageNumber="458" value="122.0">122</number>
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</specimenCode>
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, specimen code:
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<specimenCode box="[817,1004,461,483]" pageId="11" pageNumber="458">CASENT0317589</specimenCode>
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(
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<collectionCode box="[1017,1089,461,483]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="11" pageNumber="458">CASC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="458" type="diagnosis">
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<paragraph blockId="11.[151,1436,527,623]" pageId="11" pageNumber="458">
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<emphasis bold="true" box="[151,423,527,552]" pageId="11" pageNumber="458">WORKER. Diagnosis:</emphasis>
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Anteromedian margin of clypeus broadly convex; posterolateral margins of propodeum and petiolar node smooth, without sharp teeth or tubercles; anterior surface of first gastral segment concave, forming a shallow impression to lodge the posterior surface of petiole.
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</paragraph>
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</subSubSection>
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<caption httpUri="https://zenodo.org/record/247531/files/figure.png" pageId="11" pageNumber="458" targetBox="[151,1433,679,1900]" targetPageId="11">
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<paragraph blockId="11.[151,1436,1938,1991]" pageId="11" pageNumber="458">
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<emphasis bold="true" box="[151,340,1938,1961]" pageId="11" pageNumber="458">FIGURES 28–30.</emphasis>
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Worker caste of
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<taxonomicName box="[513,732,1939,1961]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="458" phylum="Arthropoda" rank="species" species="agnivo">
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<emphasis box="[513,732,1939,1961]" italics="true" pageId="11" pageNumber="458">Pachycondyla agnivo</emphasis>
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</taxonomicName>
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: lateral and dorsal view of body and full-face view of head, holotype CASENT0317589.
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</paragraph>
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</caption>
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<subSubSection pageId="12" pageNumber="459" type="description">
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">
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<emphasis bold="true" box="[199,557,151,176]" pageId="12" pageNumber="459">
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Measurements (
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<specimenCount box="[393,545,151,176]" pageId="12" pageNumber="459" type="generic">
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<number box="[393,408,151,176]" pageId="12" pageNumber="459" value="3.0">3</number>
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specimens
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</specimenCount>
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):
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</emphasis>
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HW:
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<number box="[631,746,151,175]" pageId="12" pageNumber="459" value="1.145" valueMax="1.15" valueMin="1.14">1.14–1.15</number>
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, HL:
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<number box="[815,931,151,175]" pageId="12" pageNumber="459" value="1.38" valueMax="1.4" valueMin="1.36">1.36–1.40</number>
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, CI:
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<number box="[991,1064,151,175]" pageId="12" pageNumber="459" value="83.0" valueMax="84.0" valueMin="82.0">82–84</number>
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, SL:
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<number box="[1128,1243,151,175]" pageId="12" pageNumber="459" value="1.02" valueMax="1.03" valueMin="1.01">1.01–1.03</number>
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, SI:
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<number box="[1300,1372,151,175]" pageId="12" pageNumber="459" value="89.0" valueMax="90.0" valueMin="88.0">88–90</number>
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, PW:
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<number box="[151,265,185,209]" pageId="12" pageNumber="459" value="1.02" valueMax="1.03" valueMin="1.01">1.01–1.03</number>
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, WL:
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<number box="[335,450,185,209]" pageId="12" pageNumber="459" value="2.0" valueMax="2.04" valueMin="1.96">1.96–2.04</number>
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, NH:
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<number box="[516,631,185,209]" pageId="12" pageNumber="459" value="0.88" valueMax="0.89" valueMin="0.87">0.87–0.89</number>
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, NL:
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<number box="[694,809,185,209]" pageId="12" pageNumber="459" value="0.655" valueMax="0.68" valueMin="0.63">0.63–0.68</number>
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, NW:
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<number box="[881,996,185,209]" pageId="12" pageNumber="459" value="0.95" valueMax="0.96" valueMin="0.94">0.94–0.96</number>
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, DNI:
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<number box="[1071,1172,185,209]" pageId="12" pageNumber="459" value="146.5" valueMax="150.0" valueMin="143.0">143–150</number>
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, LNI:
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<number box="[1245,1346,185,209]" pageId="12" pageNumber="459" value="133.5" valueMax="138.0" valueMin="129.0">129–138</number>
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.
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</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">
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<emphasis bold="true" box="[199,347,218,243]" pageId="12" pageNumber="459">Description:</emphasis>
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In full face-view, head broader posteriorly; sides slightly convex; posterior margin feebly concave. Dorsum of head reticulate-punctate anteriorly and densely punctate behind from about the posterior third. Eyes large, maximum diameter nearly the same as the greatest width of antennal scape. Scape barely reaching posterior cephalic margin. Anteromedian clypeal margin convex. Mandibles sparsely punctate, the space between punctures smooth and shiny; masticatory margin armed with
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<number box="[836,864,362,386]" pageId="12" pageNumber="459" value="12.0">12</number>
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teeth or denticles.
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</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">With mesosoma in dorsal view, metanotal groove obsolete; in profile, mesopleural suture absent or indistinct; posterior magin of propodeum without a series of sharp teeth or tubercles; junction of lateral propodeal surface and declivity angulate, angles fairly emarginate. Dorsum of mesosoma with punctate sculpture. Petiole nodiform, anterodorsal angle faintly overhanging anterior margin in lateral view; subpetiolar process consisting of convex anterior face and broadly subquadrate posterior margin; in dorsal view node slightly broader than long, lateral border roughly convex and converging in a rounded line to the more or less straight anterior face; posterior margin medially broadly convex; dorsum of node sparsely punctate.</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">Anterior face of first gastral segment with shallow cavity, which is deeper near the insertion of petiole and becomes superficial towards the anterodorsal angle. With gaster in ventral view, anterior half of first gastral sternite with impressions on both sides of helcium; in profile anteroventral portion rounded and anteriorly projecting as hooklike process (the prora). Dorsum of first two gastral tergites smooth between piligerous punctures.</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">Dorsum of head and body covered with erect hairs; pubescence quite abundant on head dorsum and scarce or absent on the rest of body dorsum. Body color black with bluish reflection; appendages and tip of gaster brown.</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" box="[199,405,866,891]" pageId="12" pageNumber="459">
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<emphasis bold="true" box="[199,286,866,891]" pageId="12" pageNumber="459">Queen:</emphasis>
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unknown.
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</paragraph>
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</subSubSection>
|
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<subSubSection pageId="12" pageNumber="459" type="discussion">
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">
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<emphasis bold="true" box="[199,334,902,927]" pageId="12" pageNumber="459">Discussion:</emphasis>
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<taxonomicName box="[345,591,902,927]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="459" phylum="Arthropoda" rank="species" species="agnivo">
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<emphasis box="[345,591,902,927]" italics="true" pageId="12" pageNumber="459">Pachycondyla agnivo</emphasis>
|
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</taxonomicName>
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can be recognized easily by the medially convex anterior margin of its clypeus and the presence of a shallow impression on the anterior face of its first gastral tergite. No striking morphological variation is observed in this species despite the great distance between collection localities in the south and the center of
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<collectingCountry box="[413,551,1010,1034]" name="Madagascar" pageId="12" pageNumber="459">Madagascar</collectingCountry>
|
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.
|
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</paragraph>
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">The shallow impression on the anterior face of the first gastral segment matches the shape of the posterior section of the petiolar node, suggesting this hollow plays a role in lodging the posterior face of the node.</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="12" pageNumber="459" type="distribution">
|
||
<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">
|
||
<emphasis bold="true" box="[199,499,1118,1143]" pageId="12" pageNumber="459">Distribution and biology:</emphasis>
|
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<taxonomicName box="[506,747,1118,1142]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="459" phylum="Arthropoda" rank="species" species="agnivo">
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||
<emphasis box="[506,747,1118,1142]" italics="true" pageId="12" pageNumber="459">Pachycondyla agnivo</emphasis>
|
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</taxonomicName>
|
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is an endemic species found in mid-elevation areas of eastern
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<collectingCountry box="[151,289,1154,1178]" name="Madagascar" pageId="12" pageNumber="459">Madagascar</collectingCountry>
|
||
. In addition to the
|
||
<typeStatus box="[496,544,1155,1179]" pageId="12" pageNumber="459">type</typeStatus>
|
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locality, the species occurs in the PN Mantadia in the central east and in the PN Andohahela in the southernmost portion of the island. It was collected by sampling leaf litter in the central east of
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<collectingCountry box="[151,286,1226,1250]" name="Madagascar" pageId="12" pageNumber="459">Madagascar</collectingCountry>
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and was only recorded once from a pitfall trap in Andohahela.
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="12" pageNumber="459" type="materials_examined">
|
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<paragraph blockId="12.[151,1437,151,1358]" pageId="12" pageNumber="459">
|
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<emphasis bold="true" box="[199,577,1262,1287]" pageId="12" pageNumber="459">Additional material examined:</emphasis>
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Province Toamasina: PN Mantadia,
|
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<quantity box="[1016,1095,1262,1286]" metricMagnitude="2" metricUnit="m" metricValue="8.95" pageId="12" pageNumber="459" unit="m" value="895.0">895 m</quantity>
|
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, rainforest (H.J. Ratsirarson) (
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<collectionCode box="[159,238,1298,1322]" pageId="12" pageNumber="459">CASC</collectionCode>
|
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); Province Toliara: PN Andohahela, Col du Sedro,
|
||
<quantity box="[840,922,1298,1322]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="459" unit="km" value="3.8">3.8 km</quantity>
|
||
<number box="[932,975,1298,1322]" pageId="12" pageNumber="459" value="113.0">113</number>
|
||
° ESE Mahamavo,
|
||
<quantity box="[1199,1295,1298,1322]" metricMagnitude="4" metricUnit="m" metricValue="3.76" pageId="12" pageNumber="459" unit="km" value="37.6">37.6 km</quantity>
|
||
<number box="[1305,1349,1298,1322]" pageId="12" pageNumber="459" value="341.0">341</number>
|
||
° NNW Tolagnaro,
|
||
<quantity box="[278,351,1334,1358]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="459" unit="m" value="900.0">900 m</quantity>
|
||
, montane rainforest (Fisher-Griswold Arthropod Team) (
|
||
<collectionCode box="[989,1068,1334,1358]" pageId="12" pageNumber="459">CASC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |