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<document id="D5D193DEFAE04273A742278CA002B26F" ID="10.11646/zootaxa.4242.3.4SLASH10466" ID-CLB-Dataset="36295" ID-DOI="10.11646/zootaxa.4242.3.4" ID-GBIF-Dataset="2fed5d1a-9893-4b46-a15d-6db3dde8b286" ID-ISSN="1175-5326" ID-Zenodo-Dep="376925" ID-ZooBank="4FCEE7BF-C562-4550-9D42-F7B86678DBBB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" checkinTime="1489394797390" checkinUser="plazi" docAuthor="Machado, Angelo B. M. &amp; Soldati Lacerda, Déborah S." docDate="2017" docId="891587DDFFCE9B28D2A2FB61FB5A60A8" docLanguage="en" docName="zootaxa.4242.3.4SLASH10466.pdf" docOrigin="Zootaxa 4242 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Platystigma pronoti" docType="treatment" docVersion="9" lastPageNumber="502" masterDocId="752CFFA5FFC69B21D235FFCDFFB16712" masterDocTitle="Revalidation of Platystigma Kennedy, 1920, with a synopsis of the quadratum species group and the description of three new species (Odonata: Pseudostigmatidae)" masterLastPageNumber="516" masterPageNumber="493" pageNumber="501" updateTime="1728985839563" updateUser="ExternalLinkService">
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<mods:title id="9FE1768A56A3F87FC147FA510283850E">Revalidation of Platystigma Kennedy, 1920, with a synopsis of the quadratum species group and the description of three new species (Odonata: Pseudostigmatidae)</mods:title>
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<mods:namePart id="004C41081B73EFE62FC0887AD369871E">Machado, Angelo B. M.</mods:namePart>
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<mods:namePart id="D7540ED50A969E488E135C45A620E2F9">Soldati Lacerda, Déborah S.</mods:namePart>
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<treatment id="891587DDFFCE9B28D2A2FB61FB5A60A8" ID-DOI="http://doi.org/10.5281/zenodo.6042207" ID-GBIF-Taxon="127843801" ID-Zenodo-Dep="6042207" LSID="urn:lsid:plazi:treatment:891587DDFFCE9B28D2A2FB61FB5A60A8" httpUri="http://treatment.plazi.org/id/891587DDFFCE9B28D2A2FB61FB5A60A8" lastPageId="9" lastPageNumber="502" pageId="8" pageNumber="501">
<subSubSection id="49A66540FFCE9B29D2A2FB61FE3D63F5" pageId="8" pageNumber="501" type="nomenclature">
<paragraph id="010336CBFFCE9B29D2A2FB61FD0E63D4" blockId="8.[151,703,1196,1255]" box="[151,703,1196,1222]" pageId="8" pageNumber="501">
<heading id="5A4B81A7FFCE9B29D2A2FB61FD0E63D4" bold="true" box="[151,703,1196,1222]" fontSize="11" level="1" pageId="8" pageNumber="501" reason="1">
<emphasis id="33C8EAD9FFCE9B29D2A2FB61FD0E63D4" bold="true" box="[151,703,1196,1222]" pageId="8" pageNumber="501">
<taxonomicName id="C6BC4D48FFCE9B29D2A2FB61FE3563D4" box="[151,388,1196,1222]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCE9B29D2A2FB61FE3563D4" bold="true" box="[151,388,1196,1222]" italics="true" pageId="8" pageNumber="501">Platystigma pronoti</emphasis>
</taxonomicName>
(Sj ӧstedt, 1918) n. comb.
</emphasis>
</heading>
</paragraph>
<paragraph id="010336CBFFCE9B29D2A2FB03FE3D63F5" blockId="8.[151,703,1196,1255]" box="[151,396,1230,1255]" pageId="8" pageNumber="501">
<figureCitation id="99872A4EFFCE9B29D2A2FB03FEA163F5" box="[151,272,1230,1255]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="8" pageNumber="501">Figures 12</figureCitation>
,
<figureCitation id="99872A4EFFCE9B29D329FB02FE8B63F5" box="[284,314,1231,1255]" captionStart="FIGURES 14 19" captionStartId="11.[151,264,1903,1926]" captionTargetBox="[195,1401,236,1826]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 14 19. Pterothorax in lateral view: in P. astictum male, Itatiaia, Rio de Janeiro (14); P. martinezi female, holotype (15); P. humaita sp. nov. male, holotype (16); P. minimum sp. nov. male, holotype (17); P. pronoti female, holotype (18); P. quadratum male, holotype (19)." figureDoi="http://doi.org/10.5281/zenodo.376930" httpUri="https://zenodo.org/record/376930/files/figure.png" pageId="8" pageNumber="501">18</figureCitation>
,
<figureCitation id="99872A4EFFCE9B29D373FB03FE3D63F5" box="[326,396,1230,1255]" captionStart="FIGURES 33 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="8" pageNumber="501">3336</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCE9B29D2A2FADBFC2C6278" pageId="8" pageNumber="501" type="description">
<paragraph id="010336CBFFCE9B29D2A2FADBFC2C6278" blockId="8.[151,1436,1302,1386]" pageId="8" pageNumber="501">
Sj ӧstedt (1918: 31);
<bibRefCitation id="652D4B3AFFCE9B29D35DFADBFD31623E" author="Davies" box="[360,640,1302,1324]" pageId="8" pageNumber="501" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. 1 - Zygoptera and Anisozygoptera. Societas Internationalis Odonatologica, Utrecht, 127 pp." type="book" year="1984">Davies &amp; Tobin (1984: 57)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D0BEFADBFC5A623E" author="Garrison" box="[651,1003,1302,1324]" pageId="8" pageNumber="501" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2016) A synonymic list of the new world Odonata, 73 pp. Available from: http: // www. odonatacentral. org / (accessed 7 February 2017)" type="book" year="2016">Garrison &amp; Ellenrieder (2016: 20)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D1C3FADBFB79623E" author="Machado" box="[1014,1224,1302,1324]" pageId="8" pageNumber="501" refString="Machado, A. B. M. (1992) Regional report: Brazil. In: Moore, N. W. (Org) Report of the 7 th Meeting of the IUCN. Odonata specialist Group. Reports of the Odonata Specialist Group, International Union for Conservation of Nature and Natural Resources, 9, 18 - 21." type="journal article" year="1992">Machado (1992: 19)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D6E1FADBFA26623E" author="Bridges" box="[1236,1431,1302,1324]" pageId="8" pageNumber="501" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world. 3 rd Edition. Privately published, Urbana, Illinois, 995 pp." type="book" year="1994">Bridges (1994: 39)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D2F2FAF8FECA6259" author="Tsuda" box="[199,379,1333,1355]" pageId="8" pageNumber="501" refString="Tsuda, S. (2000) A distributional List of World Odonata. Published by author, Osaka, 430 pp." type="book" year="2000">Tsuda (2000: 58)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D3BDFAF8FC976259" author="Machado" box="[392,806,1333,1355]" pageId="8" pageNumber="501" refString="Machado, A. B. M., Drummond, G. M. &amp; Martins, C. S. (2005) Lista da Fauna Brasileira Ameacada de Extincao. Fundacao Biodiversitas, Belo Horizonte, 157 pp." type="book" year="2005">
Machado,
<bibRefCitation id="652D4B3AFFCE9B29D3C0FAF8FC976259" author="Drummond" box="[501,806,1333,1355]" pageId="8" pageNumber="501" refString="Machado, A. B. M., Drummond, G. M. &amp; Martins, C. S. (2005) Lista da Fauna Brasileira Ameacada de Extincao. Fundacao Biodiversitas, Belo Horizonte, 157 pp." type="book" year="2005">Drummond &amp; Martins (2005</bibRefCitation>
</bibRefCitation>
: 96);
<bibRefCitation id="652D4B3AFFCE9B29D156FAF8FBF86259" author="Heckman" box="[867,1097,1333,1355]" pageId="8" pageNumber="501" refString="Heckman, C. W. (2008) Encyclopedia of South American aquatic insects: Odonata - Zygoptera. Springer, Heidelberg, 687 pp. https: // doi. org / 10.1007 / 978 - 1 - 4020 - 8176 - 7" type="book" year="2008">Heckman (2008: 205)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D660FAF8FEB76278" author="Garrison" pageId="8" pageNumber="501" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly Genera of the New World. The Johns Hopkins University Press, Baltimore, Maryland, 490 pp." type="book" year="2010">
Garrison,
<bibRefCitation id="652D4B3AFFCE9B29D688FAF8FEB76278" author="Ellenrieder" pageId="8" pageNumber="501" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly Genera of the New World. The Johns Hopkins University Press, Baltimore, Maryland, 490 pp." type="book" year="2010">Ellenrieder &amp; Louton (2010</bibRefCitation>
</bibRefCitation>
: 391);
<bibRefCitation id="652D4B3AFFCE9B29D37EFA99FDFA6278" author="Schorr" box="[331,587,1364,1386]" pageId="8" pageNumber="501" refString="Schorr, M. &amp; Paulson, D. (2016) World Odonata List. Revision 3. University of Puget Sound. Available from: http: // www. pugetsound. edu / academics / academic-resources / slater-museum / biodiversity-resources / dragonflies / world-odonatalist 2 / (accessed 14 September 2016)" type="journal article" year="2016">Schorr &amp; Paulson (2016)</bibRefCitation>
;
<bibRefCitation id="652D4B3AFFCE9B29D063FA99FC286278" author="Machado" box="[598,921,1364,1386]" pageId="8" pageNumber="501" refString="Machado, A. B. M. &amp; Lacerda, D. S. S. (2016) Redescription of the holotype of Mecistogaster pronoti Sjostedt, 1918 (Zygoptera: Pseudostigmatidae), International Journal of Odonatology, 19 (1 - 2), 63 - 68." type="journal article" year="2016">Machado &amp; Lacerda (2016: 63)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCE9B29D2A2FA5CFEA862DC" pageId="8" pageNumber="501" type="materials_examined">
<paragraph id="010336CBFFCE9B29D2A2FA5CFEA862DC" blockId="8.[151,1437,1425,2026]" pageId="8" pageNumber="501">
<materialsCitation id="B1D43C96FFCE9B29D2A2FA5CFB2962B8" ID-GBIF-Occurrence="1439511253" box="[151,1176,1425,1450]" country="Brazil" location="Espirito Santo" pageId="8" pageNumber="501" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo" typeStatus="holotype">
<emphasis id="33C8EAD9FFCE9B29D2A2FA5CFE3062B8" bold="true" box="[151,385,1425,1450]" pageId="8" pageNumber="501">Material examined.</emphasis>
<typeStatus id="DE078869FFCE9B29D3BCFA5CFE4062B8" box="[393,497,1425,1450]" pageId="8" pageNumber="501" type="holotype">Holotype</typeStatus>
<specimenCount id="17BAFD42FFCE9B29D3CCFA5FFDA162BB" box="[505,528,1426,1449]" pageId="8" pageNumber="501" type="female"></specimenCount>
:
<collectingCountry id="79AB765BFFCE9B29D02BFA5CFDD462B8" box="[542,613,1425,1450]" name="Brazil" pageId="8" pageNumber="501">Brazil</collectingCountry>
,
<collectingRegion id="C378F829FFCE9B29D044FA5CFCA062B8" box="[625,785,1425,1450]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="501">Espírito Santo</collectingRegion>
(no further information available)
</materialsCitation>
. Holotype deposited in the NHMS.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCE9B29D2F2FA14FAFA60D4" pageId="8" pageNumber="501" type="diagnosis">
<paragraph id="010336CBFFCE9B29D2F2FA14FAFA60D4" blockId="8.[151,1437,1425,2026]" pageId="8" pageNumber="501">
<emphasis id="33C8EAD9FFCE9B29D2F2FA14FEF062E0" bold="true" box="[199,321,1497,1522]" pageId="8" pageNumber="501">Diagnosis.</emphasis>
Head with dorsum black and orange-white spots anteriorly and another medially to antennae base. Pronotum dark brown with pair of pale yellow stripes anteriorly and laterally to middle lobe. Border of medial and apex of lateral portions of posterior prothorax lobe pale yellow. Median portion straight, separated from lateral portions by deep incision at each side (
<figureCitation id="99872A4EFFCE9B29D07BF988FD13614C" box="[590,674,1605,1630]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="8" pageNumber="501">Fig. 12</figureCitation>
); lateral portions with apex rounded (
<figureCitation id="99872A4EFFCE9B29D67CF988FB2C614C" box="[1097,1181,1605,1630]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="8" pageNumber="501">Fig. 12</figureCitation>
). Mesepisternum (
<figureCitation id="99872A4EFFCE9B29D75AF988FF056190" captionStart="FIGURES 14 19" captionStartId="11.[151,264,1903,1926]" captionTargetBox="[195,1401,236,1826]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 14 19. Pterothorax in lateral view: in P. astictum male, Itatiaia, Rio de Janeiro (14); P. martinezi female, holotype (15); P. humaita sp. nov. male, holotype (16); P. minimum sp. nov. male, holotype (17); P. pronoti female, holotype (18); P. quadratum male, holotype (19)." figureDoi="http://doi.org/10.5281/zenodo.376930" httpUri="https://zenodo.org/record/376930/files/figure.png" pageId="8" pageNumber="501">Fig. 18</figureCitation>
) black with antehumeral grayish-yellow stripe occupying whole extension of sclerite. Mesepimeron dark brown with grayish-yellow stripe not reaching lower part of sclerite. Metapleuron (
<figureCitation id="99872A4EFFCE9B29D63FF940FBEF61B4" box="[1034,1118,1677,1702]" captionStart="FIGURES 14 19" captionStartId="11.[151,264,1903,1926]" captionTargetBox="[195,1401,236,1826]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 14 19. Pterothorax in lateral view: in P. astictum male, Itatiaia, Rio de Janeiro (14); P. martinezi female, holotype (15); P. humaita sp. nov. male, holotype (16); P. minimum sp. nov. male, holotype (17); P. pronoti female, holotype (18); P. quadratum male, holotype (19)." figureDoi="http://doi.org/10.5281/zenodo.376930" httpUri="https://zenodo.org/record/376930/files/figure.png" pageId="8" pageNumber="501">Fig. 18</figureCitation>
) whitish yellow with black stripe adjacent to metapleural suture occupying dorsal ¾ of sclerite. Venter of pterothorax whitish yellow, with narrow middle dark stripe occupying its anterior ½. It is impossible to know whether this stripe reaches the posterior part of the venter, because it is damaged. Supplementary tooth of tarsal claws absent. Wings dominantly hyaline with apical area milky-white reaching a row of cells posterior to RP1. Pseudostigma in both wings weakly indicate pale yellow color (
<figureCitation id="99872A4EFFCE9B29D3EDF88CFDD96048" box="[472,616,1857,1882]" captionStart="FIGURES 33 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="8" pageNumber="501">Figs. 33, 35</figureCitation>
). At high saturation, the color of apical area changes; the white color becomes pale blue and the pseudostigma appears very evident, yellow, with six cells in Fw and eight in Hw (
<figureCitation id="99872A4EFFCE9B29D751F8A8FF5260B0" captionStart="FIGURES 33 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="8" pageNumber="501">Figs. 34, 36</figureCitation>
). Costal vein becomes orange and RA to RP1 yellow (
<figureCitation id="99872A4EFFCE9B29D163F844FC5260B0" box="[854,995,1929,1954]" captionStart="FIGURES 33 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="8" pageNumber="501">Figs. 34, 36</figureCitation>
). Abdomen dominantly brown. Cercus dark, with apex rounded. Ovipositor (stylus lacking) reaching posteriorly about level of hind border of S10.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCE9B28D2F2F81CFE3267A2" lastPageId="9" lastPageNumber="502" pageId="8" pageNumber="501" type="description">
<paragraph id="010336CBFFCE9B29D2F2F81CFD0E60F8" blockId="8.[151,1437,1425,2026]" box="[199,703,2001,2026]" pageId="8" pageNumber="501">
<emphasis id="33C8EAD9FFCE9B29D2F2F81CFE0F60F8" bold="true" box="[199,446,2001,2026]" pageId="8" pageNumber="501">Measurements (mm)</emphasis>
: Hw 40. Abdomen 54.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FF5AFE3267A2" blockId="9.[151,1437,151,504]" box="[199,387,151,176]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FF5AFEBC67A2" bold="true" box="[199,269,151,176]" pageId="9" pageNumber="502">Male:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCF9B28D2F2FF71FDEF6594" pageId="9" pageNumber="502" type="discussion">
<paragraph id="010336CBFFCF9B28D2F2FF71FBBF66E5" blockId="9.[151,1437,151,504]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FF71FE8A67C7" bold="true" box="[199,315,188,213]" pageId="9" pageNumber="502">Remarks.</emphasis>
<taxonomicName id="C6BC4D48FFCF9B28D37CFF70FDF767C6" box="[329,582,189,212]" class="Insecta" family="Pseudostigmatidae" genus="Mecistogaster" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCF9B28D37CFF70FDF767C6" box="[329,582,189,212]" italics="true" pageId="9" pageNumber="502">Mecistogaster pronoti</emphasis>
</taxonomicName>
is transferred here to
<taxonomicName id="C6BC4D48FFCF9B28D16AFF71FC5467C7" box="[863,997,188,213]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="33C8EAD9FFCF9B28D16AFF71FC5467C7" box="[863,997,188,213]" italics="true" pageId="9" pageNumber="502">Platystigma</emphasis>
</taxonomicName>
based on its small size, absence of supplementary tooth of tarsal claw and presence of pale apical areas on the wings. The species was described by Sj ӧstedt (1918) based on a single female specimen from the state of
<collectingRegion id="C378F829FFCF9B28D183FEC9FBEB660E" box="[950,1114,260,285]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="502">Espírito Santo</collectingRegion>
,
<collectingCountry id="79AB765BFFCF9B28D65CFEC9FB01660F" box="[1129,1200,260,285]" name="Brazil" pageId="9" pageNumber="502">Brazil</collectingCountry>
, and has never been found again. According to
<bibRefCitation id="652D4B3AFFCF9B28D3FBFEEAFD206652" author="Heckman" box="[462,657,295,320]" pageId="9" pageNumber="502" refString="Heckman, C. W. (2008) Encyclopedia of South American aquatic insects: Odonata - Zygoptera. Springer, Heidelberg, 687 pp. https: // doi. org / 10.1007 / 978 - 1 - 4020 - 8176 - 7" type="book" year="2008">Heckman (2008)</bibRefCitation>
, “
<taxonomicName id="C6BC4D48FFCF9B28D09EFEE4FC956652" box="[683,804,297,320]" class="Insecta" family="Pseudostigmatidae" genus="Mecistogaster" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCF9B28D09EFEE4FC956652" box="[683,804,297,320]" italics="true" pageId="9" pageNumber="502">M. pronoti</emphasis>
</taxonomicName>
is a rare or unrecognized species.” The
<typeStatus id="DE078869FFCF9B28D6CEFEEAFAEC6652" box="[1275,1373,295,320]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C6BC4D48FFCF9B28D7B2FEE4FF596676" class="Insecta" family="Pseudostigmatidae" genus="Mecistogaster" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCF9B28D7B2FEE4FF596676" italics="true" pageId="9" pageNumber="502">P. pronoti</emphasis>
</taxonomicName>
has been recently redescribed and illustrated by
<bibRefCitation id="652D4B3AFFCF9B28D10FFE81FB376677" author="Machado" box="[826,1158,332,357]" pageId="9" pageNumber="502" refString="Machado, A. B. M. &amp; Lacerda, D. S. S. (2016) Redescription of the holotype of Mecistogaster pronoti Sjostedt, 1918 (Zygoptera: Pseudostigmatidae), International Journal of Odonatology, 19 (1 - 2), 63 - 68." type="journal article" year="2016">Machado &amp; Lacerda (2016)</bibRefCitation>
, thus allowing its easy identification. The main character separating
<taxonomicName id="C6BC4D48FFCF9B28D0ADFEBCFCB7669A" box="[664,774,369,392]" class="Insecta" family="Pseudostigmatidae" genus="Mecistogaster" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCF9B28D0ADFEBCFCB7669A" box="[664,774,369,392]" italics="true" pageId="9" pageNumber="502">P. pronoti</emphasis>
</taxonomicName>
from the other
<taxonomicName id="C6BC4D48FFCF9B28D18DFEA2FB8F669A" box="[952,1086,367,392]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="33C8EAD9FFCF9B28D18DFEA2FB8F669A" box="[952,1086,367,392]" italics="true" pageId="9" pageNumber="502">Platystigma</emphasis>
</taxonomicName>
species is the peculiar form of the hind prothoracic lobe, whose median portion is clearly separated from the lateral ones.
<taxonomicName id="C6BC4D48FFCF9B28D688FE58FA9C66BE" box="[1213,1325,405,428]" class="Insecta" family="Pseudostigmatidae" genus="Mecistogaster" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFCF9B28D688FE58FA9C66BE" box="[1213,1325,405,428]" italics="true" pageId="9" pageNumber="502">P. pronoti</emphasis>
</taxonomicName>
has been listed as Critically Endangered in Brazils, Portarias MMA n o 444/445 (2014) and IUCN (2006) red list, and a strategy for its conservation has been proposed by
<bibRefCitation id="652D4B3AFFCF9B28D0FBFE13FBBB66E5" author="Machado" box="[718,1034,478,503]" pageId="9" pageNumber="502" refString="Machado, A. B. M. &amp; Lacerda, D. S. S. (2016) Redescription of the holotype of Mecistogaster pronoti Sjostedt, 1918 (Zygoptera: Pseudostigmatidae), International Journal of Odonatology, 19 (1 - 2), 63 - 68." type="journal article" year="2016">Machado &amp; Lacerda (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="010336CBFFCF9B28D2A2FD87FDA36576" blockId="9.[151,606,586,646]" box="[151,530,586,612]" pageId="9" pageNumber="502">
<taxonomicName id="C6BC4D48FFCF9B28D2A2FD87FE006576" box="[151,433,586,612]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="quadratum" status="sp. nov.">
<emphasis id="33C8EAD9FFCF9B28D2A2FD87FE006576" bold="true" box="[151,433,586,612]" italics="true" pageId="9" pageNumber="502">Platystigma quadratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28FB57A2FFCF9B28D38CFD86FDA36576" box="[441,530,587,612]" pageId="9" pageNumber="502" rank="species">
<emphasis id="33C8EAD9FFCF9B28D38CFD86FDA36576" bold="true" box="[441,530,587,612]" pageId="9" pageNumber="502">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="010336CBFFCF9B28D2A2FDA0FDEF6594" blockId="9.[151,606,586,646]" box="[151,606,621,646]" pageId="9" pageNumber="502">
<figureCitation id="99872A4EFFCF9B28D2A2FDA0FEA16594" box="[151,272,621,646]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="9" pageNumber="502">Figures 13</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D329FDA3FE8B6594" box="[284,314,622,646]" captionStart="FIGURES 14 19" captionStartId="11.[151,264,1903,1926]" captionTargetBox="[195,1401,236,1826]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 14 19. Pterothorax in lateral view: in P. astictum male, Itatiaia, Rio de Janeiro (14); P. martinezi female, holotype (15); P. humaita sp. nov. male, holotype (16); P. minimum sp. nov. male, holotype (17); P. pronoti female, holotype (18); P. quadratum male, holotype (19)." figureDoi="http://doi.org/10.5281/zenodo.376930" httpUri="https://zenodo.org/record/376930/files/figure.png" pageId="9" pageNumber="502">19</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D373FDA3FED56594" box="[326,356,622,646]" captionStart="FIGURES 20 24" captionStartId="12.[151,264,1945,1968]" captionTargetBox="[280,1335,276,1822]" captionTargetId="figure@12.[151,1436,193,1924]" captionTargetPageId="12" captionText="FIGURES 20 24. Venter in P. astictum male, Itatiaia, Rio de Janeiro (20); P. martinezi female, holotype (21); P. humaita sp. nov. male, holotype (22). P. minimum sp. nov. male, holotype (23); P. quadratum sp. nov. male, holotype (24)." figureDoi="http://doi.org/10.5281/zenodo.376931" httpUri="https://zenodo.org/record/376931/files/figure.png" pageId="9" pageNumber="502">24</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D345FDA3FE3E6594" box="[368,399,622,646]" captionStart="FIGURES 29 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="9" pageNumber="502">32</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D3AFFDA3FE096594" box="[410,440,622,646]" captionStart="FIGURES 37 41" captionStartId="16.[151,264,1976,1999]" captionTargetBox="[254,1339,240,1915]" captionTargetId="figure@16.[165,1422,141,1955]" captionTargetPageId="16" captionText="FIGURES 37 41. Genitalia of second segment in P. astictum, Tinguá, Rio de Janeiro (37); P. martinezi, Bolívia, Santa Cruz (38); P. humaita sp. nov., holotype (39); P. minimum sp. nov., holotype (40); P. quadratum sp. nov., holotype (41)." figureDoi="http://doi.org/10.5281/zenodo.376935" httpUri="https://zenodo.org/record/376935/files/figure.png" pageId="9" pageNumber="502">41</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D3F1FDA0FDBD6594" box="[452,524,621,646]" captionStart="FIGURES 42 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="9" pageNumber="502">50, 51</figureCitation>
,
<figureCitation id="99872A4EFFCF9B28D02DFDA0FDEF6594" box="[536,606,621,646]" captionStart="FIGURES 54 61" captionStartId="19.[151,264,1836,1859]" captionTargetBox="[208,1385,237,1720]" captionTargetId="figure@19.[151,1436,193,1815]" captionTargetPageId="19" captionText="FIGURES 54 61. Male cercus in P. martinezi, Bolívia, Santa Cruz, in dorsal (54) and lateral (55) views; P. humaita sp. nov., holotype, in dorsal (56) and lateral (57) views; P. minimum sp. nov. in dorsal (58) and lateral (59) views; P. quadratum sp. nov. in dorsal (60) and lateral (61) views." figureDoi="http://doi.org/10.5281/zenodo.376938" httpUri="https://zenodo.org/record/376938/files/figure.png" pageId="9" pageNumber="502">60, 61</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCF9B28D2A2FD78FB4F65E0" pageId="9" pageNumber="502" type="materials_examined">
<paragraph id="010336CBFFCF9B28D2A2FD78FB4F65E0" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<materialsCitation id="B1D43C96FFCF9B28D2A2FD78FD4A65E0" ID-GBIF-Occurrence="1439511249" collectingDate="2003-09" collectionCode="ABMM" collectorName="Mielke &amp; Casagrande" country="Brazil" county="Material" latitude="-10.066667" location="Reserva Catuaba" longLatPrecision="21" longitude="-67.599724" municipality="Senador Guiomard" pageId="9" pageNumber="502" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="holotype">
<emphasis id="33C8EAD9FFCF9B28D2A2FD78FE3065DC" bold="true" box="[151,385,693,718]" pageId="9" pageNumber="502">
<collectingCounty id="E8624E47FFCF9B28D2A2FD78FF4E65DC" box="[151,255,693,718]" pageId="9" pageNumber="502">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="DE078869FFCF9B28D3BFFD78FE4365DC" box="[394,498,693,718]" pageId="9" pageNumber="502" type="holotype">Holotype</typeStatus>
<specimenCount id="17BAFD42FFCF9B28D3CFFD7BFDA365DF" box="[506,530,694,717]" pageId="9" pageNumber="502" type="male"></specimenCount>
:
<collectingCountry id="79AB765BFFCF9B28D02AFD78FDD765DC" box="[543,614,693,718]" name="Brazil" pageId="9" pageNumber="502">Brazil</collectingCountry>
,
<collectingRegion id="C378F829FFCF9B28D046FD7BFD1865DC" box="[627,681,694,718]" country="Brazil" name="Acre" pageId="9" pageNumber="502">Acre</collectingRegion>
State,
<collectingMunicipality id="E167ACB1FFCF9B28D0CDFD78FC6065DC" box="[760,977,693,718]" pageId="9" pageNumber="502">Senador Guiomard</collectingMunicipality>
,
<location id="04636010FFCF9B28D1E8FD7BFB2A65DC" LSID="urn:lsid:plazi:treatment:891587DDFFCE9B28D2A2FB61FB5A60A8:04636010FFCF9B28D1E8FD7BFB2A65DC" box="[989,1179,693,718]" country="Brazil" county="Material" latitude="-10.066667" longLatPrecision="21" longitude="-67.599724" municipality="Senador Guiomard" name="Reserva Catuaba" pageId="9" pageNumber="502" stateProvince="Acre">Reserva Catuaba</location>
(
<geoCoordinate id="6488500CFFCF9B28D699FD7BFA8F65DC" box="[1196,1342,693,718]" direction="south" orientation="latitude" pageId="9" pageNumber="502" precision="15" value="-10.066667">10° 03' 60&quot;S</geoCoordinate>
,
<geoCoordinate id="6488500CFFCF9B28D779FD78FF6D65E0" direction="west" orientation="longitude" pageId="9" pageNumber="502" precision="15" value="-67.599724">67° 35' 59&quot;W</geoCoordinate>
),
<collectingDate id="6546E9E3FFCF9B28D2C5FD17FEE765E0" box="[240,342,730,754]" bridgedPair="-" pageId="9" pageNumber="502" value="2003-09">
<date id="7502100BFFCF9B28D2C5FD17FEE765E0" box="[240,342,730,754]" bridgedPair="-" pageId="9" pageNumber="502" value="2003-09">IX2003</date>
</collectingDate>
,
<collectorName id="AC49531DFFCF9B28D357FD14FE0065E0" box="[354,433,729,754]" pageId="9" pageNumber="502">Mielke</collectorName>
&amp;
<collectorName id="AC49531DFFCF9B28D3E0FD14FDE965E0" box="[469,600,729,754]" pageId="9" pageNumber="502">Casagrande</collectorName>
leg. (
<collectionCode id="67ADAE0EFFCF9B28D0A3FD17FD4265E0" box="[662,755,730,754]" pageId="9" pageNumber="502">ABMM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B1D43C96FFCF9B28D0CBFD17FB4F65E0" ID-GBIF-Occurrence="1439511252" box="[766,1278,729,754]" collectingDate="2003-09" collectionCode="ABMM" collectorName="Mielke &amp; Casagrande" country="Brazil" county="Material" latitude="-10.066667" location="Reserva Catuaba" longLatPrecision="21" longitude="-67.599724" municipality="Senador Guiomard" pageId="9" pageNumber="502" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="paratype">
Two
<specimenCount id="17BAFD42FFCF9B28D10CFD17FCFF65E3" box="[825,846,730,753]" pageId="9" pageNumber="502" type="male"></specimenCount>
<typeStatus id="DE078869FFCF9B28D160FD17FC7165E0" box="[853,960,730,754]" pageId="9" pageNumber="502" type="paratype">paratypes</typeStatus>
with same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCF9B28D2F2FD30FF626428" pageId="9" pageNumber="502" type="etymology">
<paragraph id="010336CBFFCF9B28D2F2FD30FF626428" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FD30FEFF6404" bold="true" box="[199,334,765,790]" pageId="9" pageNumber="502">Etymology.</emphasis>
From Latin
<emphasis id="33C8EAD9FFCF9B28D3D6FD30FD106404" box="[483,673,765,790]" italics="true" pageId="9" pageNumber="502">quadratum-a-um</emphasis>
, quadrate, an allusion to the shape of its penis segment
<quantity id="C6449B2EFFCF9B28D705FD33FAEC6404" box="[1328,1373,765,790]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="502" unit="in" value="3.0">3 in</quantity>
ectal view.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCF9B28D2F2FC88FE2F6060" pageId="9" pageNumber="502" type="description">
<paragraph id="010336CBFFCF9B28D2F2FC88FC0964B4" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FC88FDE4644C" bold="true" box="[199,597,837,862]" pageId="9" pageNumber="502">
Description of the male
<typeStatus id="DE078869FFCF9B28D3D2FC88FDE1644C" box="[487,592,837,862]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="33C8EAD9FFCF9B28D068FC88FD28644C" box="[605,665,837,862]" italics="true" pageId="9" pageNumber="502">Head</emphasis>
. Labium whitish yellow, base of mandible and genae bluish yellow, anteclypeus and postclypeus grayish blue, antefrons brownish. Upper part of head black with metallic green luster. Postocular lobe and occipital crest yellow. Rear of head whitish yellow.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FC7CFBC0636C" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FC7CFEA964D8" box="[199,280,945,970]" italics="true" pageId="9" pageNumber="502">Thorax</emphasis>
. Prothorax: pronotum black. Anterior lobe with pair of pale oblong transverse spots. Middle lobe with large pale marking dilated posteriorly (
<figureCitation id="99872A4EFFCF9B28D065FC18FD1564FC" box="[592,676,981,1006]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="9" pageNumber="502">Fig. 13</figureCitation>
). Posterior lobe bordered with yellow (
<figureCitation id="99872A4EFFCF9B28D657FC18FB0764FC" box="[1122,1206,981,1006]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="9" pageNumber="502">Fig. 13</figureCitation>
). Propleuron yellow with black markings (
<figureCitation id="99872A4EFFCF9B28D391FC34FDB16300" box="[420,512,1017,1042]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="9" pageNumber="502">Fig. 13</figureCitation>
). Pterothorax: mesepisternum (
<figureCitation id="99872A4EFFCF9B28D14DFC34FC636300" box="[888,978,1017,1042]" captionStart="FIGURES 14 19" captionStartId="11.[151,264,1903,1926]" captionTargetBox="[195,1401,236,1826]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 14 19. Pterothorax in lateral view: in P. astictum male, Itatiaia, Rio de Janeiro (14); P. martinezi female, holotype (15); P. humaita sp. nov. male, holotype (16); P. minimum sp. nov. male, holotype (17); P. pronoti female, holotype (18); P. quadratum male, holotype (19)." figureDoi="http://doi.org/10.5281/zenodo.376930" httpUri="https://zenodo.org/record/376930/files/figure.png" pageId="9" pageNumber="502">Fig. 19</figureCitation>
) black with middorsal carina yellow, antehumeral yellow stripe reduced to triangular marking at about 1/3 of sclerite. Posthumeral yellow stripe not reaching lower part of sclerite. Mesepimeron black. Metapleuron yellow with black stripe along metapleural suture. Venter with anterior triangular and posterior subcylindrical black spots (
<figureCitation id="99872A4EFFCF9B28D625FBA8FBD4636C" box="[1040,1125,1125,1150]" captionStart="FIGURES 20 24" captionStartId="12.[151,264,1945,1968]" captionTargetBox="[280,1335,276,1822]" captionTargetId="figure@12.[151,1436,193,1924]" captionTargetPageId="12" captionText="FIGURES 20 24. Venter in P. astictum male, Itatiaia, Rio de Janeiro (20); P. martinezi female, holotype (21); P. humaita sp. nov. male, holotype (22). P. minimum sp. nov. male, holotype (23); P. quadratum sp. nov. male, holotype (24)." figureDoi="http://doi.org/10.5281/zenodo.376931" httpUri="https://zenodo.org/record/376931/files/figure.png" pageId="9" pageNumber="502">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FB47FED46247" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FB47FEBB63B3" box="[199,266,1162,1185]" italics="true" pageId="9" pageNumber="502">Wings</emphasis>
. Fw hyaline with light brown pseudostigma, with six cells. Hw with large apical area bluish white reaching RP2; pseudostigma blue with eight cells (
<figureCitation id="99872A4EFFCF9B28D0E0FB60FC9863D4" box="[725,809,1197,1222]" captionStart="FIGURES 29 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="9" pageNumber="502">Fig. 32</figureCitation>
). At high saturation, bluish white area becomes whitish blue, surrounded by blue and pseudostigma more intensely blue. Venation: Px in Fw 31, in Hw 33, RP
<quantity id="C6449B2EFFCF9B28D722FB1FFAF363F8" box="[1303,1346,1233,1258]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="502" unit="in" value="2.0">2 in</quantity>
Fw and in Hw originating at Px 11. IR1 originating at Px
<quantity id="C6449B2EFFCF9B28D0FFFB38FCB5621C" box="[714,772,1269,1294]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="9" pageNumber="502" unit="in" value="16.0">16 in</quantity>
Fw, at Px
<quantity id="C6449B2EFFCF9B28D14BFB3BFC09621C" box="[894,952,1269,1294]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="502" unit="in" value="14.0">14 in</quantity>
Hw. RP2 branching from RP at about half distance from subnodus to tip of wing. Petiolation originates distal to CuP by two times length of CuP in Fw and three times in Hw.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FAAFFC50628C" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FAAFFF4D626B" box="[199,252,1378,1401]" italics="true" pageId="9" pageNumber="502">Legs</emphasis>
. Femora with proximal 1/3 yellow, distally black on extensor surface and yellow on flexural surface. Tibiae with extensor surface yellow and flexural surface black. Tarsi black.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FA64FCCD62D0" blockId="9.[151,1437,693,1978]" box="[199,892,1449,1474]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FA64FE8062D0" box="[199,305,1449,1474]" italics="true" pageId="9" pageNumber="502">Abdomen</emphasis>
. S110 dorsally black, laterally yellow. Cerci black.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2FA00FA3561F0" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2FA00FE7D62F4" bold="true" box="[199,460,1485,1510]" pageId="9" pageNumber="502">Structural characters</emphasis>
. Median and lateral portions of posterior prothoracic lobe not separated. Posterior border slightly convex (
<figureCitation id="99872A4EFFCF9B28D393FA3CFE4B6118" box="[422,506,1521,1546]" captionStart="FIGURES 8 13" captionStartId="10.[151,264,1930,1953]" captionTargetBox="[207,1392,211,1857]" captionTargetId="figure@10.[159,1428,127,1941]" captionTargetPageId="10" captionText="FIGURES 8 13. Prothorax in dorsal view: in P. astictum male, Itatiaia, Rio de Janeiro (8); P. martinezi female, holotype, Bolívia, Santa Cruz (9); P. humaita sp. nov. male, holotype, Porto Acre (10); P. minimum sp. nov. male, holotype (11); P. pronoti female, holotype (12); P. quadratum sp. nov. male, holotype (13)." figureDoi="http://doi.org/10.5281/zenodo.376929" httpUri="https://zenodo.org/record/376929/files/figure.png" pageId="9" pageNumber="502">Fig. 13</figureCitation>
). Cleft of anterior lamina shaped as ogival Gothic arch (
<figureCitation id="99872A4EFFCF9B28D648FA3CFB636118" box="[1149,1234,1521,1546]" captionStart="FIGURES 37 41" captionStartId="16.[151,264,1976,1999]" captionTargetBox="[254,1339,240,1915]" captionTargetId="figure@16.[165,1422,141,1955]" captionTargetPageId="16" captionText="FIGURES 37 41. Genitalia of second segment in P. astictum, Tinguá, Rio de Janeiro (37); P. martinezi, Bolívia, Santa Cruz (38); P. humaita sp. nov., holotype (39); P. minimum sp. nov., holotype (40); P. quadratum sp. nov., holotype (41)." figureDoi="http://doi.org/10.5281/zenodo.376935" httpUri="https://zenodo.org/record/376935/files/figure.png" pageId="9" pageNumber="502">Fig. 41</figureCitation>
). Anterior hamuli with ear-like basal plate. Posterior hamuli with apex curved anteromedially (
<figureCitation id="99872A4EFFCF9B28D1CCF9D8FBFC613C" box="[1017,1101,1557,1582]" captionStart="FIGURES 37 41" captionStartId="16.[151,264,1976,1999]" captionTargetBox="[254,1339,240,1915]" captionTargetId="figure@16.[165,1422,141,1955]" captionTargetPageId="16" captionText="FIGURES 37 41. Genitalia of second segment in P. astictum, Tinguá, Rio de Janeiro (37); P. martinezi, Bolívia, Santa Cruz (38); P. humaita sp. nov., holotype (39); P. minimum sp. nov., holotype (40); P. quadratum sp. nov., holotype (41)." figureDoi="http://doi.org/10.5281/zenodo.376935" httpUri="https://zenodo.org/record/376935/files/figure.png" pageId="9" pageNumber="502">Fig. 41</figureCitation>
). Distal portion of batilliform lamina very small, apex sclerotized (
<figureCitation id="99872A4EFFCF9B28D00BF9F4FD226140" box="[574,659,1593,1618]" captionStart="FIGURES 37 41" captionStartId="16.[151,264,1976,1999]" captionTargetBox="[254,1339,240,1915]" captionTargetId="figure@16.[165,1422,141,1955]" captionTargetPageId="16" captionText="FIGURES 37 41. Genitalia of second segment in P. astictum, Tinguá, Rio de Janeiro (37); P. martinezi, Bolívia, Santa Cruz (38); P. humaita sp. nov., holotype (39); P. minimum sp. nov., holotype (40); P. quadratum sp. nov., holotype (41)." figureDoi="http://doi.org/10.5281/zenodo.376935" httpUri="https://zenodo.org/record/376935/files/figure.png" pageId="9" pageNumber="502">Fig. 41</figureCitation>
). Penis segment
<quantity id="C6449B2EFFCF9B28D162F9F7FC356140" box="[855,900,1593,1618]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="502" unit="in" value="3.0">3 in</quantity>
ectal view subquadrate with small tongue-like apical lobe and tuft of spines at anterolateral angle (
<figureCitation id="99872A4EFFCF9B28D0DCF990FC8E6164" box="[745,831,1629,1654]" captionStart="FIGURES 42 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="9" pageNumber="502">Fig. 50</figureCitation>
); in lateral view with medial and distal tuft of spines (
<figureCitation id="99872A4EFFCF9B28D2AAF94CFF456188" box="[159,244,1665,1690]" captionStart="FIGURES 42 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="9" pageNumber="502">Fig. 51</figureCitation>
). Penis segment 2 with large tuft of spines near commissure (
<figureCitation id="99872A4EFFCF9B28D181F94CFB8E6188" box="[948,1087,1665,1690]" captionStart="FIGURES 42 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="9" pageNumber="502">Figs. 5051</figureCitation>
). Hind border of S
<quantity id="C6449B2EFFCF9B28D721F94FFAFE6188" box="[1300,1359,1665,1690]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="502" unit="in" value="10.0">10 in</quantity>
dorsal view with medial concavity (
<figureCitation id="99872A4EFFCF9B28D3C7F968FDFA61AC" box="[498,587,1701,1726]" captionStart="FIGURES 54 61" captionStartId="19.[151,264,1836,1859]" captionTargetBox="[208,1385,237,1720]" captionTargetId="figure@19.[151,1436,193,1815]" captionTargetPageId="19" captionText="FIGURES 54 61. Male cercus in P. martinezi, Bolívia, Santa Cruz, in dorsal (54) and lateral (55) views; P. humaita sp. nov., holotype, in dorsal (56) and lateral (57) views; P. minimum sp. nov. in dorsal (58) and lateral (59) views; P. quadratum sp. nov. in dorsal (60) and lateral (61) views." figureDoi="http://doi.org/10.5281/zenodo.376938" httpUri="https://zenodo.org/record/376938/files/figure.png" pageId="9" pageNumber="502">Fig. 60</figureCitation>
); in lateral view with distal ½ elevated (
<figureCitation id="99872A4EFFCF9B28D603F968FB3E61AC" box="[1078,1167,1701,1726]" captionStart="FIGURES 54 61" captionStartId="19.[151,264,1836,1859]" captionTargetBox="[208,1385,237,1720]" captionTargetId="figure@19.[151,1436,193,1815]" captionTargetPageId="19" captionText="FIGURES 54 61. Male cercus in P. martinezi, Bolívia, Santa Cruz, in dorsal (54) and lateral (55) views; P. humaita sp. nov., holotype, in dorsal (56) and lateral (57) views; P. minimum sp. nov. in dorsal (58) and lateral (59) views; P. quadratum sp. nov. in dorsal (60) and lateral (61) views." figureDoi="http://doi.org/10.5281/zenodo.376938" httpUri="https://zenodo.org/record/376938/files/figure.png" pageId="9" pageNumber="502">Fig. 61</figureCitation>
). Cerci in lateral view cylindrical, with distal 1/3 tapering and directed ventrally (
<figureCitation id="99872A4EFFCF9B28D11FF904FCCF61F0" box="[810,894,1737,1762]" captionStart="FIGURES 54 61" captionStartId="19.[151,264,1836,1859]" captionTargetBox="[208,1385,237,1720]" captionTargetId="figure@19.[151,1436,193,1815]" captionTargetPageId="19" captionText="FIGURES 54 61. Male cercus in P. martinezi, Bolívia, Santa Cruz, in dorsal (54) and lateral (55) views; P. humaita sp. nov., holotype, in dorsal (56) and lateral (57) views; P. minimum sp. nov. in dorsal (58) and lateral (59) views; P. quadratum sp. nov. in dorsal (60) and lateral (61) views." figureDoi="http://doi.org/10.5281/zenodo.376938" httpUri="https://zenodo.org/record/376938/files/figure.png" pageId="9" pageNumber="502">Fig. 61</figureCitation>
). In dorsal view, distally convergent (
<figureCitation id="99872A4EFFCF9B28D716F904FAC661F0" box="[1315,1399,1737,1762]" captionStart="FIGURES 54 61" captionStartId="19.[151,264,1836,1859]" captionTargetBox="[208,1385,237,1720]" captionTargetId="figure@19.[151,1436,193,1815]" captionTargetPageId="19" captionText="FIGURES 54 61. Male cercus in P. martinezi, Bolívia, Santa Cruz, in dorsal (54) and lateral (55) views; P. humaita sp. nov., holotype, in dorsal (56) and lateral (57) views; P. minimum sp. nov. in dorsal (58) and lateral (59) views; P. quadratum sp. nov. in dorsal (60) and lateral (61) views." figureDoi="http://doi.org/10.5281/zenodo.376938" httpUri="https://zenodo.org/record/376938/files/figure.png" pageId="9" pageNumber="502">Fig. 60</figureCitation>
).
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2F920FD066014" blockId="9.[151,1437,693,1978]" box="[199,695,1773,1798]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2F920FE0F6014" bold="true" box="[199,446,1773,1798]" pageId="9" pageNumber="502">Measurements (mm)</emphasis>
: Hw 47, abdomen 60.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2F8DCFEE0605C" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2F8DCFE556038" bold="true" box="[199,484,1809,1834]" pageId="9" pageNumber="502">
Variations in
<typeStatus id="DE078869FFCF9B28D352F8DCFE516038" box="[359,480,1809,1834]" pageId="9" pageNumber="502" type="paratype">paratypes</typeStatus>
.
</emphasis>
In one of the
<typeStatus id="DE078869FFCF9B28D04BF8DFFD5B6038" box="[638,746,1810,1834]" pageId="9" pageNumber="502" type="paratype">paratypes</typeStatus>
the tuft of spines at the anterolateral angle of penis segment 3 is hardly visible.
</paragraph>
<paragraph id="010336CBFFCF9B28D2F2F894FE2F6060" blockId="9.[151,1437,693,1978]" box="[199,414,1881,1906]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2F894FE966060" bold="true" box="[199,295,1881,1906]" pageId="9" pageNumber="502">Female:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="49A66540FFCF9B28D2F2F8B0FB5A60A8" pageId="9" pageNumber="502" type="discussion">
<paragraph id="010336CBFFCF9B28D2F2F8B0FB5A60A8" blockId="9.[151,1437,693,1978]" pageId="9" pageNumber="502">
<emphasis id="33C8EAD9FFCF9B28D2F2F8B0FE8A6084" bold="true" box="[199,315,1917,1942]" pageId="9" pageNumber="502">Remarks.</emphasis>
<taxonomicName id="C6BC4D48FFCF9B28D370F8B2FE6E6084" box="[325,479,1917,1942]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="quadratum">
<emphasis id="33C8EAD9FFCF9B28D370F8B2FE6E6084" box="[325,479,1917,1942]" italics="true" pageId="9" pageNumber="502">P. quadratum</emphasis>
</taxonomicName>
shares with
<taxonomicName id="C6BC4D48FFCF9B28D041F8B2FD5E6084" box="[628,751,1917,1942]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="humaita">
<emphasis id="33C8EAD9FFCF9B28D041F8B2FD5E6084" box="[628,751,1917,1942]" italics="true" pageId="9" pageNumber="502">P. humaita</emphasis>
</taxonomicName>
the penis segment 3 subquadrate, with a small tongue-like apical lobe. It differs from it by having a pale apical area only in Hw (in both wings in
<taxonomicName id="C6BC4D48FFCF9B28D651F86FFB6A60A8" box="[1124,1243,1953,1978]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="humaita">
<emphasis id="33C8EAD9FFCF9B28D651F86FFB6A60A8" box="[1124,1243,1953,1978]" italics="true" pageId="9" pageNumber="502">P. humaita</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>