214 lines
23 KiB
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214 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.451.7210" ID-GBIF-Dataset="961a3a60-22ec-4e5e-a3b4-b9cf3715da77" ID-PMC="PMC4258623" ID-Pensoft-Pub="1313-2970-451-109" ID-PubMed="25493050" ID-ZBK="1DA5E3D462414384AFD2472DA3505E5F" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-451-109" ModsDocOrigin="ZooKeys 451" ModsDocTitle="Phylogenetic relationships in the Niviventer-Chiromyscus complex (Rodentia, Muridae) inferred from molecular data, with description of a new species" checkinTime="1451245162554" checkinUser="pensoft" docAuthor="Balakirev, Alexander E., Abramov, Alexei V. & Rozhnov, Viatcheslav V." docDate="2014" docId="82397024122A5A102B76B5B92507BDEA" docLanguage="en" docName="ZooKeys 451: 109-136" docOrigin="ZooKeys 451" docSource="http://dx.doi.org/10.3897/zookeys.451.7210" docTitle="Chiromyscus thomasi Balakirev, Abramov & Rozhnov, 2014, sp. n." docType="treatment" docUuid="8127C488-5D01-4FFC-9556-0986A1198A26" docUuidSource="ZooBank" docVersion="4" lastPageNumber="124" masterDocId="53350837FFD1FFA7FF92FFBA721DF74B" masterDocTitle="Phylogenetic relationships in the Niviventer-Chiromyscus complex (Rodentia, Muridae) inferred from molecular data, with description of a new species" masterLastPageNumber="136" masterPageNumber="109" pageNumber="121" updateTime="1668159542287" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogenetic relationships in the Niviventer-Chiromyscus complex (Rodentia, Muridae) inferred from molecular data, with description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balakirev, Alexander E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Abramov, Alexei V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rozhnov, Viatcheslav V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>451</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>109</mods:start>
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<mods:end>136</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.451.7210</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.451.7210</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-451-109</mods:identifier>
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<mods:identifier type="ZBK">1DA5E3D462414384AFD2472DA3505E5F</mods:identifier>
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<mods:identifier type="ZooBank">1DA5E3D462414384AFD2472DA3505E5F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152056199" LSID="urn:lsid:zoobank.org:act:8127C488-5D01-4FFC-9556-0986A1198A26" httpUri="http://treatment.plazi.org/id/82397024122A5A102B76B5B92507BDEA" lastPageId="15" lastPageNumber="124" pageId="12" pageNumber="121">
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<subSubSection pageId="12" pageNumber="121" type="multiple">
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<paragraph pageId="12" pageNumber="121">Taxon classification Animalia Rodentia Muridae</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="121" type="nomenclature">
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<paragraph pageId="12" pageNumber="121">
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<taxonomicName LSID="http://zoobank.org/8127C488-5D01-4FFC-9556-0986A1198A26" class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="12" pageNumber="121" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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<taxonomicNameLabel pageId="12" pageNumber="121">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="121" type="holotype">
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<paragraph pageId="12" pageNumber="121">Holotype.</paragraph>
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<paragraph pageId="12" pageNumber="121">ZMMU S-191982, body in ethanol, skull extracted, genetic code MC68, adult male, collected 17 December 2011 by Alexander E. Balakirev. GenBank IDs: JQ755933, JQ755964, KF154025, KF154068.</paragraph>
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<caption pageId="12" pageNumber="121">
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<paragraph pageId="12" pageNumber="121">
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Figure 8. The holotype of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="12" pageNumber="121" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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sp. n., Son La Province, northern Vietnam, skull ZMMU S-191982, genetic voucher MC68.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="121" type="type locality">
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<paragraph pageId="12" pageNumber="121">Type locality.</paragraph>
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<paragraph pageId="12" pageNumber="121">
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Vietnam, Son La Province, Muong Thai Village, near Lung Lo pass,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="21.30861">21°18'31"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="104.69278">104°41'34"E</geoCoordinate>
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, elevation ~ 450 m above sea level.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="121" type="paratype">
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<paragraph pageId="12" pageNumber="121">Paratype.</paragraph>
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<paragraph pageId="12" pageNumber="121">ZIN 101651, body in ethanol, skull extracted, genetic code MC80, adult female, collected 17 December 2011 by Alexander E. Balakirev from the type locality. GenBank IDs: JQ755934, JQ755965, KF154069).</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="122" type="referred material">
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<paragraph pageId="13" pageNumber="122">
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<pageBreakToken pageId="13" pageNumber="122" start="start">Referred</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="13" pageNumber="122">BMNH 25.1.1.110, skin and skull, male, Bao Ha, Lao Cai Province, Vietnam; BMNH 26.10.4.167, skin and skull, female, Dak To, Kon Tum Province, Vietnam; BMNH 26.10.4.166, skin, male, Xieng Kuang, Laos.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="123" pageId="13" pageNumber="122" type="diagnosis">
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<paragraph pageId="13" pageNumber="122">Diagnosis.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="123" pageId="13" pageNumber="122">
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This species is set apart from all other described species within the genus
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="14" pageNumber="123" phylum="Chordata" rank="genus">
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<pageBreakToken pageId="14" pageNumber="123" start="start">Chiromyscus</pageBreakToken>
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</taxonomicName>
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by the following combination of morphological traits: (1) Appreciably larger size. This species is the largest in size of any species of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="14" pageNumber="123" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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. Head and body length is 145-180 mm, tail length 200-231 mm, length of hind foot 27-29 mm, ear length 18-20 mm, greatest skull length 41.0-43.0 mm, upper molar lengths 7.0-8.0 mm; the supraorbital ridges are more developed than in other species, forming a distinct pointed triangle shelf at the point where the frontal and palatal bones come into contact. This shelf is very perceptible in the frontal view of the skull. (2) The upper parts are orange-brown. From the face to behind the ears, the pelage is bright orange, with a prominent darker ring around the eye forming a
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<normalizedToken originalValue="“mask”">"mask"</normalizedToken>
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on the face. The under parts are pure white and sharply demarcated from the upper parts. The feet and toes are generally white with orange hairs on top. The tail is bicolored, dark on top and appreciably lighter below, where there is a pinkish hue. The hallux is shortened with rounded nails instead of pointed claws. The species is well differentiated genetically from other
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="14" pageNumber="123" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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. The DNA sequences that are deposited in GenBank under IDs JQ755933-JQ755934, JQ755964-JQ755965, KF154025 and KF154068-KF154069 may be used as genetic vouchers for this species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="123" type="description">
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<paragraph pageId="14" pageNumber="123">Description.</paragraph>
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<paragraph pageId="14" pageNumber="123">
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The fur is dense, smooth and downy. The coloration of the upper side is a bright fulvous with a perceptible orange hue, which is most prominent in the humeral area. On the underside, the belly is pure white without patches or creamy hues. The sides are more brightly colored than the back. The cheek, lateral surface of the neck and the front legs are a bright yellowish-orange. The rump, hips, and base of tail are also, like the cheek, a rich ochraceous color. A very prominent black strip passes over the eye, forming a very characteristic
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<normalizedToken originalValue="“mask”">"mask"</normalizedToken>
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on the face. The vibrissae are long (over 60 mm), both black- and white-colored, and the ears are small (18-20 mm), pale-brown colored and rounded. The dorsal sides of both the front and hind feet are completely buffy-orange. The pads both in the front and hind feet are well developed. The claws are large (4.2-5.0 mm in length), curved and appreciably sharp. The hallux bears a plain nail instead of a claw. The tail is very long, slender and hairy; it is much longer than the body (128-132% of body length). It is rather thick and almost uniformly tinged pale-brown from the proximal part to the tip.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="124" pageId="14" pageNumber="123" type="comparisons">
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<paragraph pageId="14" pageNumber="123">Comparisons.</paragraph>
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<paragraph pageId="15" pageNumber="124">
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">
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<pageBreakToken pageId="15" pageNumber="124" start="start">Chiromyscus</pageBreakToken>
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thomasi
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</taxonomicName>
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is a brightly colored species, a feature that obviously distinguishes it from
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus langbianis" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="langbianis">Chiromyscus langbianis</taxonomicName>
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, which is generally dull in coloration. With its bright fulvous or orange coloration
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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is similar to
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus chiropus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="chiropus">Chiromyscus chiropus</taxonomicName>
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but may be distinguished from it by its dorso-ventral coloration demarcation line. In
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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, the white-colored belly replaces the bright orange ventral side coloration abruptly, without any intermediate zone, whereas a lighter-colored fulvous intermediate zone (0.5-1.0 cm in width) is perceptible on the back sides of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus chiropus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="chiropus">Chiromyscus chiropus</taxonomicName>
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. However, the most apparent distinguishing feature of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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is a dark
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<normalizedToken originalValue="“mask”">"mask"</normalizedToken>
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on the face around the eyes, which may be used to visually separate it from any another
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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or
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<taxonomicName class="Mammalia" family="Muridae" genus="Niviventer" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Niviventer" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Niviventer</taxonomicName>
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species.
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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is the largest species in the genus, appreciably bigger than
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus chiropus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="chiropus">Chiromyscus chiropus</taxonomicName>
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and
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus langbianis" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="langbianis">Chiromyscus langbianis</taxonomicName>
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. Its skull well exceeds the known range of size variation for other
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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as well as for the majority of
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<taxonomicName class="Mammalia" family="Muridae" genus="Niviventer" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Niviventer" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Niviventer</taxonomicName>
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species, with the exception of
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<taxonomicName class="Mammalia" family="Muridae" genus="Niviventer" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Niviventer tenaster" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="tenaster">Niviventer tenaster</taxonomicName>
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and the "
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<taxonomicName class="Insecta" family="Calliphoridae" genus="Muroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Muroidea andersoni" order="Diptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="andersoni">andersoni</taxonomicName>
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" group, both of which are roughly equal in size to, or larger than,
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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. In comparison with other
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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species, the skull of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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is also the most
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<normalizedToken originalValue="“heavily-built”">"heavily-built"</normalizedToken>
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, with supraorbital ridges that are more developed, forming prominent wide shelves. The skull of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus langbianis" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="langbianis">Chiromyscus langbianis</taxonomicName>
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is much smaller and gracile, and the shelves are not so apparent, whereas in
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus chiropus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="chiropus">Chiromyscus chiropus</taxonomicName>
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the skull has an obviously convex profile (when viewed from the side), in contrast with
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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, which appears rather flattened when viewed from the side.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="124" type="etymology">
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<paragraph pageId="15" pageNumber="124">Etymology.</paragraph>
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<paragraph pageId="15" pageNumber="124">
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The new species is named in honor of Oldfield Thomas (1858-1929), the British zoologist who named and described the genus
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="genus">Chiromyscus</taxonomicName>
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and the species
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<taxonomicName class="Insecta" family="Calliphoridae" genus="Muroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Muroidea chiropus" order="Diptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="chiropus">chiropus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="124" type="common name">
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<paragraph pageId="15" pageNumber="124">Common name.</paragraph>
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<paragraph pageId="15" pageNumber="124">Thomas' masked tree rat.</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="124" type="distribution">
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<paragraph pageId="15" pageNumber="124">Distribution.</paragraph>
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<paragraph pageId="15" pageNumber="124">
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Confirmed specimens of
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus thomasi" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="thomasi">Chiromyscus thomasi</taxonomicName>
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have been recorded from the provinces of Son La and Lao Cai in northern Vietnam, the provinces of Kon Tum and Nhge An in central Vietnam, and the provinces of Xieng Khouang and Luang Prabang in northern Laos, based on published data and our (BAE) most recent and unpublished data. This species may have a wider distribution in central Vietnam (
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<bibRefCitation author="Dang, Huy Huynh" journalOrPublisher="Publishing House " Science and Technics ", Hanoi" pageId="18" pageNumber="127" title="Checklist of Mammals in Vietnam." year="1994">Dang Huy Huynh et al. 1994</bibRefCitation>
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,
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<bibRefCitation author="Dang, Ngoc Can" journalOrPublisher="Institute of Ecology and Biological Resources, Hanoi" pageId="17" pageNumber="126" title="Checklist of Wild Mammal Species of Vietnam." year="2008">Dang Ngoc Can et al. 2008</bibRefCitation>
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) and in northern and central Laos (
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<bibRefCitation author="Aplin, K" journalOrPublisher="IUCN Red List of Threatened Species" pageId="17" pageNumber="126" title="Chiromyscuschiropus." url="www.iucnredlist.org" volumeTitle="IUCN 2011" year="2008">Aplin et al. 2008</bibRefCitation>
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,
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<bibRefCitation author="Musser, GG" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="20" pageNumber="129" pagination="225 - 334" title="Results of the Archbold Expeditions. No. 105. Notes on systematics of Indo-Malayan murid rodents, and descriptions of new genera and species from Ceylon, Sulawesi and the Philippines." volume="168" year="1981">Musser 1981</bibRefCitation>
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,
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<bibRefCitation author="Corbet, GB" journalOrPublisher="Oxford University Press, Oxford" pageId="18" pageNumber="127" title="The Mammals of the Indo-Malayan Region: A Systematic Review." year="1992">Corbet and Hill 1992</bibRefCitation>
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) where similar
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<normalizedToken originalValue="“mask-bearing”">"mask-bearing"</normalizedToken>
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specimens have been reported. It is also likely distributed in south-western China (see
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<bibRefCitation author="Wang, YX" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="21" pageNumber="130" title="A Complete Checklist of Mammals Species and Subspecies of China: A Taxonomic and Geographic References." year="2003">Wang 2003</bibRefCitation>
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) and northern Thailand (see
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<bibRefCitation author="Marshall, JT Jr" editor="Lekagul, B" journalOrPublisher="Government Printing Office, Kurusapha Ladprao, Bangkok" pageId="19" pageNumber="128" pagination="396 - 487" title="Family Muridae (rats and mice)." volumeTitle="Mammals of Thailand" year="1977">Marshall 1977</bibRefCitation>
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) but clarifying comparisons are needed to rule out alternative identifications (
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus chiropus" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="chiropus">Chiromyscus chiropus</taxonomicName>
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and
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<taxonomicName class="Mammalia" family="Muridae" genus="Chiromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chiromyscus langbianis" order="Rodentia" pageId="15" pageNumber="124" phylum="Chordata" rank="species" species="langbianis">Chiromyscus langbianis</taxonomicName>
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) before this wider potential geographic distribution is confirmed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |