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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1125.86905" ID-Pensoft-Pub="1313-2970-1125-87" ID-Pensoft-UUID="759BB9D7B7525B799D653AC1912E08EC" ID-ZooBank="8B173267C53A4018BFF2E8397DD6EA1F" ModsDocID="1313-2970-1125-87" checkinTime="1666302377346" checkinUser="pensoft" docAuthor="Li, Song-lin, Liu, Ping &amp; Peng, Xian-jin" docDate="2022" docId="8920B59C0B8C58F5A07DB9B5D6FE6B2B" docLanguage="en" docName="ZooKeys 1125: 87-101" docOrigin="ZooKeys 1125" docPubDate="2022-10-20" docSource="http://dx.doi.org/10.3897/zookeys.1125.86905" docTitle="Tetragnatha tortilis Li &amp; Liu 2022, sp. nov." docType="treatment" docUuid="FD801E2B-0F51-44B4-84A0-C57EF7519B93" docUuidSource="ZooBank" docVersion="1" id="759BB9D7B7525B799D653AC1912E08EC" lastPageNumber="87" masterDocId="759BB9D7B7525B799D653AC1912E08EC" masterDocTitle="Three new species of the genus Tetragnatha Latreille, 1804 (Araneae, Tetragnathidae) from China" masterLastPageNumber="101" masterPageNumber="87" pageNumber="87" updateTime="1666302377346" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Three new species of the genus Tetragnatha Latreille, 1804 (Araneae, Tetragnathidae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Song-lin</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Ping</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China</mods:affiliation>
<mods:nameIdentifier type="email">pingzi129@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peng, Xian-jin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2614-3910</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-10-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1125</mods:number>
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<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>101</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1125.86905</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1125.86905</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1125-87</mods:identifier>
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<subSection pageId="0" pageNumber="87" type="taxonomic account">
<treatment LSID="urn:lsid:zoobank.org:act:FD801E2B-0F51-44B4-84A0-C57EF7519B93" httpUri="http://treatment.plazi.org/id/8920B59C0B8C58F5A07DB9B5D6FE6B2B" lastPageNumber="87" pageId="0" pageNumber="87">
<subSubSection pageId="0" pageNumber="87" type="nomenclature">
<paragraph pageId="0" pageNumber="87">
<taxonomicName LSID="https://zoobank.org/FD801E2B-0F51-44B4-84A0-C57EF7519B93" authority="Li &amp; Liu" authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha tortilis" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">Tetragnatha tortilis Li &amp; Liu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">Figs 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tetragnatha tortilis sp. nov., paratype ♀ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, H female genitalia G epigynal fold, ventral view H vulva, dorsal view. Scale bars: 1 mm (A-C); 0.5 mm (D-F); 0.2 mm (G, H)." figureDoi="10.3897/zookeys.1125.86905.figure6" httpUri="https://binary.pensoft.net/fig/760191" pageId="0" pageNumber="87">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Collection localities for Tetragnatha bifurcata sp. nov., Tetragnatha bimaculata sp. nov., and Tetragnatha tortilis sp. nov. in China." figureDoi="10.3897/zookeys.1125.86905.figure7" httpUri="https://binary.pensoft.net/fig/760192" pageId="0" pageNumber="87">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="type material">
<paragraph pageId="0" pageNumber="87">Type material.</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Holotype</emphasis>
♂: China, Yunnan Province, Tengchong County: Jietou Township, Datang Village:
<geoCoordinate degrees="25.4277" direction="north" orientation="latitude" precision="5" value="25.4277">25.4277°N</geoCoordinate>
,
<geoCoordinate degrees="98.4129" direction="east" orientation="longitude" precision="5" value="98.4129">98.4129°E</geoCoordinate>
, 1952 m, 18.V.2006, P. Hu leg. (Hu060518).
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Paratypes</emphasis>
: 1♂ 1♀, same data as holotype (Hu060518); 1♀,
<geoCoordinate degrees="25.7456" direction="north" orientation="latitude" precision="5" value="25.7456">25.7456°N</geoCoordinate>
,
<geoCoordinate degrees="98.6963" direction="east" orientation="longitude" precision="5" value="98.6963">98.6963°E</geoCoordinate>
, 2030 m, 20.V.2006, X.J. Peng &amp; P. Hu leg. (Peng060520); 1♂,
<geoCoordinate degrees="25.4202" direction="north" orientation="latitude" precision="5" value="25.4202">25.4202°N</geoCoordinate>
,
<geoCoordinate degrees="98.4095" direction="east" orientation="longitude" precision="5" value="98.4095">98.4095°E</geoCoordinate>
, 1870 m, 17.V.2006 (Peng060517), 2♂♂,
<geoCoordinate degrees="25.7456" direction="north" orientation="latitude" precision="5" value="25.7456">25.7456°N</geoCoordinate>
,
<geoCoordinate degrees="98.6963" direction="east" orientation="longitude" precision="5" value="98.6963">98.6963°E</geoCoordinate>
, 2030 m, 15.V.2006 (Peng060515), 3♂♂ 2♀♀,
<geoCoordinate degrees="25.7572" direction="north" orientation="latitude" precision="5" value="25.7572">25.7572°N</geoCoordinate>
,
<geoCoordinate degrees="98.6946" direction="east" orientation="longitude" precision="5" value="98.6946">98.6946°E</geoCoordinate>
, 2078 m, 16.V.2006 (Peng060516), 1♂,
<geoCoordinate degrees="25.4202" direction="north" orientation="latitude" precision="5" value="25.4202">25.4202°N</geoCoordinate>
,
<geoCoordinate degrees="98.4095" direction="east" orientation="longitude" precision="5" value="98.4095">98.4095°E</geoCoordinate>
, 1878 m, 19.V.2006 (Peng060519), X. J. Peng et al. leg.; 1♂, Houqiao Township:
<geoCoordinate degrees="25.3539" direction="north" orientation="latitude" precision="5" value="25.3539">25.3539°N</geoCoordinate>
,
<geoCoordinate degrees="98.2549" direction="east" orientation="longitude" precision="5" value="98.2549">98.2549°E</geoCoordinate>
, 1785 m, 28.V.2006 (Wang060528-1), 1♀, Gaoshidong Village,
<geoCoordinate degrees="25.3986" direction="north" orientation="latitude" precision="5" value="25.3986">25.3986°N</geoCoordinate>
,
<geoCoordinate degrees="98.3053" direction="east" orientation="longitude" precision="5" value="98.3053">98.3053°E</geoCoordinate>
, 2374 m, 27.V.2006 (Wang060527-2), 3♀♀, Zhaobitang Village,
<geoCoordinate degrees="25.5380" direction="north" orientation="latitude" precision="5" value="25.538">25.5380°N</geoCoordinate>
,
<geoCoordinate degrees="98.2094" direction="east" orientation="longitude" precision="5" value="98.2094">98.2094°E</geoCoordinate>
, 2480 m, 29.V.2006 (Wang060529-1), X.P. Wang &amp; P. Hu leg.; 2♂♂ 1♀, Mingguang Township: Zizhi Village, Cizhu River,
<geoCoordinate degrees="25.7666" direction="north" orientation="latitude" precision="5" value="25.7666">25.7666°N</geoCoordinate>
,
<geoCoordinate degrees="98.6174" direction="east" orientation="longitude" precision="5" value="98.6174">98.6174°E</geoCoordinate>
, 2120 m, 21.V.2006, C.M. Yin &amp; J.F. Hu leg. (YHY09).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="etymology">
<paragraph pageId="0" pageNumber="87">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="87">
The specific epithet is derived from the Latin adjective
<emphasis italics="true" pageId="0" pageNumber="87">tortilis</emphasis>
(twisted), referring to the twisted distal end of conductor.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="diagnosis">
<paragraph pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="87">
The new species resembles
<taxonomicName lsidName="T. pinicola" pageId="0" pageNumber="87" rank="species" species="pinicola">
<emphasis italics="true" pageId="0" pageNumber="87">T. pinicola</emphasis>
</taxonomicName>
L. Koch, 1870 (
<bibRefCitation author="Zhu, MS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="87" refId="B10" refString="Zhu, MS, Song, DX, Zhang, JX, 2003. Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae." year="2003">Zhu et al. 2003</bibRefCitation>
: figs 87A-G, 88A-G;
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.19.3.07" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="87" pagination="199 - 202" refId="B5" refString="Marusik, YM, 2010. A new species of Tetragnatha Latreille, 1904 (Aranei: Tetragnathidae) from western Kazakhstan. Arthropoda Selecta 19 (3): 199 - 202, DOI: https://doi.org/10.15298/arthsel.19.3.07" title="A new species of Tetragnatha Latreille, 1904 (Aranei: Tetragnathidae) from western Kazakhstan." url="https://doi.org/10.15298/arthsel.19.3.07" volume="19" year="2010">Marusik 2010</bibRefCitation>
: fig. 7). Males of the two species are similar in the presence of an elongated tooth in the upper row of the chelicera in males (
<emphasis italics="true" pageId="0" pageNumber="87">T</emphasis>
), the conductor having 2 folds, and the shape of paracymbium, but can be distinguished by: (1) the dorsal spur of chelicera is straight in ventral view in
<taxonomicName lsidName="T. tortilis" pageId="0" pageNumber="87" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">T. tortilis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">5E</figureCitation>
) (vs. distal end curved; fig. 88C in
<bibRefCitation author="Zhu, MS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="87" refId="B10" refString="Zhu, MS, Song, DX, Zhang, JX, 2003. Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae." year="2003">Zhu et al. [2003]</bibRefCitation>
); (2) the absence of an auxiliary guide tooth of the lower row of the chelicera (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">5E, F</figureCitation>
) (vs. present; fig. 88C in
<bibRefCitation author="Zhu, MS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="87" refId="B10" refString="Zhu, MS, Song, DX, Zhang, JX, 2003. Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae." year="2003">Zhu et al. [2003]</bibRefCitation>
); (3) the distal portion of conductor is twisted and has a small knot (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">5G</figureCitation>
) (vs. hook-shaped; fig. 7 in
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.19.3.07" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="87" pagination="199 - 202" refId="B5" refString="Marusik, YM, 2010. A new species of Tetragnatha Latreille, 1904 (Aranei: Tetragnathidae) from western Kazakhstan. Arthropoda Selecta 19 (3): 199 - 202, DOI: https://doi.org/10.15298/arthsel.19.3.07" title="A new species of Tetragnatha Latreille, 1904 (Aranei: Tetragnathidae) from western Kazakhstan." url="https://doi.org/10.15298/arthsel.19.3.07" volume="19" year="2010">Marusik [2010]</bibRefCitation>
). Females of the two species are similar in the presence of 2 pairs of spermathecae and the absence of a central membranous sac in the vulva, but can be distinguished by the anterior spermathecae 4
<normalizedToken originalValue="×">x</normalizedToken>
larger than posterior in
<taxonomicName lsidName="T. tortilis" pageId="0" pageNumber="87" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">T. tortilis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tetragnatha tortilis sp. nov., paratype ♀ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, H female genitalia G epigynal fold, ventral view H vulva, dorsal view. Scale bars: 1 mm (A-C); 0.5 mm (D-F); 0.2 mm (G, H)." figureDoi="10.3897/zookeys.1125.86905.figure6" httpUri="https://binary.pensoft.net/fig/760191" pageId="0" pageNumber="87">6H</figureCitation>
) (vs. anterior spermathecae ~ 1/2
<normalizedToken originalValue="×">x</normalizedToken>
posterior spermathecae; fig. 87G in
<bibRefCitation author="Zhu, MS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="87" refId="B10" refString="Zhu, MS, Song, DX, Zhang, JX, 2003. Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata Vol. 35, Arachnida: Araneae: Tetragnathidae." year="2003">Zhu et al. [2003]</bibRefCitation>
). Both sexes of the two species can be distinguished by the sternum which is yellowish brown and without a stripe in
<taxonomicName lsidName="T. tortilis" pageId="0" pageNumber="87" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">T. tortilis</emphasis>
</taxonomicName>
sp. nov. (vs. dark brown with yellow stripe).
</paragraph>
<caption doi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 5.</emphasis>
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha tortilis" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">Tetragnatha tortilis</emphasis>
</taxonomicName>
sp. nov., holotype ♂
<emphasis bold="true" pageId="0" pageNumber="87">A-C</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="87">C</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="87">D-F</emphasis>
left chelicera
<emphasis bold="true" pageId="0" pageNumber="87">D</emphasis>
upper view
<emphasis bold="true" pageId="0" pageNumber="87">E</emphasis>
inner view
<emphasis bold="true" pageId="0" pageNumber="87">F</emphasis>
lower view
<emphasis bold="true" pageId="0" pageNumber="87">G, K</emphasis>
conductor and embolus detail
<emphasis bold="true" pageId="0" pageNumber="87">H-J</emphasis>
left palp
<emphasis bold="true" pageId="0" pageNumber="87">H</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="87">I</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="87">J</emphasis>
retrolateral view. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="87">A-C</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="87">D-F, H-J</emphasis>
); 0. 1 mm (
<emphasis bold="true" pageId="0" pageNumber="87">G, K</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">Description.</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Male</emphasis>
(holotype) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">5A-F</figureCitation>
). Total length 4.97. Carapace 1.72 long, 1.05 wide, yellowish brown, fovea, cervical, and radial grooves distinct. Eye sizes and interdistances: AME 0.06, ALE 0.07, PME 0.08, PLE 0.09; AME-AME 0.10, AME-ALE 0.14, PME-PME 0.15, PME-PLE 0.13, MO anterior width 0.24, posterior width 0.31, length 0.27. Clypeus 0.10 high. Labium brown, with thickened edge. Sternum yellowish brown with dark edge. Legs yellowish brown, with sparse spines. Leg measurements: I, 15.59 (4.04, 5.27, 4.87, 1.41); II, 9.96 (2.90, 3.10, 2.98, 0.98); III, 4.46 (1.50, 1.24, 1.12, 0.60); IV, 8.96 (2.93, 2.64, 2.66, 0.73). Chelicera: yellow, ~ 2/3 length of carapace; dorsal spur tapered;
<emphasis italics="true" pageId="0" pageNumber="87">AXu</emphasis>
absent; upper row with 7 teeth:
<emphasis italics="true" pageId="0" pageNumber="87">Gu</emphasis>
curved and almost equal to
<emphasis italics="true" pageId="0" pageNumber="87">sl</emphasis>
in size,
<emphasis italics="true" pageId="0" pageNumber="87">T</emphasis>
present, 4
<emphasis italics="true" pageId="0" pageNumber="87">rsu</emphasis>
decreasing in size gradually;
<emphasis italics="true" pageId="0" pageNumber="87">AXl</emphasis>
absent; lower row with 7 teeth:
<emphasis italics="true" pageId="0" pageNumber="87">Gl</emphasis>
tiny,
<emphasis italics="true" pageId="0" pageNumber="87">L2</emphasis>
slightly smaller than
<emphasis italics="true" pageId="0" pageNumber="87">L3</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="87">L3</emphasis>
largest, all other teeth smaller than
<emphasis italics="true" pageId="0" pageNumber="87">L2</emphasis>
and almost equal in size. Abdomen 3.23 long, 0.85 wide, dorsum grayish yellow, dark folium covering almost complete dorsum, with scattered pale spots and a brown longitudinal line medially, 2 pairs of sigillae; venter grayish yellow, anterior part with sparse pale spots.
</paragraph>
<paragraph pageId="0" pageNumber="87">
Palp (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tetragnatha tortilis sp. nov., holotype ♂ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, K conductor and embolus detail H-J left palp H prolateral view I ventral view J retrolateral view. Scale bars: 1 mm (A-C); 0.2 mm (D-F, H-J); 0. 1 mm (G, K)." figureDoi="10.3897/zookeys.1125.86905.figure5" httpUri="https://binary.pensoft.net/fig/760190" pageId="0" pageNumber="87">5G-K</figureCitation>
). Paracymbium with blunt distal end, notch inconspicuous, translucent lobe elongated, extended to the end, ~ 1/3 of the width of paracymbium, knob spherical. Tegulum oval, ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Conductor with 2 folds, distal portion twisted considerably, and directed to dorsal side. Embolus partially enveloped by conductor.
</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Female</emphasis>
(paratype Hu060518) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tetragnatha tortilis sp. nov., paratype ♀ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, H female genitalia G epigynal fold, ventral view H vulva, dorsal view. Scale bars: 1 mm (A-C); 0.5 mm (D-F); 0.2 mm (G, H)." figureDoi="10.3897/zookeys.1125.86905.figure6" httpUri="https://binary.pensoft.net/fig/760191" pageId="0" pageNumber="87">6A-F</figureCitation>
). Total length 5.70. Carapace 1.77 long, 1.24 wide. Eye sizes and interdistances: AME 0.07, ALE 007, PME 0.08, PLE 0.08; AME-AME 0.10, AME-ALE 0.18, PME-PME 0.12, PME-PLE 0.14. MO anterior width 0.25, posterior width 0.33, length 0.3. Clypeus 0.07 high. Leg measurements: I, 15.33 (4.21, 5.05, 4.73, 1.34); II, 9.48 (2.91, 2.94, 2.70, 0.93); III, 4.4 (1.48, 1.25, 1.00, 0.67); IV, 8.25 (2.72, 2.50, 2.28, 0.75). Chelicera:
<emphasis italics="true" pageId="0" pageNumber="87">AXu</emphasis>
absent; upper row with 7 teeth:
<emphasis italics="true" pageId="0" pageNumber="87">Gu</emphasis>
curved,
<emphasis italics="true" pageId="0" pageNumber="87">U2</emphasis>
largest, other teeth decreasing in size gradually;
<emphasis italics="true" pageId="0" pageNumber="87">AXl</emphasis>
absent; lower row with 6 teeth:
<emphasis italics="true" pageId="0" pageNumber="87">Gl</emphasis>
almost equal to
<emphasis italics="true" pageId="0" pageNumber="87">L3</emphasis>
in size,
<emphasis italics="true" pageId="0" pageNumber="87">L2</emphasis>
largest, other teeth decreasing in size gradually. Abdomen 3.92 long, 1.32 wide, dorsum pale yellow; venter with pale spots on both sides; otherwise, remaining pattern same as in male.
</paragraph>
<caption doi="10.3897/zookeys.1125.86905.figure6" httpUri="https://binary.pensoft.net/fig/760191" pageId="0" pageNumber="87" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 6.</emphasis>
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha tortilis" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">Tetragnatha tortilis</emphasis>
</taxonomicName>
sp. nov., paratype ♀
<emphasis bold="true" pageId="0" pageNumber="87">A-C</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="87">C</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="87">D-F</emphasis>
left chelicera
<emphasis bold="true" pageId="0" pageNumber="87">D</emphasis>
upper view
<emphasis bold="true" pageId="0" pageNumber="87">E</emphasis>
inner view
<emphasis bold="true" pageId="0" pageNumber="87">F</emphasis>
lower view
<emphasis bold="true" pageId="0" pageNumber="87">G, H</emphasis>
female genitalia
<emphasis bold="true" pageId="0" pageNumber="87">G</emphasis>
epigynal fold, ventral view
<emphasis bold="true" pageId="0" pageNumber="87">H</emphasis>
vulva, dorsal view. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="87">A-C</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="87">D-F</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="87">G, H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="87">
Epigyne (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tetragnatha tortilis sp. nov., paratype ♀ A-C habitus A dorsal view B lateral view C ventral view D-F left chelicera D upper view E inner view F lower view G, H female genitalia G epigynal fold, ventral view H vulva, dorsal view. Scale bars: 1 mm (A-C); 0.5 mm (D-F); 0.2 mm (G, H)." figureDoi="10.3897/zookeys.1125.86905.figure6" httpUri="https://binary.pensoft.net/fig/760191" pageId="0" pageNumber="87">6G, H</figureCitation>
). Fold ~ 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Vulva composed of 2 pairs of spermathecae, anterior pair larger, oval and almost touched each other, posterior pair smaller, spherical, and spaced by 3 diameters. Central membranous sac absent.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="87">
Known only from the type locality (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Collection localities for Tetragnatha bifurcata sp. nov., Tetragnatha bimaculata sp. nov., and Tetragnatha tortilis sp. nov. in China." figureDoi="10.3897/zookeys.1125.86905.figure7" httpUri="https://binary.pensoft.net/fig/760192" pageId="0" pageNumber="87">7</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1125.86905.figure7" httpUri="https://binary.pensoft.net/fig/760192" pageId="0" pageNumber="87" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 7.</emphasis>
Collection localities for
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha bifurcata" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis italics="true" pageId="0" pageNumber="87">Tetragnatha bifurcata</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha bimaculata" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis italics="true" pageId="0" pageNumber="87">Tetragnatha bimaculata</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2022" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha tortilis" order="Araneae" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="tortilis">
<emphasis italics="true" pageId="0" pageNumber="87">Tetragnatha tortilis</emphasis>
</taxonomicName>
sp. nov. in China.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>