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<document ID-DOI="10.11646/zootaxa.3683.4.8" ID-GBIF-Dataset="790ed03f-43eb-409c-8fad-3bbbffa0910f" ID-ISSN="1175-5326" ID-Zenodo-Dep="247516" ID-ZooBank="A7A91D16-85AD-40DB-9627-FD02101289A4" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1460645395031" checkinUser="plazi" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2013" docId="8931A65BFFAE4251FF5441C7FA92FDA6" docLanguage="en" docName="zt03683p485.pdf" docOrigin="Zootaxa 3683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachycondyla tahary Rakotonirina and Fisher, sp. nov." docType="treatment" docVersion="10" lastPageNumber="478" masterDocId="7508DE23FFB3424EFFC34533FFA5FF9A" masterDocTitle="Revision of the Pachycondyla sikorae species-group (Hymenoptera: Formicidae) in Madagascar" masterLastPageNumber="485" masterPageNumber="447" pageNumber="476" updateTime="1683651725442" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Pachycondyla sikorae species-group (Hymenoptera: Formicidae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakotonirina, Jean Claude</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-07-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3683</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>447</mods:start>
<mods:end>485</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.3683.4.8</mods:identifier>
<mods:identifier type="GBIF-Dataset">790ed03f-43eb-409c-8fad-3bbbffa0910f</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">247516</mods:identifier>
<mods:identifier type="ZooBank">A7A91D16-85AD-40DB-9627-FD02101289A4</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6162626" ID-GBIF-Taxon="119571906" ID-Zenodo-Dep="6162626" LSID="urn:lsid:plazi:treatment:8931A65BFFAE4251FF5441C7FA92FDA6" httpUri="http://treatment.plazi.org/id/8931A65BFFAE4251FF5441C7FA92FDA6" lastPageId="31" lastPageNumber="478" pageId="29" pageNumber="476">
<subSubSection box="[151,827,1267,1294]" pageId="29" pageNumber="476" type="nomenclature">
<paragraph blockId="29.[151,827,1267,1326]" box="[151,827,1267,1294]" pageId="29" pageNumber="476">
<heading bold="true" box="[151,827,1267,1294]" fontSize="11" level="1" pageId="29" pageNumber="476" reason="1">
<emphasis bold="true" box="[151,827,1267,1294]" pageId="29" pageNumber="476">
<taxonomicName authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[151,721,1267,1294]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="476" phylum="Arthropoda" rank="species" species="tahary" status="sp. nov.">
<emphasis bold="true" box="[151,405,1268,1294]" italics="true" pageId="29" pageNumber="476">Pachycondyla tahary</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel box="[734,827,1267,1293]" pageId="29" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,456,1302,1326]" pageId="29" pageNumber="476" type="description">
<paragraph blockId="29.[151,827,1267,1326]" box="[151,456,1302,1326]" pageId="29" pageNumber="476">
(
<figureCitation box="[158,281,1302,1326]" captionStart="FIGURES 26 27" captionStartId="10.[151,264,1846,1869]" captionTargetBox="[151,1436,1321,1812]" captionTargetId="figure@10.[151,1436,1321,1825]" captionTargetPageId="10" captionText="FIGURES 26 27. Mesosoma in lateral view; Fig. 26: mesopleural suture distinctly continuous and narrowly impressed (CASENT 0317590); Fig. 27: mesopleural suture indistinct (CASENT 0162668: holotype)." httpUri="https://zenodo.org/record/247530/files/figure.png" pageId="29" pageNumber="476">Figures 27</figureCitation>
,
<figureCitation box="[293,407,1302,1326]" captionStart="FIGURES 61 63" captionStartId="30.[151,264,1732,1755]" captionTargetBox="[151,1436,260,1704]" captionTargetId="figure@30.[151,1436,260,1721]" captionTargetPageId="30" captionText="FIGURES 61 63. Worker caste of Pachycondyla tahary: lateral and dorsal view of body and full-face view of head, holotype CASENT 0162668." httpUri="https://zenodo.org/record/247542/files/figure.png" pageId="29" pageNumber="476">61, 62, 63</figureCitation>
,
<figureCitation box="[419,448,1302,1326]" captionStart="FIGURES 76 81" captionStartId="36.[151,264,1974,1997]" captionTargetBox="[178,1407,349,1935]" captionTargetId="figure@36.[161,1425,334,1987]" captionTargetPageId="36" captionText="FIGURES 76 81. Distribution maps of the P. sikorae species-group in Madagascar. Fig. 76: P. m i a l y (triangle) and P. n o s y (circle); Fig. 77: P. ro v a n a; Fig. 78: P. sikorae; Fig. 79: P. t a h a r y; Fig. 80: P. vohitravo; Fig. 81: P. z o ro." httpUri="https://zenodo.org/record/247546/files/figure.png" pageId="29" pageNumber="476">
<number box="[419,448,1302,1326]" pageId="29" pageNumber="476" value="79.0">79</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="476" type="materials_examined">
<paragraph blockId="29.[151,1436,1374,1428]" pageId="29" pageNumber="476">
<materialsCitation ID-GBIF-Occurrence="3744230364" collectingDate="2010-02-20" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="450" latitude="-16.7727" location="Sandrangato River" longLatPrecision="7" longitude="49.2655" municipality="RS Ambatovaky" pageId="29" pageNumber="476" specimenCode="CASENT0162668" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis box="[151,321,1375,1397]" italics="true" pageId="29" pageNumber="476">
<typeStatus box="[151,244,1375,1397]" pageId="29" pageNumber="476">Holotype</typeStatus>
<specimenCount box="[249,321,1375,1397]" pageId="29" pageNumber="476" type="worker">worker</specimenCount>
</emphasis>
:
<collectingCountry box="[334,458,1374,1396]" name="Madagascar" pageId="29" pageNumber="476">Madagascar</collectingCountry>
,
<collectingRegion box="[467,579,1374,1396]" country="Madagascar" name="Toamasina" pageId="29" pageNumber="476">Toamasina</collectingRegion>
,
<collectingMunicipality box="[589,757,1374,1396]" pageId="29" pageNumber="476">RS Ambatovaky</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:8931A65BFFAE4251FF5441C7FA92FDA6:04474196FFAE4253FD3D406DFC18FAEE" box="[766,957,1374,1396]" country="Madagascar" latitude="-16.7727" longLatPrecision="7" longitude="49.2655" municipality="RS Ambatovaky" name="Sandrangato River" pageId="29" pageNumber="476" stateProvince="Toamasina">Sandrangato River</location>
,
<geoCoordinate box="[966,1059,1374,1396]" degrees="16.7727" direction="south" orientation="latitude" pageId="29" pageNumber="476" precision="5" value="-16.7727">
-
<number box="[973,1059,1374,1396]" pageId="29" pageNumber="476" value="16.7727">16.7727</number>
</geoCoordinate>
,
<number box="[1069,1154,1374,1396]" pageId="29" pageNumber="476" value="49.2655">
<geoCoordinate box="[1069,1154,1374,1396]" degrees="49.2655" direction="east" orientation="longitude" pageId="29" pageNumber="476" precision="5" value="49.2655">49.2655</geoCoordinate>
</number>
,
<quantity box="[1163,1229,1374,1396]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="29" pageNumber="476" unit="m" value="450.0">
<elevation box="[1163,1229,1374,1396]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="29" pageNumber="476" unit="m" value="450.0">450 m</elevation>
</quantity>
, rainforest, ex rotten log,
<date box="[244,370,1405,1427]" pageId="29" pageNumber="476" value="2010-02-20">
<collectingDate box="[244,370,1405,1427]" pageId="29" pageNumber="476" value="2010-02-20">
<number box="[244,269,1406,1427]" pageId="29" pageNumber="476" value="20.0">20</number>
Feb
<number box="[320,370,1406,1427]" pageId="29" pageNumber="476" value="2010.0">2010</number>
</collectingDate>
</date>
(
<collectorName box="[385,500,1405,1427]" pageId="29" pageNumber="476">B.L. Fisher</collectorName>
<emphasis box="[506,557,1405,1427]" italics="true" pageId="29" pageNumber="476">et al.</emphasis>
), collection code: BLF
<number box="[792,857,1406,1427]" pageId="29" pageNumber="476" value="24323.0">24323</number>
, specimen code:
<specimenCode box="[1030,1218,1405,1427]" pageId="29" pageNumber="476">CASENT0162668</specimenCode>
(
<collectionCode box="[1231,1303,1406,1428]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="29" pageNumber="476">CASC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="476" type="diagnosis">
<paragraph blockId="29.[151,1437,1471,2037]" pageId="29" pageNumber="476">
<emphasis bold="true" box="[151,421,1471,1496]" pageId="29" pageNumber="476">WORKER. Diagnosis:</emphasis>
Anteromedial margin of clypeus projecting anteriorly as a blunt angle; with head in fullface view, lateral margins before level of anterior margin of eye strongly curving towards base of mandibles; in lateral view, distance between eye and base of mandible roughly less than twice the maximum diameter of eye; antennal scape attaining posterior cephalic margin; posterolateral margins of propodeum and petiolar node with a series of sharp teeth or denticles; mesopleural suture indistinct, while broad and discontinuous lines in different directions may be present; second gastral tergite punctate to sparsely punctate, pubescence absent or reduced on first and second gastral tergites; head behind the level of eye mostly smooth and shiny on faintly effaced reticulatepunctate sculptures.
</paragraph>
</subSubSection>
<subSubSection lastPageId="31" lastPageNumber="478" pageId="29" pageNumber="476" type="description">
<paragraph blockId="29.[151,1437,1471,2037]" pageId="29" pageNumber="476">
<emphasis bold="true" box="[199,557,1759,1784]" pageId="29" pageNumber="476">
Measurements (
<specimenCount box="[393,545,1759,1784]" pageId="29" pageNumber="476" type="generic">
<number box="[393,408,1759,1784]" pageId="29" pageNumber="476" value="8.0">8</number>
specimens
</specimenCount>
):
</emphasis>
HW:
<number box="[631,746,1759,1783]" pageId="29" pageNumber="476" value="1.445" valueMax="1.57" valueMin="1.32">1.321.57</number>
, HL:
<number box="[815,930,1759,1783]" pageId="29" pageNumber="476" value="1.5299999999999998" valueMax="1.67" valueMin="1.39">1.391.67</number>
, CI:
<number box="[991,1064,1759,1783]" pageId="29" pageNumber="476" value="96.5" valueMax="99.0" valueMin="94.0">9499</number>
, SL:
<number box="[1128,1243,1759,1783]" pageId="29" pageNumber="476" value="1.1400000000000001" valueMax="1.3" valueMin="0.98">0.981.30</number>
, SI:
<number box="[1300,1373,1759,1783]" pageId="29" pageNumber="476" value="77.0" valueMax="83.0" valueMin="71.0">7183</number>
, PW:
<number box="[151,265,1796,1820]" pageId="29" pageNumber="476" value="1.075" valueMax="1.17" valueMin="0.98">0.981.17</number>
, WL:
<number box="[335,450,1796,1820]" pageId="29" pageNumber="476" value="2.065" valueMax="2.34" valueMin="1.79">1.792.34</number>
, NH:
<number box="[516,631,1796,1820]" pageId="29" pageNumber="476" value="0.96" valueMax="1.08" valueMin="0.84">0.841.08</number>
, NL:
<number box="[694,809,1796,1820]" pageId="29" pageNumber="476" value="0.74" valueMax="0.85" valueMin="0.63">0.630.85</number>
, NW:
<number box="[881,995,1796,1820]" pageId="29" pageNumber="476" value="0.9500000000000001" valueMax="1.09" valueMin="0.81">0.811.09</number>
, DNI:
<number box="[1071,1172,1796,1820]" pageId="29" pageNumber="476" value="127.0" valueMax="132.0" valueMin="122.0">122132</number>
, LNI:
<number box="[1245,1346,1796,1820]" pageId="29" pageNumber="476" value="131.0" valueMax="139.0" valueMin="123.0">123139</number>
.
</paragraph>
<paragraph blockId="29.[151,1437,1471,2037]" lastBlockId="30.[151,1436,151,212]" lastPageId="30" lastPageNumber="477" pageId="29" pageNumber="476">
<emphasis bold="true" box="[199,347,1831,1856]" pageId="29" pageNumber="476">Description:</emphasis>
Head subquadrate, roughly as long as broad, sides nearly parallel posteriorly before curving abruptly from level of eyes towards base of mandibles; posterior border with weak median excision. Head dorsum reticulate-punctate, with large, smooth, and shiny spaces between punctures; sculptures become punctate and sparsely punctate near posterior margin and sides, respectively. Eyes medium, protruding from head surface, with more than
<number box="[272,300,1976,2000]" pageId="29" pageNumber="476" value="25.0">25</number>
ommatidia; distance between eye and base of mandible approximately less than twice the maximum diameter of eye. Antennal scape barely surpassing posterior margin of head. Median lobe of clypeus convex, anterior margin projecting anteriorly into a median convexity or blunt angle. Mandibles smooth and shiny, with scattered piliferous punctures; masticatory margins with ten teeth or denticles.
</paragraph>
<caption httpUri="https://zenodo.org/record/247542/files/figure.png" pageId="30" pageNumber="477" targetBox="[151,1436,260,1704]" targetPageId="30">
<paragraph blockId="30.[151,1436,1732,1785]" pageId="30" pageNumber="477">
<emphasis bold="true" box="[151,340,1732,1755]" pageId="30" pageNumber="477">FIGURES 6163.</emphasis>
Worker caste of
<taxonomicName box="[514,730,1733,1755]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="477" phylum="Arthropoda" rank="species" species="tahary">
<emphasis box="[514,730,1733,1755]" italics="true" pageId="30" pageNumber="477">Pachycondyla tahary</emphasis>
</taxonomicName>
: lateral and dorsal view of body and full-face view of head, holotype CASENT0162668.
</paragraph>
</caption>
<paragraph blockId="30.[151,1436,1830,2035]" lastBlockId="31.[151,1436,151,572]" lastPageId="31" lastPageNumber="478" pageId="30" pageNumber="477">With mesosoma in dorsal view, metanotal groove absent; mesosoma dorsum mostly smooth between scattered punctures; blunt rugae or tubercles present on pronotum. In lateral view, mesopleural sulcus indistinct, and a broad superficial impression may be present; posterolateral border of propodeum with a series of teeth or tubercles. In profile, lower half of posterolateral margin of petiole node with a series of teeth or tubercles; dorsum and lateral surface of node coarsely bluntly tuberculate. In dorsal view, lateral margins of node forming a convex line to generally rounded anterior border; posterior margin slightly convex. Anterior face of first gastral segment without shallow cavity; first and second gastral tergites punctate laterally, the punctures becoming widely spaced dorsally. Erect hairs long and thin; pubescence abundant on head and pronotum, but gradually reduced or absent from rest of mesosoma dorsum to second gastral tergite. Color black with bluish or opalescent reflection; appendages and tip of gaster brown.</paragraph>
<paragraph blockId="31.[151,1436,151,572]" box="[199,405,295,320]" pageId="31" pageNumber="478">
<emphasis bold="true" box="[199,286,295,320]" pageId="31" pageNumber="478">Queen:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="478" type="multiple">
<paragraph blockId="31.[151,1436,151,572]" pageId="31" pageNumber="478">
<emphasis bold="true" box="[199,503,331,356]" pageId="31" pageNumber="478">Distribution and biology:</emphasis>
Endemic to
<collectingCountry box="[648,785,332,356]" name="Madagascar" pageId="31" pageNumber="478">Madagascar</collectingCountry>
,
<taxonomicName box="[797,899,332,356]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="478" phylum="Arthropoda" rank="species" species="tahary">
<emphasis box="[797,899,332,356]" italics="true" pageId="31" pageNumber="478">P. tahary</emphasis>
</taxonomicName>
is only recorded from the rainforests of the RS Ambatovaky in the east of the island (
<figureCitation box="[577,661,367,391]" captionStart="FIGURES 76 81" captionStartId="36.[151,264,1974,1997]" captionTargetBox="[178,1407,349,1935]" captionTargetId="figure@36.[161,1425,334,1987]" captionTargetPageId="36" captionText="FIGURES 76 81. Distribution maps of the P. sikorae species-group in Madagascar. Fig. 76: P. m i a l y (triangle) and P. n o s y (circle); Fig. 77: P. ro v a n a; Fig. 78: P. sikorae; Fig. 79: P. t a h a r y; Fig. 80: P. vohitravo; Fig. 81: P. z o ro." httpUri="https://zenodo.org/record/247546/files/figure.png" pageId="31" pageNumber="478">
Fig.
<number box="[629,661,367,391]" pageId="31" pageNumber="478" value="79.0">79</number>
</figureCitation>
). It forages mainly through the leaf litter and nests in rotten logs.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="478" type="materials_examined">
<paragraph blockId="31.[151,1436,151,572]" pageId="31" pageNumber="478">
<emphasis bold="true" box="[199,566,403,428]" pageId="31" pageNumber="478">Additional material examined:</emphasis>
Province
<emphasis bold="true" box="[679,809,403,428]" pageId="31" pageNumber="478">Toamasina</emphasis>
: RS Ambatovaky, Sandrangato river,
<quantity box="[1234,1307,404,428]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="478" unit="m" value="360.0">360 m</quantity>
, rainforest (B.L. Fisher
<emphasis box="[303,364,440,464]" italics="true" pageId="31" pageNumber="478">et al.</emphasis>
) (
<collectionCode box="[393,472,439,463]" pageId="31" pageNumber="478">CASC</collectionCode>
); RS Ambatovaky, Sandrangato River, -
<number box="[943,1038,439,463]" pageId="31" pageNumber="478" value="16.7727">16.7727</number>
,
<number box="[1054,1148,439,463]" pageId="31" pageNumber="478" value="49.2655">49.2655</number>
,
<quantity box="[1163,1241,439,463]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="478" unit="m" value="450.0">450 m</quantity>
, rainforest (B.L. Fisher
<emphasis box="[229,287,476,500]" italics="true" pageId="31" pageNumber="478">et al.</emphasis>
) (
<collectionCode box="[314,393,476,500]" pageId="31" pageNumber="478">CASC</collectionCode>
); RS Ambatovaky, Sandrangato River, -
<number box="[849,945,476,500]" pageId="31" pageNumber="478" value="16.7633">16.7633</number>
,
<number box="[957,1051,476,500]" pageId="31" pageNumber="478" value="49.2669">49.2669</number>
,
<quantity box="[1064,1139,476,500]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="31" pageNumber="478" unit="m" value="520.0">520 m</quantity>
, rainforest (B.L. Fisher
<emphasis italics="true" pageId="31" pageNumber="478">et al.</emphasis>
) (
<collectionCode box="[204,283,511,535]" pageId="31" pageNumber="478">CASC</collectionCode>
); RS Ambatovaky, Sandrangato River, -
<number box="[727,823,511,535]" pageId="31" pageNumber="478" value="16.8175">16.8175</number>
, 49.295, 360 m, rainforest (B.L. Fisher
<emphasis box="[1265,1320,512,536]" italics="true" pageId="31" pageNumber="478">et al.</emphasis>
) (
<collectionCode box="[1344,1423,511,535]" pageId="31" pageNumber="478">CASC</collectionCode>
); Sahafina Forest
<quantity box="[333,424,548,572]" metricMagnitude="4" metricUnit="m" metricValue="1.14" pageId="31" pageNumber="478" unit="km" value="11.4">11.4 km</quantity>
W Brickaville, -
<number box="[610,706,548,572]" pageId="31" pageNumber="478" value="18.8144">18.8144</number>
,
<number box="[716,810,548,572]" pageId="31" pageNumber="478" value="48.9621">48.9621</number>
,
<quantity box="[821,894,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="478" unit="m" value="140.0">140 m</quantity>
, rainforest (B.L. Fisher
<emphasis box="[1164,1220,548,572]" italics="true" pageId="31" pageNumber="478">et al.</emphasis>
) (
<collectionCode box="[1244,1323,548,572]" pageId="31" pageNumber="478">CASC</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>