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<document id="A06FBA2B86AC7B2FA0925CCDD63A657D" ID-CLB-Dataset="27606" ID-DOI="10.11646/zootaxa.4551.2.4" ID-GBIF-Dataset="d17f25eb-6875-4fcb-b296-d4b6fe0057b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2622786" ID-ZooBank="3EDA729F-1FAA-4972-8B25-DFD23894F723" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554269821878" checkinUser="plazi" docAuthor="Cruz, Paulo Vilela &amp; Hamada, Neusa" docDate="2019" docId="83330B58FFBDFF9FEFFAFA9FFB0CFBE9" docLanguage="en" docName="zootaxa.4551.2.4.pdf" docOrigin="Zootaxa 4551 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpagobaetis brigada Cruz &amp; Hamada 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="200" masterDocId="7F0A7320FFBFFF9AEF6DFFA1FFBAFF95" masterDocTitle="A new species of Harpagobaetis Mol, with amended diagnosis of the genus, and new considerations on predatory species of Baetidae (Ephemeroptera) from South America" masterLastPageNumber="210" masterPageNumber="195" pageNumber="197" updateTime="1698716063100" updateUser="ExternalLinkService">
<mods:mods id="B03B08A7ED9DC02E11AA0A1F2E3A218D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="92879EAEF54AFD5857A5F3CD994E4245">
<mods:title id="BD0D4CB68F6B54F0707C09DAC70C093F">A new species of Harpagobaetis Mol, with amended diagnosis of the genus, and new considerations on predatory species of Baetidae (Ephemeroptera) from South America</mods:title>
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<mods:name id="3A96D69E109DABEF3F1D8C7623AAA411" type="personal">
<mods:role id="C1B2103D8E69043512509DCB162CE4D2">
<mods:roleTerm id="3E22507E3A6631B6859BD41B1C2DD302">Author</mods:roleTerm>
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<mods:namePart id="A358C23BCFD7AB854F32638C7667B2F3">Cruz, Paulo Vilela</mods:namePart>
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<mods:name id="81C51C319E96A60825325D09A25911FC" type="personal">
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<mods:namePart id="BE4EBDB4C45CBCAF09AD2F216475B3A7">Hamada, Neusa</mods:namePart>
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<mods:typeOfResource id="A7C8C76E99681D3CAD25B7A7A7033984">text</mods:typeOfResource>
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<mods:titleInfo id="E89820943F708705C51CBE151B027006">
<mods:title id="0085766C7A9AA08F310C666FE07C94A6">Zootaxa</mods:title>
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<mods:part id="251292DD868AB11ACC5AB6E249E3CABE">
<mods:date id="6176ACE4DCE87F9D80F3C38FB3A5884C">2019</mods:date>
<mods:detail id="5D19C64FC3C788EFC753FD550A526F99" type="pubDate">
<mods:number id="76B4088983D65C9254EDB83720F9B0DF">2019-01-31</mods:number>
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<mods:detail id="F3AE089E560BBC6ED2D330805A49B01C" type="volume">
<mods:number id="3F9D71117FCEC3F02D3F9560511EC88D">4551</mods:number>
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<mods:number id="145F556C8AB4BFA1B0B8CCA23AFC6236">2</mods:number>
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<mods:start id="2CEFAEA23616CFE7A60C6611AF6517C7">195</mods:start>
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<mods:classification id="1D7CBD184DF53F5145EBF2C9CE803FD2">journal article</mods:classification>
<mods:identifier id="0155E2766864108D72D18B769CEB174D" type="CLB-Dataset">27606</mods:identifier>
<mods:identifier id="8D50E13CBD229AB07ACC6D13041D8375" type="DOI">10.11646/zootaxa.4551.2.4</mods:identifier>
<mods:identifier id="589D7BC2568282B66F20850F86F4C8F8" type="GBIF-Dataset">d17f25eb-6875-4fcb-b296-d4b6fe0057b4</mods:identifier>
<mods:identifier id="AD25308F802D5617E362B734E7C6ED76" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="248FE58CA438D02BE337162337A1F901" type="Zenodo-Dep">2622786</mods:identifier>
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<treatment id="83330B58FFBDFF9FEFFAFA9FFB0CFBE9" ID-DOI="http://doi.org/10.5281/zenodo.5311054" ID-GBIF-Taxon="154533209" ID-Zenodo-Dep="5311054" LSID="urn:lsid:plazi:treatment:83330B58FFBDFF9FEFFAFA9FFB0CFBE9" httpUri="http://treatment.plazi.org/id/83330B58FFBDFF9FEFFAFA9FFB0CFBE9" lastPageId="5" lastPageNumber="200" pageId="2" pageNumber="197">
<subSubSection id="4380E9C5FFBDFF98EFFAFA9FFDABFACD" box="[151,529,1342,1368]" pageId="2" pageNumber="197" type="nomenclature">
<paragraph id="0B25BA4EFFBDFF98EFFAFA9FFDABFACD" blockId="2.[151,529,1342,1402]" box="[151,529,1342,1368]" pageId="2" pageNumber="197">
<heading id="506D0D22FFBDFF98EFFAFA9FFDABFACD" bold="true" box="[151,529,1342,1368]" fontSize="11" level="1" pageId="2" pageNumber="197" reason="1">
<emphasis id="39EE665CFFBDFF98EFFAFA9FFDABFACD" bold="true" box="[151,529,1342,1368]" pageId="2" pageNumber="197">
<taxonomicName id="CC9AC1CDFFBDFF98EFFAFA9FFE17FACD" authority="Cruz &amp; Hamada, 2019" authorityName="Cruz &amp; Hamada" authorityYear="2019" box="[151,429,1342,1368]" class="Insecta" family="Baetidae" genus="Harpagobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="brigada" status="sp. nov.">
<emphasis id="39EE665CFFBDFF98EFFAFA9FFE17FACD" bold="true" box="[151,429,1342,1368]" italics="true" pageId="2" pageNumber="197">Harpagobaetis brigada</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22DDDB27FFBDFF98EEDEFA9FFDABFACD" box="[435,529,1342,1368]" pageId="2" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4380E9C5FFBDFF98EFFAFAC0FEB1FAEF" box="[151,267,1377,1402]" pageId="2" pageNumber="197" type="description">
<paragraph id="0B25BA4EFFBDFF98EFFAFAC0FEB1FAEF" blockId="2.[151,529,1342,1402]" box="[151,267,1377,1402]" pageId="2" pageNumber="197">
(
<figureCitation id="93A1A6CBFFBDFF98EFF2FAC0FEB8FAEF" box="[159,258,1377,1402]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1969,1991]" captionStartId-1="4.[151,250,1434,1456]" captionStartId-2="6.[151,250,1713,1735]" captionStartId-3="8.[151,250,1987,2009]" captionStartId-4="9.[151,250,1961,1983]" captionStartId-5="10.[151,250,1973,1995]" captionTargetBox-0="[281,1312,193,1960]" captionTargetBox-1="[165,1421,195,1416]" captionTargetBox-2="[155,1426,225,1648]" captionTargetBox-3="[163,1425,204,1962]" captionTargetBox-4="[188,1389,203,1936]" captionTargetBox-5="[179,1413,219,1944]" captionTargetId-0="figure@3.[281,1312,193,1960]" captionTargetId-1="figure@4.[165,1423,195,1416]" captionTargetId-2="figure@6.[150,1436,225,1663]" captionTargetId-3="figure@8.[163,1425,193,1977]" captionTargetId-4="figure@9.[178,1420,193,1949]" captionTargetId-5="figure@10.[174,1425,193,1961]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." captionText-1="FIGURE 2. Harpagobaetis brigada sp. nov. A. gut content; B. details of gut content; C. details of cephalic capsule of prey (larva of Chironomidae); D. lateral view of head, predatory habit; E. frontal view of head, predatory habit." captionText-2="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." captionText-3="FIGURE 4. Harpagobaetis brigada sp. nov. Holotype. A. hypopharynx (left v.v., right d.v.); B. labium (left v.v., right d.v.)." captionText-4="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." captionText-5="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri-0="https://zenodo.org/record/2622788/files/figure.png" httpUri-1="https://zenodo.org/record/2622790/files/figure.png" httpUri-2="https://zenodo.org/record/2622793/files/figure.png" httpUri-3="https://zenodo.org/record/2622795/files/figure.png" httpUri-4="https://zenodo.org/record/2622797/files/figure.png" httpUri-5="https://zenodo.org/record/2622799/files/figure.png" pageId="2" pageNumber="197">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4380E9C5FFBDFF98EFFAFA08FB9DF9C7" pageId="2" pageNumber="197" type="diagnosis">
<paragraph id="0B25BA4EFFBDFF98EFFAFA08FB9DF9C7" blockId="2.[151,1437,1449,2014]" pageId="2" pageNumber="197">
<emphasis id="39EE665CFFBDFF98EFFAFA08FE65FA57" bold="true" box="[151,479,1449,1474]" pageId="2" pageNumber="197">Diagnosis. Mature nymph.</emphasis>
1) labrum without setae along lateral margin (
<figureCitation id="93A1A6CBFFBDFF98EB7BFA08FBCEFA57" box="[1046,1140,1449,1474]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3C</figureCitation>
); 2) maxillary palp twosegmented (
<figureCitation id="93A1A6CBFFBDFF98EE72FA6CFECDFA73" box="[287,375,1485,1510]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3E</figureCitation>
); 3) second maxillary canine as long as others (
<figureCitation id="93A1A6CBFFBDFF98ECFBFA6CFC54FA73" box="[918,1006,1485,1510]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3E</figureCitation>
); 4) paraglossa with base overlapping the glossa (
<figureCitation id="93A1A6CBFFBDFF98EE74FA50FEC9F99F" box="[281,371,1521,1546]" captionStart="FIGURE 4" captionStartId="8.[151,250,1987,2009]" captionTargetBox="[163,1425,204,1962]" captionTargetId="figure@8.[163,1425,193,1977]" captionTargetPageId="8" captionText="FIGURE 4. Harpagobaetis brigada sp. nov. Holotype. A. hypopharynx (left v.v., right d.v.); B. labium (left v.v., right d.v.)." httpUri="https://zenodo.org/record/2622795/files/figure.png" pageId="2" pageNumber="197">Fig. 4B</figureCitation>
); 5) middle part of anterior surface of femur without setae (
<figureCitation id="93A1A6CBFFBDFF98EB4DFA50FBC6F99F" box="[1056,1148,1521,1546]" captionStart="FIGURE 5" captionStartId="9.[151,250,1961,1983]" captionTargetBox="[188,1389,203,1936]" captionTargetId="figure@9.[178,1420,193,1949]" captionTargetPageId="9" captionText="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." httpUri="https://zenodo.org/record/2622797/files/figure.png" pageId="2" pageNumber="197">Fig. 5A</figureCitation>
); 6) claws without apical seta (
<figureCitation id="93A1A6CBFFBDFF98EFBFF9B4FE31F9BB" box="[210,395,1557,1582]" captionStart="FIGURE 5" captionStartId="9.[151,250,1961,1983]" captionTargetBox="[188,1389,203,1936]" captionTargetId="figure@9.[178,1420,193,1949]" captionTargetPageId="9" captionText="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." httpUri="https://zenodo.org/record/2622797/files/figure.png" pageId="2" pageNumber="197">Figs 5D, 5E, 5F</figureCitation>
); 7) rows of denticles less than half of length of claw (
<figureCitation id="93A1A6CBFFBDFF98EC98F9B4FBF5F9BB" box="[1013,1103,1557,1582]" captionStart="FIGURE 5" captionStartId="9.[151,250,1961,1983]" captionTargetBox="[188,1389,203,1936]" captionTargetId="figure@9.[178,1420,193,1949]" captionTargetPageId="9" captionText="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." httpUri="https://zenodo.org/record/2622797/files/figure.png" pageId="2" pageNumber="197">Fig. 5D</figureCitation>
); 8) posterior margin of terga IV with rounded projections (
<figureCitation id="93A1A6CBFFBDFF98EE8EF998FD87F9C7" box="[483,573,1593,1618]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="2" pageNumber="197">Fig. 6A</figureCitation>
); 9) gills with tracheae translucid (
<figureCitation id="93A1A6CBFFBDFF98ECAFF998FBA3F9C7" box="[962,1049,1593,1618]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="2" pageNumber="197">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4380E9C5FFBDFF9FEFAAF9FCFE1FFC15" lastPageId="5" lastPageNumber="200" pageId="2" pageNumber="197" type="description">
<paragraph id="0B25BA4EFFBDFF9FEFAAF9FCFE1FFC15" blockId="2.[151,1437,1449,2014]" lastBlockId="5.[151,1437,151,1149]" lastPageId="5" lastPageNumber="200" pageId="2" pageNumber="197">
<emphasis id="39EE665CFFBDFF98EFAAF9FCFDA7F9E3" bold="true" box="[199,541,1629,1654]" pageId="2" pageNumber="197">Description. Mature nymph</emphasis>
(
<figureCitation id="93A1A6CBFFBDFF98ED5DF9FCFD20F9E3" box="[560,666,1629,1654]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1969,1991]" captionStartId-1="4.[151,250,1434,1456]" captionStartId-2="6.[151,250,1713,1735]" captionStartId-3="8.[151,250,1987,2009]" captionStartId-4="9.[151,250,1961,1983]" captionStartId-5="10.[151,250,1973,1995]" captionTargetBox-0="[281,1312,193,1960]" captionTargetBox-1="[165,1421,195,1416]" captionTargetBox-2="[155,1426,225,1648]" captionTargetBox-3="[163,1425,204,1962]" captionTargetBox-4="[188,1389,203,1936]" captionTargetBox-5="[179,1413,219,1944]" captionTargetId-0="figure@3.[281,1312,193,1960]" captionTargetId-1="figure@4.[165,1423,195,1416]" captionTargetId-2="figure@6.[150,1436,225,1663]" captionTargetId-3="figure@8.[163,1425,193,1977]" captionTargetId-4="figure@9.[178,1420,193,1949]" captionTargetId-5="figure@10.[174,1425,193,1961]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." captionText-1="FIGURE 2. Harpagobaetis brigada sp. nov. A. gut content; B. details of gut content; C. details of cephalic capsule of prey (larva of Chironomidae); D. lateral view of head, predatory habit; E. frontal view of head, predatory habit." captionText-2="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." captionText-3="FIGURE 4. Harpagobaetis brigada sp. nov. Holotype. A. hypopharynx (left v.v., right d.v.); B. labium (left v.v., right d.v.)." captionText-4="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." captionText-5="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri-0="https://zenodo.org/record/2622788/files/figure.png" httpUri-1="https://zenodo.org/record/2622790/files/figure.png" httpUri-2="https://zenodo.org/record/2622793/files/figure.png" httpUri-3="https://zenodo.org/record/2622795/files/figure.png" httpUri-4="https://zenodo.org/record/2622797/files/figure.png" httpUri-5="https://zenodo.org/record/2622799/files/figure.png" pageId="2" pageNumber="197">Figs 1–6</figureCitation>
). Body length
<quantity id="CC6217ABFFBDFF98EC21F9FCFC4EF9E0" box="[844,1012,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="3.515" metricValueMax="3.54" metricValueMin="3.49" pageId="2" pageNumber="197" unit="mm" value="3.515" valueMax="3.54" valueMin="3.49">3.49–3.54 mm</quantity>
(n=2); cercus length
<quantity id="CC6217ABFFBDFF98EB98F9FFFA26F9E0" box="[1269,1436,1630,1654]" metricMagnitude="-3" metricUnit="m" metricValue="1.795" metricValueMax="1.81" metricValueMin="1.78" pageId="2" pageNumber="197" unit="mm" value="1.795" valueMax="1.81" valueMin="1.78">1.78–1.81 mm</quantity>
(n=2). Head yellowish (
<figureCitation id="93A1A6CBFFBDFF98EECCF920FE59F90F" box="[417,483,1665,1690]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="2" pageNumber="197">Figs 1</figureCitation>
D–F). Antenna short, 2.0 length of head capsule and with spines and simple setae on the apices of the segments (
<figureCitation id="93A1A6CBFFBDFF98EEBBF904FD88F92B" box="[470,562,1701,1726]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3A</figureCitation>
). Turbinate eye of the nymph with orange-brown surface (
<figureCitation id="93A1A6CBFFBDFF98EBBCF904FA97F92B" box="[1233,1325,1701,1726]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="2" pageNumber="197">Fig. 1D</figureCitation>
). Labrum length about 0.40 the maximum width; distal margin on ventral surface with one row of small blunt setae medially; one row of small spine-like setae continuous from distal margin to distolateral and lateral margins; distal margin on dorsal surface with one row of small thin setae and one row of setae continuous from distal margin to distolateral and lateral margins; dorsal surface covered with thin simple setae (
<figureCitation id="93A1A6CBFFBDFF98ECE8F894FBABF8DB" box="[901,1041,1845,1870]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Figs 3B, 3C</figureCitation>
). Left mandible (
<figureCitation id="93A1A6CBFFBDFF98EBBEF894FA91F8DB" box="[1235,1323,1845,1870]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3D</figureCitation>
) long and narrow, incisor completely fused, with four denticles; prostheca robust, apically with seven denticles; mola shaped like an incisor, tuft of setae between prostheca and mola absent; lateral margins almost straight, bare dorsally. Right mandible (
<figureCitation id="93A1A6CBFFBDFF98EE7DF800FEDCF82F" box="[272,358,1953,1978]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="2" pageNumber="197">Fig. 3F</figureCitation>
) long and narrow, incisor completely fused, with five denticles, prostheca developed, apically with three denticles; mola shaped like an incisor, tuft of setae between prostheca and mola absent; lateral margins almost straight, bare dorsally. Maxilla (
<figureCitation id="93A1A6CBFFBBFF9EED3DFA5AFD1EF981" box="[592,676,1531,1556]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="4" pageNumber="199">Fig. 3E</figureCitation>
) with three canines long and slender plus one distal dentiseta in the same direction; row of setae proximad of canines reduced to two setae and two teeth-like setae; medially, on inner margin, with one spine-like seta. Maxillary palp two-segmented and long, 1.31 length of galea-lacinia; length of segment I 0.17 length of segment II; segments I and II covered with short and thin setae. Hypopharynx (
<figureCitation id="93A1A6CBFFBBFF9EEA56F9C6FA2EF9EA" box="[1339,1428,1639,1664]" captionStart="FIGURE 4" captionStartId="8.[151,250,1987,2009]" captionTargetBox="[163,1425,204,1962]" captionTargetId="figure@8.[163,1425,193,1977]" captionTargetPageId="8" captionText="FIGURE 4. Harpagobaetis brigada sp. nov. Holotype. A. hypopharynx (left v.v., right d.v.); B. labium (left v.v., right d.v.)." httpUri="https://zenodo.org/record/2622795/files/figure.png" pageId="4" pageNumber="199">Fig. 4A</figureCitation>
) large; lingua longer than superlingua, covered with short and thin setae; superlingua with lateral expansion and covered with short and thin setae over inner and distal margin. Labium (
<figureCitation id="93A1A6CBFFBBFF9EEC90F90EFBE2F952" box="[1021,1112,1711,1736]" captionStart="FIGURE 4" captionStartId="8.[151,250,1987,2009]" captionTargetBox="[163,1425,204,1962]" captionTargetId="figure@8.[163,1425,193,1977]" captionTargetPageId="8" captionText="FIGURE 4. Harpagobaetis brigada sp. nov. Holotype. A. hypopharynx (left v.v., right d.v.); B. labium (left v.v., right d.v.)." httpUri="https://zenodo.org/record/2622795/files/figure.png" pageId="4" pageNumber="199">Fig. 4B</figureCitation>
) with paraglossa with base overlapping the glossa; glossa narrow, longer than paraglossa, inner margin bare, distal margin with five robust spine-like setae, outer margin with one row of robust spine-like setae, ventral surface with three simple setae; paraglossa ventral surface with one simple seta, distal margin with four long robust spine-like setae, inner margin with one row of long robust spine-like setae on apical third, outer margin with one row of long robust spine-like setae on apical half. Labial palp three-segmented, with length of segment I 0.61 lengths of segments II and III combined, outer margin with few thin setae; segment II outer margin with long thin setae, inner margin of segment II without distomedial projection; segment III globular-oval, outer margin with many long thin setae. Thorax. Dark brown, fore and hind protoptera with cuticle colorless (
<figureCitation id="93A1A6CBFFBBFF9EEC64F86EFCF6F872" box="[777,844,1999,2024]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="4" pageNumber="199">Figs 1</figureCitation>
A–C). Legs. Femur (
<figureCitation id="93A1A6CBFFBBFF9EEB50F86EFB39F87D" box="[1085,1155,1999,2024]" captionStart="FIGURE 5" captionStartId="9.[151,250,1961,1983]" captionTargetBox="[188,1389,203,1936]" captionTargetId="figure@9.[178,1420,193,1949]" captionTargetPageId="9" captionText="FIGURE 5. Harpagobaetis brigada sp. nov. Holotype. A. foreleg anterior surface; B. detail of dorsal margin of foreleg; C. detail of ventral margin of foreleg; D. foreclaw; E. mid claw; F. hind claw." httpUri="https://zenodo.org/record/2622797/files/figure.png" pageId="4" pageNumber="199">Figs 5</figureCitation>
A–F) yellowish, anterior surface with black mark at base and two yellowish marks, one medially and other apically; anterior surface covered with pores, dorsal margin with one row of small blunt and concave setae (which sometimes resemble spine-like setae), anterior surface near dorsal margin with one row of small blunt setae, anterior surface near ventral margin with one row of small blunt setae, ventral margin bare; tibia with one row of robust spine-like setae on ventral margin and few robust spine-like setae on anterior surface, patella-tibial suture present on all legs and shorter than half the length of tibia; ventral margin of tarsus with one row of robust spine-like setae; claw short, less than the half the length of tarsus, with two rows of triangular denticles increasing in size distally, denticles apically directed, without setae. Mid and hind legs similar to foreleg, except claws with the two apical pairs of denticle disproportionately larger than others. Abdomen with terga I, II, VIII, IX and X white, with brown medial mark (
<figureCitation id="93A1A6CBFFBAFF9FEFF2FE7DFF5EFE61" box="[159,228,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1</figureCitation>
A–C); terga III–VII dark brown (
<figureCitation id="93A1A6CBFFBAFF9FED06FE7DFD0AFE61" box="[619,688,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1</figureCitation>
A–C). Terga surface creased, covered with pores (
<figureCitation id="93A1A6CBFFBAFF9FEB96FE7DFAE3FE61" box="[1275,1369,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1C</figureCitation>
,
<figureCitation id="93A1A6CBFFBAFF9FEA04FE7DFA35FE60" box="[1385,1423,476,501]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="5" pageNumber="200">6A</figureCitation>
), without scales or scale-like setae, posterior margin with irregular blunt projections in segments III–VII, absent in I– II, and almost pointed in VIII-X. Gills yellowish, on segments I–VII, length of gill I (
<figureCitation id="93A1A6CBFFBAFF9FEBFAFD85FB4BFDA8" box="[1175,1265,548,573]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="5" pageNumber="200">Fig. 6B</figureCitation>
) 2.0 length of segment II, length of gill IV (
<figureCitation id="93A1A6CBFFBAFF9FEE83FDE6FDF2FDF5" box="[494,584,583,608]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="5" pageNumber="200">Fig. 6C</figureCitation>
) 1.50 length of segment V; all gills with tracheae translucid (
<figureCitation id="93A1A6CBFFBAFF9FEA63FDE6FAEFFDF5" box="[1294,1365,583,608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1713,1735]" captionTargetBox="[155,1426,225,1648]" captionTargetId="figure@6.[150,1436,225,1663]" captionTargetPageId="6" captionText="FIGURE 3. Harpagobaetis brigada sp. nov. Holotype. A. antenna (medial third); B. labrum (left v.v., right d.v.); C. detail of distal margin of labrum (left v.v., right d.v.); D. left mandible (d.v.); E. maxilla (d.v.); F. right mandible (d.v)." httpUri="https://zenodo.org/record/2622793/files/figure.png" pageId="5" pageNumber="200">Figs 3</figureCitation>
A–C), margins with short and thin setae, surface covered with pores. Paraproct (
<figureCitation id="93A1A6CBFFBAFF9FEC8FFDCDFB84FD11" box="[994,1086,620,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1C</figureCitation>
,
<figureCitation id="93A1A6CBFFBAFF9FEB20FDCDFBCAFD11" box="[1101,1136,620,644]" captionStart="FIGURE 8" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[225,1363,193,1948]" captionTargetId="figure@12.[225,1365,193,1948]" captionTargetPageId="12" captionText="FIGURE 8. A. Map showing the Campos Amazônicos National Park and the savanna area; AM = Amazonas state; MT = Mato Grosso state; RO = Rondônia state. B. habitat of Harpagobaetis brigada sp. nov. C. detail of habitat." httpUri="https://zenodo.org/record/2622803/files/figure.png" pageId="5" pageNumber="200">8D</figureCitation>
) covered with pores, with about 20 spines, postero-lateral extension with marginal spines. Cerci (
<figureCitation id="93A1A6CBFFBAFF9FECD7FD2EFBA2FD3D" box="[954,1048,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1A</figureCitation>
,
<figureCitation id="93A1A6CBFFBAFF9FEB48FD2EFBFCFD3D" box="[1061,1094,655,680]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="5" pageNumber="200">6E</figureCitation>
) white, with dark brown band on apical third, segments with spines on outer margin, posterior margin without spines; primary swimming setae on basal third segments present only in apex of inner margin; all other segments with primary swimming setae all along inner margin, short, colorless; secondary swimming setae absent. Paracercus (
<figureCitation id="93A1A6CBFFBAFF9FEB20FD5DFB11FC81" box="[1101,1195,764,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[281,1312,193,1960]" captionTargetId="figure@3.[281,1312,193,1960]" captionTargetPageId="3" captionText="FIGURE 1. Harpagobaetis brigada sp. nov. A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts." httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="5" pageNumber="200">Figs 1A</figureCitation>
,
<figureCitation id="93A1A6CBFFBAFF9FEBD5FD5DFB6DFC80" box="[1208,1239,764,789]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[179,1413,219,1944]" captionTargetId="figure@10.[174,1425,193,1961]" captionTargetPageId="10" captionText="FIGURE 6. Harpagobaetis brigada sp. nov. Holotype. A. posterior margin of abdominal tergum IV; B. gill I; C. gill IV; D. paraproct; E. paracercus (base section); F. paracercus (medial section); G. cercus (base section); H. (medial section)." httpUri="https://zenodo.org/record/2622799/files/figure.png" pageId="5" pageNumber="200">6F</figureCitation>
) as long as cerci, white, with dark brown band on apical third, segments without spines; primary swimming setae on basal third segments present only in apex of inner margin; all other segments with primary swimming setae all along inner margin, short, colorless.
</paragraph>
</subSubSection>
<caption id="5FE5EAC6FFBCFF99EFFAF810FB57F873" httpUri="https://zenodo.org/record/2622788/files/figure.png" pageId="3" pageNumber="198" startId="3.[151,250,1969,1991]" targetBox="[281,1312,193,1960]" targetPageId="3">
<paragraph id="0B25BA4EFFBCFF99EFFAF810FB57F873" blockId="3.[151,1436,1969,2022]" pageId="3" pageNumber="198">
<emphasis id="39EE665CFFBCFF99EFFAF810FEAFF852" bold="true" box="[151,277,1969,1991]" pageId="3" pageNumber="198">FIGURE 1.</emphasis>
<taxonomicName id="CC9AC1CDFFBCFF99EE73F810FDB4F852" authority="Cruz &amp; Hamada, 2019" authorityName="Cruz &amp; Hamada" authorityYear="2019" box="[286,526,1969,1991]" class="Insecta" family="Baetidae" genus="Harpagobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="brigada" status="sp. nov.">
<emphasis id="39EE665CFFBCFF99EE73F810FDB4F852" box="[286,526,1969,1991]" italics="true" pageId="3" pageNumber="198">Harpagobaetis brigada</emphasis>
</taxonomicName>
<emphasis id="39EE665CFFBCFF99ED7AF810FDD0F852" bold="true" box="[535,618,1969,1991]" pageId="3" pageNumber="198">
<taxonomicNameLabel id="22DDDB27FFBCFF99ED7AF810FDD0F852" box="[535,618,1969,1991]" pageId="3" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. dorsal habitus, B. details of body (dorsal habitus); C. details of abdominal terga and gills; D. frontal view of head; E. lateral view of head; F. frontal view of head, details of mouthparts.
</paragraph>
</caption>
<caption id="5FE5EAC6FFBBFF9EEFFAFA3BFB76FA5B" httpUri="https://zenodo.org/record/2622790/files/figure.png" pageId="4" pageNumber="199" startId="4.[151,250,1434,1456]" targetBox="[165,1421,195,1416]" targetPageId="4">
<paragraph id="0B25BA4EFFBBFF9EEFFAFA3BFB76FA5B" blockId="4.[151,1436,1434,1486]" pageId="4" pageNumber="199">
<emphasis id="39EE665CFFBBFF9EEFFAFA3BFEAEFA25" bold="true" box="[151,276,1434,1456]" pageId="4" pageNumber="199">FIGURE 2.</emphasis>
<taxonomicName id="CC9AC1CDFFBBFF9EEE71FA3BFDB6FA25" authority="Cruz &amp; Hamada, 2019" authorityName="Cruz &amp; Hamada" authorityYear="2019" box="[284,524,1434,1456]" class="Insecta" family="Baetidae" genus="Harpagobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="brigada" status="sp. nov.">
<emphasis id="39EE665CFFBBFF9EEE71FA3BFDB6FA25" box="[284,524,1434,1456]" italics="true" pageId="4" pageNumber="199">Harpagobaetis brigada</emphasis>
</taxonomicName>
<emphasis id="39EE665CFFBBFF9EED79FA3BFDDDFA25" bold="true" box="[532,615,1434,1456]" pageId="4" pageNumber="199">
<taxonomicNameLabel id="22DDDB27FFBBFF9EED79FA3BFDDDFA25" box="[532,615,1434,1456]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. gut content; B. details of gut content; C. details of cephalic capsule of prey (larva of
<taxonomicName id="CC9AC1CDFFBBFF9EEF9EFA19FE33FA5B" box="[243,393,1464,1486]" class="Insecta" family="Chironomidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
); D. lateral view of head, predatory habit; E. frontal view of head, predatory habit.
</paragraph>
</caption>
<subSubSection id="4380E9C5FFBAFF9FEFAAFC2DFABBFC79" pageId="5" pageNumber="200" type="etymology">
<paragraph id="0B25BA4EFFBAFF9FEFAAFC2DFABBFC79" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="200">
<emphasis id="39EE665CFFBAFF9FEFAAFC2DFEF4FC30" bold="true" box="[199,334,908,933]" pageId="5" pageNumber="200">Etymology.</emphasis>
The specific epithet, in apposition, is in honor of the men who fight criminal fire in the Brazilian national parks, especially those in the PNCA, who always joyfully provided logistical support for collection of this new taxon, making our field work pleasant. The name of this fire brigade in Portuguese is &quot;
<emphasis id="39EE665CFFBAFF9FEBFBFC75FB4DFC78" box="[1174,1271,980,1005]" italics="true" pageId="5" pageNumber="200">
<taxonomicName id="CC9AC1CDFFBAFF9FEBFBFC75FB48FC78" authorityName="Cruz &amp; Hamada" authorityYear="2019" box="[1174,1266,980,1005]" class="Insecta" family="Baetidae" genus="Harpagobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="brigada">brigada</taxonomicName>
.
</emphasis>
&quot;
</paragraph>
</subSubSection>
<subSubSection id="4380E9C5FFBAFF9FEFAAFC56FB0CFBE9" pageId="5" pageNumber="200" type="materials_examined">
<paragraph id="0B25BA4EFFBAFF9FEFAAFC56FB0CFBE9" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="200">
<emphasis id="39EE665CFFBAFF9FEFAAFC56FD82FB85" bold="true" box="[199,568,1015,1040]" pageId="5" pageNumber="200">
Material examined.
<materialsCitation id="BBF2B013FFBAFF9FEEAFFC56FD82FB85" ID-GBIF-Occurrence="2234218690" box="[450,568,1015,1040]" collectingDate="2017-07-07" collectionCode="PNCA, INPA" collectorName="P. V. Cruz" country="Brazil" county="Manicore county" latitude="-8.440955" location="Campos Amazonicos" longLatPrecision="1" longitude="-61.660397" municipality="Parque Nacional Campos Amazonicos" pageId="5" pageNumber="200" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<typeStatus id="D42104ECFFBAFF9FEEAFFC56FD89FB85" box="[450,563,1015,1040]" pageId="5" pageNumber="200" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
one nymph mounted on slide,
<collectingCountry id="738DFADEFFBAFF9FECDFFC56FC43FB85" box="[946,1017,1015,1040]" name="Brazil" pageId="5" pageNumber="200">Brazil</collectingCountry>
,
<collectingRegion id="C95E74ACFFBAFF9FEB67FC59FB3AFB85" box="[1034,1152,1016,1040]" country="Brazil" name="Amazonas" pageId="5" pageNumber="200">Amazonas</collectingRegion>
state,
<collectingCounty id="E244C2C2FFBAFF9FEBBEFC56FA22FB85" box="[1235,1432,1015,1040]" pageId="5" pageNumber="200">Manicoré county</collectingCounty>
,
<collectingMunicipality id="EB412034FFBAFF9FEFFAFBBCFDF1FBA0" box="[151,587,1052,1077]" pageId="5" pageNumber="200">
Parque Nacional
<location id="0E45EC95FFBAFF9FEE36FBBDFDF1FBA0" LSID="urn:lsid:plazi:treatment:83330B58FFBDFF9FEFFAFA9FFB0CFBE9:0E45EC95FFBAFF9FEE36FBBDFDF1FBA0" box="[347,587,1052,1077]" country="Brazil" county="Manicore county" latitude="-8.440955" longLatPrecision="1" longitude="-61.660397" municipality="Parque Nacional Campos Amazonicos" name="Campos Amazonicos" pageId="5" pageNumber="200" stateProvince="Amazonas">Campos Amazônicos</location>
</collectingMunicipality>
(PNCA),
<location id="0E45EC95FFBAFF9FEDACFBBDFCBDFBA0" LSID="urn:lsid:plazi:treatment:83330B58FFBDFF9FEFFAFA9FFB0CFBE9:0E45EC95FFBAFF9FEDACFBBDFCBDFBA0" box="[705,775,1052,1077]" country="Brazil" county="Manicore county" latitude="-8.440955" longLatPrecision="1" longitude="-61.660397" municipality="Parque Nacional Campos Amazonicos" name="Veado" pageId="5" pageNumber="200" stateProvince="Amazonas">Veado</location>
stream, tributary of
<location id="0E45EC95FFBAFF9FEC9FFBBCFBF1FBA1" LSID="urn:lsid:plazi:treatment:83330B58FFBDFF9FEFFAFA9FFB0CFBE9:0E45EC95FFBAFF9FEC9FFBBCFBF1FBA1" box="[1010,1099,1053,1076]" country="Brazil" county="Manicore county" latitude="-8.440955" longLatPrecision="1" longitude="-61.660397" municipality="Parque Nacional Campos Amazonicos" name="Macaco" pageId="5" pageNumber="200" stateProvince="Amazonas">Macaco</location>
stream,
<date id="7F249C8EFFBAFF9FEBC3FBBDFA8AFBA0" box="[1198,1328,1052,1077]" pageId="5" pageNumber="200" value="2017-07-07">
<collectingDate id="6F606566FFBAFF9FEBC3FBBDFA8AFBA0" box="[1198,1328,1052,1077]" pageId="5" pageNumber="200" value="2017-07-07">07.vii.2017</collectingDate>
</date>
,
<geoCoordinate id="6EAEDC89FFBAFF9FEA50FBBDFF58FBCD" direction="south" orientation="latitude" pageId="5" pageNumber="200" precision="1" value="-8.440955">S 8° 26’ 27.44”</geoCoordinate>
<geoCoordinate id="6EAEDC89FFBAFF9FEF87FBE1FE0FFBCD" box="[234,437,1087,1112]" direction="west" orientation="longitude" pageId="5" pageNumber="200" precision="1" value="-61.660397">W 61° 39’ 37.42”</geoCoordinate>
,
<collectorName id="A66FDF98FFBAFF9FEEACFBE1FD90FBCD" box="[449,554,1088,1112]" pageId="5" pageNumber="200">P.V. Cruz</collectorName>
col. (INPA).
<materialsCitation id="BBF2B013FFBAFF9FEDAFFB9EFB08FBE9" ID-GBIF-Occurrence="2234218739" collectingDate="2017-07-07" collectionCode="INPA" collectorName="P. V. Cruz" country="Brazil" county="Manicore county" latitude="-8.440955" location="Campos Amazonicos" longLatPrecision="1" longitude="-61.660397" municipality="Parque Nacional Campos Amazonicos" pageId="5" pageNumber="200" specimenCount="3" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="39EE665CFFBAFF9FEDAFFB9EFCF9FBCD" bold="true" box="[706,835,1087,1112]" pageId="5" pageNumber="200">
<typeStatus id="D42104ECFFBAFF9FEDAFFB9EFC84FBCD" box="[706,830,1087,1112]" pageId="5" pageNumber="200" type="paratype">Paratypes</typeStatus>
:
</emphasis>
one nymph mounted on slide, same locality and date as
<typeStatus id="D42104ECFFBAFF9FEFD8FBC5FEADFBE8" box="[181,279,1124,1149]" pageId="5" pageNumber="200" type="holotype">holotype</typeStatus>
(
<collectionCode id="6D8B228BFFBAFF9FEE4BFBC5FED6FBE9" box="[294,364,1124,1148]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="200">INPA</collectionCode>
); three nymphs in 80% ethanol, same locality and date as
<typeStatus id="D42104ECFFBAFF9FEC9EFBC5FBEFFBE8" box="[1011,1109,1124,1149]" pageId="5" pageNumber="200" type="holotype">holotype</typeStatus>
(
<collectionCode id="6D8B228BFFBAFF9FEB09FBC5FB10FBE9" box="[1124,1194,1124,1148]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="200">INPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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