treatments-xml/data/7F/AF/B9/7FAFB90D4BDE5AABA686071EA921BF47.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.98.90858" ID-Pensoft-Pub="1860-0743-2-377" ID-Pensoft-UUID="1AC39832654D5858A7369F63BD7C36F1" ID-ZooBank="CE7829AAD0304C18B7433712FE4D3128" ModsDocID="1860-0743-98-2-377" checkinTime="1663699690556" checkinUser="pensoft" docAuthor="Ballarin, Francesco &amp; Pantini, Paolo" docDate="2022" docId="7FAFB90D4BDE5AABA686071EA921BF47" docLanguage="en" docName="ZoosystEvol 98(2): 377-385" docOrigin="Zoosystematics and Evolution 98 (2)" docPubDate="2022-09-20" docSource="http://dx.doi.org/10.3897/zse.98.90858" docTitle="Cybaeopsis lodovicii Ballarin &amp; Pantini 2022, sp. nov." docType="treatment" docUuid="FF26EBC0-C226-4264-AC50-EFEC64D7F86A" docUuidSource="ZooBank" docVersion="1" id="1AC39832654D5858A7369F63BD7C36F1" lastPageNumber="377" masterDocId="1AC39832654D5858A7369F63BD7C36F1" masterDocTitle="An unexpected occurrence: discovery of the genus Cybaeopsis Strand, 1907 in Europe with the description of a new species from Italy (Arachnida, Araneae, Amaurobiidae)" masterLastPageNumber="385" masterPageNumber="377" pageNumber="377" updateTime="1663700011223" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>An unexpected occurrence: discovery of the genus Cybaeopsis Strand, 1907 in Europe with the description of a new species from Italy (Arachnida, Araneae, Amaurobiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ballarin, Francesco</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1417-2519</mods:nameIdentifier>
<mods:affiliation>Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji-shi, Tokyo, 192 - 0397, Japan &amp; Department of Zoology, Museo di Storia Naturale of Verona, Lungadige Porta Vittoria, 9, I- 37129 Verona, Italy</mods:affiliation>
<mods:nameIdentifier type="email">ballarin.francesco@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pantini, Paolo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8332-1915</mods:nameIdentifier>
<mods:affiliation>Museo Civico di Scienze Naturali &quot; E. Caffi &quot; of Bergamo, Piazza Cittadella 10, I- 24129 Bergamo, Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-20</mods:number>
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<mods:detail type="volume">
<mods:number>98</mods:number>
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<mods:number>2</mods:number>
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<mods:start>377</mods:start>
<mods:end>385</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.98.90858</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.90858</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-2-377</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:FF26EBC0-C226-4264-AC50-EFEC64D7F86A" httpUri="http://treatment.plazi.org/id/7FAFB90D4BDE5AABA686071EA921BF47" lastPageNumber="377" pageId="0" pageNumber="377">
<subSubSection pageId="0" pageNumber="377" type="nomenclature">
<paragraph pageId="0" pageNumber="377">
<taxonomicName LSID="https://zoobank.org/FF26EBC0-C226-4264-AC50-EFEC64D7F86A" authority="Ballarin &amp; Pantini, 2022" authorityName="Ballarin &amp; Pantini" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Cybaeopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybaeopsis lodovicii" order="Araneae" pageId="0" pageNumber="377" phylum="Arthropoda" rank="species" species="lodovicii" status="sp. nov.">Cybaeopsis lodovicii</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="377">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="description">
<paragraph pageId="0" pageNumber="377">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cybaeopsis lodovicii sp. nov. male palp. A prolateral view; B retrolateral view; C ventral view; D dorsal view; E tibial, dorsal view; F SEM image of the tibial apophyses, dorsal view; G SEM image of detail of the RTA tip; H SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1 - 3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (A-D); 0.1 mm (F); 0.02 mm (G); 0.05 mm (H)." figureDoi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377">Figs 1A-H</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cybaeopsis lodovicii sp. nov., male and female. A epigyne, ventral view; B ditto, ventral view after maceration; C ditto, posterior view; D vulva, dorsal view; E male habitus, dorsal view; F female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E, F)." figureDoi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377">, 2A-F</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">, 3A-D</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cybaeopsis euopla, male and female from Ontario. A male habitus, dorsal view; B female habitus, dorsal view; C epigyne, ventral view; D male palp, retrolateral view; E ditto, ventral view; F original sample labels; G distribution of the genus Cybaeopsis. Scale bars: 1 mm (A, B); 0.5 mm (C-E)." figureDoi="10.3897/zse.98.90858.figure5" httpUri="https://binary.pensoft.net/fig/746028" pageId="0" pageNumber="377">, 5G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="type material">
<paragraph pageId="0" pageNumber="377">Type material.</paragraph>
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">
<emphasis italics="true" pageId="0" pageNumber="377">Holotype</emphasis>
♂ ITALY: Liguria
</emphasis>
: Genova, Mezzanego, Giaiette, 850 m, (
<geoCoordinate degrees="44" direction="north" minutes="25" orientation="latitude" precision="15" seconds="03" value="44.4175">44°25'03&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="9" direction="east" minutes="28" orientation="longitude" precision="15" seconds="08" value="9.468888">9°28'08&quot;E</geoCoordinate>
), beechwood, pitfall trap, 31 Oct. 2009-25 May 2010, O. Lodovici, P. Pantini &amp; M. Valle leg.
</paragraph>
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">
<emphasis italics="true" pageId="0" pageNumber="377">Paratypes</emphasis>
: ITALY: Liguria
</emphasis>
: 1♂, 4♀♀, same data as the holotype; 2♀♀, same locality, 25 May-18 Aug. 2010 • 1♀, Foresta Demaniale Monte Zatta, ex colonia Devoto, 1050 m, beechwood, pitfall traps 31 Oct 2009-25 May 2010 • 2♀♀, same locality, 25 May-18 Aug. 2010, all O. Lodovici, P. Pantini &amp; M. Valle leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="etymology">
<paragraph pageId="0" pageNumber="377">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="377">The specific epithet is a patronym in honor of our colleague and friend Omar Lodovici (Museo Civico di Scienze Naturali of Bergamo, Italy).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="diagnosis">
<paragraph pageId="0" pageNumber="377">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="377">
Male of the new species can be distinguished from male of 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="C. typicus" pageId="0" pageNumber="377" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">C. typicus</emphasis>
</taxonomicName>
by the different shape and number of the dorsal apophyses of the palpal tibia (Da): four Da with Da2 and Da3 long and Da2 ending wider in 
<taxonomicName lsidName="C. lodovicii" pageId="0" pageNumber="377" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">C. lodovicii</emphasis>
</taxonomicName>
sp. nov. (vs. only three Da in 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
and Da2 shorter with a sharp end and Da3 very short in both 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="C. typicus" pageId="0" pageNumber="377" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">C. typicus</emphasis>
</taxonomicName>
). In addition, 
<taxonomicName lsidName="C. lodovicii" pageId="0" pageNumber="377" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">C. lodovicii</emphasis>
</taxonomicName>
sp. nov. can be distinguished by the absence of strongly protruding prolateral and retrolateral tegular outgrowths (Pto and Rto) (vs. strongly protruding Pto in 
<taxonomicName lsidName="C. typicus" pageId="0" pageNumber="377" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">C. typicus</emphasis>
</taxonomicName>
or Rto in 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
) (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cybaeopsis lodovicii sp. nov. male palp. A prolateral view; B retrolateral view; C ventral view; D dorsal view; E tibial, dorsal view; F SEM image of the tibial apophyses, dorsal view; G SEM image of detail of the RTA tip; H SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1 - 3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (A-D); 0.1 mm (F); 0.02 mm (G); 0.05 mm (H)." figureDoi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377">1A-D</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3A, B</figureCitation>
cf. Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cybaeopsis typicus, male and female from Kunashir Island. A male palp, retrolateral view; B ditto, prolateral view; C ditto, ventral view; D ditto, dorsal view; E epigyne, ventral view; F ditto, posterior view. Photos by Y. M. Marusik Abbreviations: Co = conductor; Da 1 - 4 = dorsal apophyses of tibia; Em = embolus; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Va = ventral apophysis." figureDoi="10.3897/zse.98.90858.figure4" httpUri="https://binary.pensoft.net/fig/746027" pageId="0" pageNumber="377">4A-D</figureCitation>
and 
<bibRefCitation DOI="https://doi.org/10.13156/arac.2021.18.8.873" author="Bosmans, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="377" pagination="873 - 882" refId="B3" refString="Bosmans, R, 2021. Notes on Amaurobiidae (Araneae) of the Western Mediterranean region, with the description of a new species. Arachnology 18 (8): 873 - 882, DOI: https://doi.org/10.13156/arac.2021.18.8.873" title="Notes on Amaurobiidae (Araneae) of the Western Mediterranean region, with the description of a new species." url="https://doi.org/10.13156/arac.2021.18.8.873" volume="18" year="2021">Bosmans 2021</bibRefCitation>
: figs 35, 36). Female of 
<taxonomicName lsidName="C. lodovicii" pageId="0" pageNumber="377" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">C. lodovicii</emphasis>
</taxonomicName>
sp. nov. can be distinguished from female of 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="C. typicus" pageId="0" pageNumber="377" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">C. typicus</emphasis>
</taxonomicName>
by the different shape of the internal pockets (Ip) and lateral lobes (Ll): rectangular Ll with comma-like and laterally elongated Ip (vs. shorter, more squared Ll with shorter and stockier Ip in 
<taxonomicName lsidName="C. theoblicki" pageId="0" pageNumber="377" rank="species" species="theoblicki">
<emphasis italics="true" pageId="0" pageNumber="377">C. theoblicki</emphasis>
</taxonomicName>
or more rounded Ll and Ip in 
<taxonomicName lsidName="C. typicus" pageId="0" pageNumber="377" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">C. typicus</emphasis>
</taxonomicName>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cybaeopsis lodovicii sp. nov., male and female. A epigyne, ventral view; B ditto, ventral view after maceration; C ditto, posterior view; D vulva, dorsal view; E male habitus, dorsal view; F female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E, F)." figureDoi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377">2A, B, D</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3C</figureCitation>
cf. 
<bibRefCitation DOI="https://doi.org/10.13156/arac.2021.18.8.873" author="Bosmans, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="377" pagination="873 - 882" refId="B3" refString="Bosmans, R, 2021. Notes on Amaurobiidae (Araneae) of the Western Mediterranean region, with the description of a new species. Arachnology 18 (8): 873 - 882, DOI: https://doi.org/10.13156/arac.2021.18.8.873" title="Notes on Amaurobiidae (Araneae) of the Western Mediterranean region, with the description of a new species." url="https://doi.org/10.13156/arac.2021.18.8.873" volume="18" year="2021">Bosmans 2021</bibRefCitation>
: figs 34-41).
</paragraph>
<paragraph pageId="0" pageNumber="377">
The different number and shape of Da and Ll quickly distinguish male and female 
<taxonomicName lsidName="C. lodovicii" pageId="0" pageNumber="377" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">C. lodovicii</emphasis>
</taxonomicName>
sp. nov. from the American congeners (for comparison see 
<bibRefCitation DOI="https://doi.org/10.4039/entm10484fv" author="Leech, RE" journalOrPublisher="Arachnology" pageId="0" pageNumber="377" refId="B4" refString="Leech, RE, 1972. A revision of the Nearctic Amaurobiidae (Arachnida: Araneida). Memoirs of the Entomological Society of Canada 84(S84): 1-182. https://doi.org/10.4039/entm10484fv" title="A revision of the Nearctic Amaurobiidae (Arachnida: Araneida). Memoirs of the Entomological Society of Canada 84 (S 84): 1 - 182." url="https://doi.org/10.4039/entm10484fv" year="1972">Leech 1972</bibRefCitation>
: figs 85-101).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="description">
<paragraph pageId="0" pageNumber="377">Description.</paragraph>
<paragraph pageId="0" pageNumber="377">
(the specimens are in rather poor condition and the coloration may be different in freshly collected samples). 
<emphasis bold="true" pageId="0" pageNumber="377">
Male (
<emphasis italics="true" pageId="0" pageNumber="377">holotype</emphasis>
).
</emphasis>
Habitus as in Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cybaeopsis lodovicii sp. nov., male and female. A epigyne, ventral view; B ditto, ventral view after maceration; C ditto, posterior view; D vulva, dorsal view; E male habitus, dorsal view; F female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E, F)." figureDoi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377">2E</figureCitation>
. Total length: 3.25. Carapace 1.55 long, 0.95 wide. Carapace brownish with dorsal slightly darker radiating strips, fovea clearly visible. Cephalic area as in Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3D</figureCitation>
, darker than thoracic part. Chelicerae dark brown, frontally swollen with 3 posterior and 6 anterior teeth. Eyes sizes and their interdistances: AME = 0.07, ALE = 0.1, PME = 0.06, PLE = 0.1, AME-ALE = 0.01, PME-PLE = 0.08. Legs uniformly brownish. Length of legs segments (for legs II-IV only femora are available for the measurements): I (0.96, 0.44, 0.88, 0.83, 0.61), II (0.95, -), III (0.93, -), IV (1.13, -). Leg formula: IV, I, II, III. Spination as in Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Leg spination of male and female C. lodovicii sp. nov. Variability in number of spines among right and left legs is reported in parentheses. Abbreviations: d = dorsal spines, p = prolateral spines, r = retrolateral spines; v = ventral spines." httpUri="http://table.plazi.org/id/8061947750521AB93CFBB88BB8324C77" pageId="0" pageNumber="377" tableUuid="8061947750521AB93CFBB88BB8324C77">1</tableCitation>
. Opisthosoma brown-greyish with lighter chevrons marks on the dorsal side. Palp as in Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cybaeopsis lodovicii sp. nov. male palp. A prolateral view; B retrolateral view; C ventral view; D dorsal view; E tibial, dorsal view; F SEM image of the tibial apophyses, dorsal view; G SEM image of detail of the RTA tip; H SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1 - 3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (A-D); 0.1 mm (F); 0.02 mm (G); 0.05 mm (H)." figureDoi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377">1A-H</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3A, B</figureCitation>
. Palpal femur and patella light brown-yellowish, tibia and tarsus dark brown. Femur approx. 1 and half the length of tibia. Tibia with several long retrolateral setae and 4 dorsal apophyses (Da1-4, Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cybaeopsis lodovicii sp. nov. male palp. A prolateral view; B retrolateral view; C ventral view; D dorsal view; E tibial, dorsal view; F SEM image of the tibial apophyses, dorsal view; G SEM image of detail of the RTA tip; H SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1 - 3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (A-D); 0.1 mm (F); 0.02 mm (G); 0.05 mm (H)." figureDoi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377">1E, F</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3A</figureCitation>
). Da1 long and curved heading first retrolaterally and then frontally, proximal part wider and flatter, distal part thin and sharp. Da2 long and thin, headed frontal-retrolaterally and curving ventrally toward the RTA, tip enlarged and with a small notch ending sharply (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cybaeopsis lodovicii sp. nov. male palp. A prolateral view; B retrolateral view; C ventral view; D dorsal view; E tibial, dorsal view; F SEM image of the tibial apophyses, dorsal view; G SEM image of detail of the RTA tip; H SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1 - 3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (A-D); 0.1 mm (F); 0.02 mm (G); 0.05 mm (H)." figureDoi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377">1G</figureCitation>
). Da3 thin and sharp, headed frontally with an S-shaped course, Da4 short and sharp, headed frontally. RTA elongated and sturdy, approx. as long as tibia, bent dorsally, slightly forked at the tip. Ventral tibial apophysis (Va) short and blunt. Cymbium as long as tibia. Bulb round and dorsoventrally flattened. Median apophysis (Ma) of bulb trapezoid and sturdy with 2 blunt subdivisions headed posteriorly and retrolaterally. Prolateral tegular outgrowth (Pto) inconspicuous. Retrolateral tegular outgrowth (Rto) wide and flat. Both tegular outgrowths only slightly protruding from tegulum. Conductor (Co) wide. Embolus (Em) short and sturdy, ribbon-like, approx. as long as the conductor, distal part with a slightly S-shaped course (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3A, B</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="8061947750521AB93CFBB88BB8324C77" httpUri="http://table.plazi.org/id/8061947750521AB93CFBB88BB8324C77" pageId="0" pageNumber="377" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Table 1.</emphasis>
Leg spination of male and female 
<taxonomicName lsidName="C. lodovicii" pageId="0" pageNumber="377" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">C. lodovicii</emphasis>
</taxonomicName>
sp. nov. Variability in number of spines among right and left legs is reported in parentheses. Abbreviations: d = dorsal spines, p = prolateral spines, r = retrolateral spines; v = ventral spines.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="377">
<table pageId="0" pageNumber="377">
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Male</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Femur</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Tibia</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Metatarsus</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">I</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1(2) r1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d2 p2 r2 V2-2</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d2 p3 r3 v2-2-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">II</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p1(2) v2-2</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p2 v2-2-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">III</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d2 p1 r1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p2 r2 v1-2-2</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d2 p3 r3 v2-2-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">IV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1(2) r1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d2 p2 r2 v1-2-2</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d3 p3 r3 v2-2-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Female</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Femur</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Tibia</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">Metatarsus</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">I</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p1 v1-2-1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p1 v2-1-1</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">II</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p1 v1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">v1-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">III</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1(2) p1 r1</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p2 r2 v1-(1)</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p3 r3 v2-3</td>
</tr>
<tr pageId="0" pageNumber="377">
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="377">IV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1(2)</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">d1 p1 r2 v1-2-2</td>
<td colspan="1" pageId="0" pageNumber="377" rowspan="1">p3 r3 v2-1-3</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zse.98.90858.figure1" httpUri="https://binary.pensoft.net/fig/746024" pageId="0" pageNumber="377" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Figure 1.</emphasis>
<taxonomicName authorityName="Ballarin &amp; Pantini" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Cybaeopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybaeopsis lodovicii" order="Araneae" pageId="0" pageNumber="377" phylum="Arthropoda" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">Cybaeopsis lodovicii</emphasis>
</taxonomicName>
sp. nov. male palp. 
<emphasis bold="true" pageId="0" pageNumber="377">A</emphasis>
prolateral view; 
<emphasis bold="true" pageId="0" pageNumber="377">B</emphasis>
retrolateral view; 
<emphasis bold="true" pageId="0" pageNumber="377">C</emphasis>
ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">D</emphasis>
dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">E</emphasis>
tibial, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">F</emphasis>
SEM image of the tibial apophyses, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">G</emphasis>
SEM image of detail of the RTA tip; 
<emphasis bold="true" pageId="0" pageNumber="377">H</emphasis>
SEM image of detail of the median apophysis, ventral view. Abbreviations: Da 1-3 = dorsal apophyses; RTA = retrolateral tibial apophysis; Va = ventral apophysis. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="377">A-D</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="377">F</emphasis>
); 0.02 mm (
<emphasis bold="true" pageId="0" pageNumber="377">G</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="377">H</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Figure 2.</emphasis>
<taxonomicName authorityName="Ballarin &amp; Pantini" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Cybaeopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybaeopsis lodovicii" order="Araneae" pageId="0" pageNumber="377" phylum="Arthropoda" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">Cybaeopsis lodovicii</emphasis>
</taxonomicName>
sp. nov., male and female. 
<emphasis bold="true" pageId="0" pageNumber="377">A</emphasis>
epigyne, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">B</emphasis>
ditto, ventral view after maceration; 
<emphasis bold="true" pageId="0" pageNumber="377">C</emphasis>
ditto, posterior view; 
<emphasis bold="true" pageId="0" pageNumber="377">D</emphasis>
vulva, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">E</emphasis>
male habitus, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">F</emphasis>
female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="377">A-D</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="377">E, F</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Figure 3.</emphasis>
<taxonomicName authorityName="Ballarin &amp; Pantini" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Cybaeopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybaeopsis lodovicii" order="Araneae" pageId="0" pageNumber="377" phylum="Arthropoda" rank="species" species="lodovicii">
<emphasis italics="true" pageId="0" pageNumber="377">Cybaeopsis lodovicii</emphasis>
</taxonomicName>
sp. nov., male and female. 
<emphasis bold="true" pageId="0" pageNumber="377">A</emphasis>
male palp, retrolateral view; 
<emphasis bold="true" pageId="0" pageNumber="377">B</emphasis>
ditto, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">C</emphasis>
epigyne, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">D</emphasis>
cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1-4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="377">A-C</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="377">D</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zse.98.90858.figure4" httpUri="https://binary.pensoft.net/fig/746027" pageId="0" pageNumber="377" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Figure 4.</emphasis>
<taxonomicName class="Arachnida" family="Amaurobiidae" genus="Cybaeopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybaeopsis typicus" order="Araneae" pageId="0" pageNumber="377" phylum="Arthropoda" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="377">Cybaeopsis typicus</emphasis>
</taxonomicName>
, male and female from Kunashir Island. 
<emphasis bold="true" pageId="0" pageNumber="377">A</emphasis>
male palp, retrolateral view; 
<emphasis bold="true" pageId="0" pageNumber="377">B</emphasis>
ditto, prolateral view; 
<emphasis bold="true" pageId="0" pageNumber="377">C</emphasis>
ditto, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">D</emphasis>
ditto, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="377">E</emphasis>
epigyne, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="377">F</emphasis>
ditto, posterior view. Photos by Y.M. Marusik Abbreviations: Co = conductor; Da 1-4 = dorsal apophyses of tibia; Em = embolus; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Va = ventral apophysis.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="377">
<emphasis bold="true" pageId="0" pageNumber="377">Female.</emphasis>
Habitus as in Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cybaeopsis lodovicii sp. nov., male and female. A epigyne, ventral view; B ditto, ventral view after maceration; C ditto, posterior view; D vulva, dorsal view; E male habitus, dorsal view; F female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E, F)." figureDoi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377">2F</figureCitation>
. Total length: 3.04. Carapace 1.35 long, 0.83 wide. Coloration as in male, chevrons on the opisthosoma more visible than in the male. Chelicera with 3 posterior and 5 anterior teeth. Eyes sizes and their interdistances: AME = 0.04, ALE = 0.1, PME = 0.07, PLE = 0.12, AME-ALE = 0.02, PME-PLE = 0.13. Length of legs segments (leg III missing): I 2.94 (0.86, 0.4, 0.65, 0.57, 0.46), II 2.68 (0.96, 0.25, 0.57, 0.53, 0.37), III (-), IV 3.04 (0.85, 0.4, 0.67, 0.72, 0.4).
</paragraph>
<paragraph pageId="0" pageNumber="377">
Leg spination as in Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Leg spination of male and female C. lodovicii sp. nov. Variability in number of spines among right and left legs is reported in parentheses. Abbreviations: d = dorsal spines, p = prolateral spines, r = retrolateral spines; v = ventral spines." httpUri="http://table.plazi.org/id/8061947750521AB93CFBB88BB8324C77" pageId="0" pageNumber="377" tableUuid="8061947750521AB93CFBB88BB8324C77">1</tableCitation>
. Calamistrum clearly visible, about 2/3 of metatarsus length. Other characters as in male. Epigyne as in Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cybaeopsis lodovicii sp. nov., male and female. A epigyne, ventral view; B ditto, ventral view after maceration; C ditto, posterior view; D vulva, dorsal view; E male habitus, dorsal view; F female habitus, dorsal view. Abbreviations: Cd = copulatory duct; Co = copulatory opening; Ip = internal pocket; Ll = lateral lobe; Sp = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E, F)." figureDoi="10.3897/zse.98.90858.figure2" httpUri="https://binary.pensoft.net/fig/746025" pageId="0" pageNumber="377">2A-D</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cybaeopsis lodovicii sp. nov., male and female. A male palp, retrolateral view; B ditto, ventral view; C epigyne, ventral view; D cephalic area of female, frontal view. Abbreviations: Cn = conductor; Da 1 - 4 = dorsal apophyses of Tibia; Em = embolus; Ip = internal pocket; Ll = lateral lobe; Ma = median apophysis; Pto = prolateral tegular outgrowth; RTA = retrolateral tibial apophysis; Rto = retrolateral tegular outgrowth; St = subtegulum; Te = tegulum; Ti = tibia; Va = ventral apophysis. Scale bars: 0.2 mm (A-C); 0.5 mm (D)." figureDoi="10.3897/zse.98.90858.figure3" httpUri="https://binary.pensoft.net/fig/746026" pageId="0" pageNumber="377">3C</figureCitation>
. Epigynal plate divided into two lateral lobes (Ll) and a septum. Copulatory openings (Co) located in the antero-median inner part of lateral lobes. Internal pockets (Ip) of lateral lobes wide, comma-like, narrowing laterally, visible by transparency through tegument of epigyne. Copulatory ducts (Cd) short, comma-like, proximal traits parallel to each other in medial part of vulva and then diverging laterally. Spermathecae (Sp) small, separated from each other more than 4 times their diameter, located in anterior side of vulva.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="distribution">
<paragraph pageId="0" pageNumber="377">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="377">Endemic to Northern Apennines, Italy.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="377" type="habitat">
<paragraph pageId="0" pageNumber="377">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="377">Litter of mountain beechwoods at mid-elevation (~800 m).</paragraph>
</subSubSection>
</treatment>
</document>