262 lines
40 KiB
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262 lines
40 KiB
XML
<document ID-CLB-Dataset="269602" ID-DOI="http://dx.doi.org/10.3897/dez.70.107425" ID-GBIF-Dataset="360868d3-9133-466c-bfae-e8de48028017" ID-Pensoft-Pub="1860-1324-2-311" ID-Pensoft-UUID="65D0B2390EBE553382C9727ECC9CBF8F" ID-ZooBank="8D3E277C424C440B8FE878085239C2A2" ModsDocID="1860-1324-70-2-311" checkinTime="1695893248727" checkinUser="pensoft" docAuthor="Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek & Schmidt, Joachim" docDate="2023" docId="78E8D3AE40CD5B96A44FFBA1906E2361" docLanguage="en" docName="DeutEntomolZeit 70(2): 311-335" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-09-27" docSource="http://dx.doi.org/10.3897/dez.70.107425" docTitle="Baehria Schmidt & Faille, gen. nov." docType="treatment" docVersion="4" id="65D0B2390EBE553382C9727ECC9CBF8F" lastPageNumber="311" masterDocId="65D0B2390EBE553382C9727ECC9CBF8F" masterDocTitle="Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)" masterLastPageNumber="335" masterPageNumber="311" pageNumber="311" updateTime="1732677706243" updateUser="ExternalLinkService">
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<mods:title id="127712B4764AE430745DC9D4DD6FA3CF">Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)</mods:title>
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<mods:namePart id="FFA05D47E104FDDD8BF6065EDF80A2FB">Faille, Arnaud</mods:namePart>
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<mods:affiliation id="52AA0DBEEF5596219752E16B687739E6">Department of Entomology, Stuttgart State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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<mods:nameIdentifier id="23A1FB6E6134732791F2DEAFBDC6DC36" type="email">arnaud.faille@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="B75479A00B2BCA61F251901971727828">Hofmann, Sylvia</mods:namePart>
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<mods:affiliation id="B5D13F97F2A5CE4BFEA80C23AD7CB105">Leibniz Institute for the Analysis of Biodiversity Change, Museum Koenig, 53113 Bonn, Germany</mods:affiliation>
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<mods:namePart id="A66BDC55C400C72D4BE76327805D61E2">Merene, Yeshitla</mods:namePart>
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<mods:nameIdentifier id="1CE3FC2D9BD329C833A5B3FF99DB8E79" type="ORCID">https://orcid.org/0000-0003-2998-822X</mods:nameIdentifier>
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<mods:affiliation id="EC17BF65F312CDA8BE2C678255D87EBD">Faculty of Biology, Philipps University Marburg, Marburg, Germany & Department of Zoological Sciences, Addis Ababa University, Addis Ababa, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
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<mods:namePart id="C96DFB495B0D97DD7B183B9F4E886F70">Hauth, David</mods:namePart>
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<mods:affiliation id="16AB2E6C77F2E8793CA6685466714051">Faculty of Biology, Philipps University Marburg, Marburg, Germany</mods:affiliation>
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<mods:namePart id="0D775511EDAC22994658C3F6332C3104">Opgenoorth, Lars</mods:namePart>
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<mods:affiliation id="EF76C596BA47D0A59AE9F5FB8CF5BE2B">Faculty of Biology, Philipps University Marburg, Marburg, Germany</mods:affiliation>
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<mods:namePart id="B7A63DA89564EECC47F4D738C62AB698">Woldehawariat, Yitbarek</mods:namePart>
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<mods:nameIdentifier id="1F1A0F10D9AC070C4B9E7BE649E2F5B3" type="ORCID">https://orcid.org/0000-0002-0918-8906</mods:nameIdentifier>
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<mods:affiliation id="5691C5C489E022655BE3CEBDAA0D7E61">Department of Zoological Sciences, Addis Ababa University, Addis Ababa, Ethiopia</mods:affiliation>
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<mods:namePart id="E191B159BF01BAA9C44FF5ADEFEB0AFD">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="3BF1D2C1DD030CAF347EDA69ED91C5C8">General and Systematic Zoology, University of Rostock, Rostock, Germany</mods:affiliation>
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<mods:nameIdentifier id="1A47746DD485A699B5FFDA77D2BEDA8E" type="email">agonumschmidt@hotmail.com</mods:nameIdentifier>
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<mods:title id="60342E00B28D1852B9358366056437B8">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="8FA26F579E418E93F20034F7081AD122">2023</mods:date>
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<mods:number id="7F64DFCD7552C6EBB3DAD6A9BB4541DA">2023-09-27</mods:number>
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<mods:number id="AFB4A313EE31B2F86BE3DA2AC8656CFE">70</mods:number>
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<mods:url id="3FFEB5513A3B77F7C8E32B556BCFA623">http://dx.doi.org/10.3897/dez.70.107425</mods:url>
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<mods:classification id="F4792BC79722285A36F092849F8A59B9">journal article</mods:classification>
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<mods:identifier id="E6E03229B281E738ABAB48F39B518496" type="DOI">http://dx.doi.org/10.3897/dez.70.107425</mods:identifier>
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<treatment id="78E8D3AE40CD5B96A44FFBA1906E2361" ID-GBIF-Taxon="213623267" LSID="urn:lsid:zoobank.org:act:71FD62CE-3922-4927-9A70-E94D7DD7A7A0" httpUri="http://treatment.plazi.org/id/78E8D3AE40CD5B96A44FFBA1906E2361" lastPageNumber="311" pageId="0" pageNumber="311" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Trechina" scope_tribe="Trechini">
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<subSubSection id="4638C8EA83A41769FD591FF6C2C105F1" pageId="0" pageNumber="311" type="nomenclature">
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<paragraph id="09D4A4AE2F8CE66CBC0DED89E7BAB046" pageId="0" pageNumber="311">
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<taxonomicName id="5CCF1EFE329806C763CE9453AB78A538" LSID="https://zoobank.org/71FD62CE-3922-4927-9A70-E94D7DD7A7A0" authority="Schmidt & Faille, gen. nov." authorityName="Schmidt & Faille, gen. nov." class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">Baehria Schmidt & Faille, gen. nov.</taxonomicName>
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</paragraph>
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<subSubSection id="4CE466C5C77A9BAF3ECF5F075E92988A" pageId="0" pageNumber="311" type="description">
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<paragraph id="90D12F67E34CACEF53D218239F11F3EC" pageId="0" pageNumber="311">
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<figureCitation id="556A1AC6419C553B56DD5A9069DDBB15" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">Figs 4-7</figureCitation>
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<figureCitation id="552A08273962CAF66B7785C8C2EF515A" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">, 8-15</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="63D9BE78899BD7C1D67851D376B14571" pageId="0" pageNumber="311" type="type species">
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<paragraph id="F9AAD633EE6B0A7948489F6D1E10E5FF" pageId="0" pageNumber="311">Type species.</paragraph>
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<paragraph id="8EF8C44D00FB6C8745F76C4BAB463830" pageId="0" pageNumber="311">
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<taxonomicName id="0A82CC9387B51DEE339F8FDD496D8063" class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria separata" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="separata">
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<emphasis id="3A014F8C39D4FC65FFF1F77D3D16AC1B" italics="true" pageId="0" pageNumber="311">Baehria separata</emphasis>
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</taxonomicName>
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sp. nov., herewith designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="F76CCC71A9A380D4E20899A7B4A94E00" pageId="0" pageNumber="311" type="diagnosis">
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<paragraph id="24615EDBE1D77896C138635902EDD65B" pageId="0" pageNumber="311">Diagnosis.</paragraph>
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<paragraph id="3D0DB007D9DA9F63664630F2F83D0C04" pageId="0" pageNumber="311">
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Representative of subtribe
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<taxonomicName id="EF145CC5FEC6C81619F4C67688C507CF" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subTribe" subTribe="Trechina">Trechina</taxonomicName>
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due to presence of bidentate mandibles (absence of retinacle) and dorsally closed aedeagal median lobe (
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<bibRefCitation id="1E3DF602B86BEB393E2A534AB6FD5387" author="Jeannel, R" journalOrPublisher="Journal d'Entomologie" pageId="0" pageNumber="311" pagination="221 - 550" refId="B20" refString="Jeannel, R, 1926. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Premiere livraison). L'Abeille. Journal d'Entomologie 32: 221 - 550" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Premiere livraison). L'Abeille." volume="32" year="1926">Jeannel 1926</bibRefCitation>
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). Comparatively large trechines characterized by robust head, large mandibles, small but markedly protruded eyes, smooth, markedly convex tempora, cordiform pronotum, straight pronotal basal margin with large, rectangular to acute laterobasal angles, slender elytra, short metepisternum, reduced hindwings, moderately slender antenna and legs, protibia with a complete longitudinal groove on external surface, presence of a row of long adhesive hairs on apical margins of 4th tarsomeres which are as long as the 5th tarsomeres, and by a markedly slender median lobe of aedeagus, which possesses a small, slightly sclerotized endophallic copulatory piece. The new genus is particularly distinguished from other representatives of
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<taxonomicName id="B44BE7E4C04E8FCDE475B20A9BF7988C" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subTribe" subTribe="Trechina">Trechina</taxonomicName>
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by unusual chaetotaxy of head capsule and elytra as follows: clypeus plurisetose, each side with 3-5 setae; submentum with three setae each side; anterior elytral discal seta located in the 4th interval, adjoined to the 4th stria.
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</paragraph>
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</subSubSection>
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<subSubSection id="928A404828FB809199A14DFC69223667" pageId="0" pageNumber="311" type="etymology">
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<paragraph id="908CE50161BA1227CD771A2803CC1DB2" pageId="0" pageNumber="311">Etymology.</paragraph>
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<paragraph id="A2138E0941608DF7AD7F6442A9618815" pageId="0" pageNumber="311">
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The new genus name is given
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<emphasis id="7600521729BAC5545AA9E76F955AC8A9" italics="true" pageId="0" pageNumber="311">in memoriam</emphasis>
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of our dear friend and colleague, the distinguished entomologist Martin Baehr, Munich (10.03.1943-17.04.2019).
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</paragraph>
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</subSubSection>
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<subSubSection id="0CD0265463436C98E47FB9C37FD8124D" pageId="0" pageNumber="311" type="description">
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<paragraph id="91793B5D740BCB8D1EE5B23F639F5C7F" pageId="0" pageNumber="311">Description.</paragraph>
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<paragraph id="A9C8EA74D5D0D0DDED0D366817D2C5CB" pageId="0" pageNumber="311">
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<emphasis id="3898404497B226C816AF931B75646552" bold="true" italics="true" pageId="0" pageNumber="311">Head</emphasis>
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: Large and robust, without pilosity. Mandibles large, moderately slender, with bidentate dentition pattern as shown in Fig.
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<figureCitation id="D3F3F868E4B8D18C34A53B12E38E2293" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">8</figureCitation>
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. Labrum with apical margin moderately emarginated, with six setae near apical margin. Clypeus each side with three or four setae (Figs
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<figureCitation id="5E58642B337DFE9FC62556C391E97154" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">5</figureCitation>
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,
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<figureCitation id="2A3F522F49517BEA4F3D69BFFF5D4B5E" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">7</figureCitation>
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; seldom only two setae at one of the sides). Eyes moderately small, as long as or slightly shorter than tempora, markedly convexly protruded (Figs
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<figureCitation id="EE6094770DAD9BCC01D6DCCADE86E4C6" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">5</figureCitation>
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,
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<figureCitation id="13B50781BC4AC546460237BEBF3DD7CF" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">7</figureCitation>
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). Two supraorbital setae each side in normal position for
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<taxonomicName id="658803BAAAC9C7C721151BB30E899D43" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subTribe" subTribe="Trechina">Trechina</taxonomicName>
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. Supraorbital furrows unevenly bent in posterior half, markedly deep in front and middle portions, slightly flatter near insertion of posterior supraorbital seta. Tempora markedly convex, markedly wrinkled to the neck, smooth. Mid of head convexly elevated, with a distinct transverse depression between supraorbital area and neck (Figs
|
||
<figureCitation id="9B474687692439281DA93734E1C3265A" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">5</figureCitation>
|
||
,
|
||
<figureCitation id="4E4F8559DF942AF53F06A65B6195A47A" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">7</figureCitation>
|
||
). Antennae slender, with third antennomere longest, 1/9-1/10 longer than first respectively fourth antennomere, and with second antennomere about 2/3 of length of third. Suborbital seta present. Apical tooth of mentum bifid, sensory pits of mentum present; submentum with three setae each side (Fig.
|
||
<figureCitation id="083E5EB738EA93A7B662E5546ED02E87" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8CC31FD4B5254128A8FCC471DE1EE878" pageId="0" pageNumber="311">
|
||
<emphasis id="33B72F8335970B6B19C5901BE94904FA" bold="true" italics="true" pageId="0" pageNumber="311">Prothorax</emphasis>
|
||
: Pronotum rather small, without pilosity, moderately transverse, cordate, broadest distinctly before middle, with lateral margin markedly concave before base, and with basal margin slightly smaller than apical margin (Figs
|
||
<figureCitation id="AFCB6C43CF49BDA0D6AD781FA1145CE4" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">4</figureCitation>
|
||
,
|
||
<figureCitation id="734547E8908D3F8809836BC1EB6AB29C" captionStart="Figures 4–7" captionStartId="F4" captionText="Figures 4 - 7. Baehria separata Schmidt & Faille, gen. nov., sp. nov., dorsal aspect of body (4, 6) and head (5, 7) of paratypes; 4, 5. Male; 6, 7. Female. The small white circles in Figs 5 and 7 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures4-7" httpUri="https://binary.pensoft.net/fig/913114" pageId="0" pageNumber="311">6</figureCitation>
|
||
). Disc moderately convex. Anterior margin straight or slightly concave in middle with anterior angles small but distinct, rounded, moderately protruded. Basal margin straight along internal 3/4, with laterobasal angles slightly shifted posteriad. Lateral margin convexly rounded in anterior 2/3 and concave towards laterobasal angles, latter large, rectangular or sharp at tip, sometimes slightly protruded laterally. Marginal gutter moderately broad throughout. Median longitudinal impression sharply incised, disappearing near apex, somewhat deepened before base. Anterior and posterior transverse impressions shallow and smooth. Laterobasal foveae large, internally and externally (towards lateral gutter) distinctly sloped, without punctures but with fine transversal wrinkles. Lateral and laterobasal setae present, with the former situated at or slightly anterad of maximum width of pronotum. Proepisternum glabrous and smooth.
|
||
</paragraph>
|
||
<paragraph id="B943F2D120D9974F77E2D8F6A7B9BF49" pageId="0" pageNumber="311">
|
||
<emphasis id="E4B9D53033A34BEB68F0B3FA59AA717D" bold="true" italics="true" pageId="0" pageNumber="311">Pterothorax</emphasis>
|
||
: Elytra without pilosity, long and very slender ovate, very slightly convex or flattened in middle of disc, in dorsal view broadest distinctly posterad middle, shoulders flatly rounded (Fig.
|
||
<figureCitation id="BA303AEF78342A45789237514920E6B1" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">13</figureCitation>
|
||
), apical sinuation distinct, apex rounded with the indication of a very obtuse apical angle. Striae 1-8 complete, moderately deep impressed, impunctate, intervals moderately convex, parascutellar stria free, 1/6-1/9 of length of elytra. Recurrent preapical stria deep, long, in most specimens reaching the apex of the fifth stria. Parascutellar seta present. Anterior discal seta located in the 4th interval, adjoined to the 4th stria, located near the end of the anterior elytral 5th (Fig.
|
||
<figureCitation id="DE3401081355D98038150A69C2BDED47" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">13</figureCitation>
|
||
); second discal seta located at the 3rd stria about at elytral middle (in most specimens, the posterior setiferous pore together with the 3rd stria is switched into the 4th interval); posterior discal seta (= subapical seta near the end of 3rd stria) present, located about 1/9 of elytral length from elytral apex; subapical seta of the recurrent stria isolated, distinctly removed from this stria by distance of 2-3 diameters of the setiferous pore. Number and positions of the setae of the marginal umbilicate series as in
|
||
<taxonomicName id="BF76BD53BE74C4BACDD46FBBEBD977A3" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BC314477B36689B10CAB6686FE245739" italics="true" pageId="0" pageNumber="311">Trechus</emphasis>
|
||
</taxonomicName>
|
||
s. str. Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin.
|
||
</paragraph>
|
||
<paragraph id="435962EE6BE99E6AE9385D6DB28D4341" pageId="0" pageNumber="311">
|
||
<emphasis id="92E7E694420ECD8EC80DCFD8CB39FC90" bold="true" italics="true" pageId="0" pageNumber="311">Legs</emphasis>
|
||
: Moderately long and robust. Protibia distinctly dilated towards apex, straight, with longitudinal groove on dorsal surface complete, and with several fine setae on anterior surface near apex (Fig.
|
||
<figureCitation id="CC15C8C4B41ACEA36351F731AF5C212C" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">12</figureCitation>
|
||
). Two basal protarsomeres of males dilated and dentoid at the inner apical border (Fig.
|
||
<figureCitation id="62CD2D7C30C12B0B3659BD6EE65DB5C1" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">10</figureCitation>
|
||
). Fourth pro-, meso- and metatarsomeres each with a row of long adhesive hairs on apical margins which are as long as the 5th tarsomeres (Figs
|
||
<figureCitation id="0E30D8AAB962C33400DC5EAFF219B6CA" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">10</figureCitation>
|
||
,
|
||
<figureCitation id="53573122B9CD92EA9DA138AE55EBD3BA" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D97BDAAAF3ABF62D4BF8325377F07C6D" pageId="0" pageNumber="311">
|
||
<emphasis id="A7C31EDFB52EB6C8711DF372ADE07E84" bold="true" italics="true" pageId="0" pageNumber="311">Male genitalia</emphasis>
|
||
(Figs
|
||
<figureCitation id="958C560699E36F57AA624B856E04077F" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">14</figureCitation>
|
||
,
|
||
<figureCitation id="7A35BC3844544A0B186C9A206E7FEDBA" captionStart="Figures 8–15" captionStartId="F5" captionText="Figures 8 - 15. Baehria separata Schmidt & Faille, gen. nov., sp. nov. 8. Left and right mandible, dorsal aspect; 9. Ventral aspect of head; the small white circles mark the insertion points of the setae on submentum; 10. Right male protarsomeres, left latero-ventral aspect; 11. Left male metatarsomeres, right lateral aspect; 12. Right male protibia, dorsal aspect; 13. Anterior part of elytra and pronotal base; the arrows point to the insertions of the anterior elytral discal setae; 14. Aedeagus, left lateral aspect; 15. Aedeagus, dorsal aspect." figureDoi="10.3897/dez.70.107425.figures8-15" httpUri="https://binary.pensoft.net/fig/913115" pageId="0" pageNumber="311">15</figureCitation>
|
||
): Aedeagal median lobe markedly elongated, slender tube-like, in lateral view slightly sinusoidal, with apex distinctly bent upwardly, simple, with apical lamella insignificant; basal bulb rather small with large sagittal aileron. Endophallus with a small, very slightly sclerotized copulatory piece. Parameres with 3-4 apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="334FE47B3481363A99300A3267BE826C" pageId="0" pageNumber="311" type="distribution">
|
||
<paragraph id="692E6AE4A0B8D83E2F37D3A844E0EC1F" pageId="0" pageNumber="311">Distribution.</paragraph>
|
||
<paragraph id="0B632320E46C9BC561895F7D204C48E7" pageId="0" pageNumber="311">
|
||
So far only known from Mt. Choke in northern Ethiopia (Fig.
|
||
<figureCitation id="FEE65F059FDAE143F3DCBACF895D613E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of Ethiopia showing the main topographic features of the country. Seven prominent volcanic massifs which are discussed in the text are highlighted. The base map was downloaded from www. freeworldmaps. net (01 - 07 - 2023)." figureDoi="10.3897/dez.70.107425.figure1" httpUri="https://binary.pensoft.net/fig/913111" pageId="0" pageNumber="311">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CE3089EBE1E8F70760AD6B487510986D" pageId="0" pageNumber="311" type="relationships and identification">
|
||
<paragraph id="BE0EA4A8C5275ED69AABAF2FB63A967F" pageId="0" pageNumber="311">Relationships and identification.</paragraph>
|
||
<paragraph id="792DEC986A0269191E1E95592EF2F3DD" pageId="0" pageNumber="311">
|
||
Based on the molecular data,
|
||
<taxonomicName id="879D9106ED448CACAD86354671D73185" class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FBBE3E0CA1847EEC063BC3B3AA2E4317" italics="true" pageId="0" pageNumber="311">Baehria</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. is representative of a well-supported clade comprising
|
||
<taxonomicName id="CD00C86946BEEFBE77709C1D8E26E409" authorityName="Jeannel" authorityYear="1928" class="Insecta" family="Carabidae" genus="Duvaliomimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Duvaliomimus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01F7D0329E436ADFA69A4B4535295EAC" italics="true" pageId="0" pageNumber="311">Duvaliomimus</emphasis>
|
||
</taxonomicName>
|
||
Jeannel from New Zealand and
|
||
<taxonomicName id="79CE3AD13C284D9819CA3881F7288371" authorityName="Jeannel" authorityYear="1920" class="Insecta" family="Carabidae" genus="Paratrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratrechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3ED17370A8797EF64760AC75A2018BCF" italics="true" pageId="0" pageNumber="311">Paratrechus</emphasis>
|
||
</taxonomicName>
|
||
Jeannel from South and Central America (Fig.
|
||
<figureCitation id="66F349C69D00A5AFE5CF9B03063849B9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ultrametric time-calibrated phylogeny of Trechini beetles and outgroups (basal part of the tree, continues in Fig. 3). The tree was obtained from BEAST 2 based on the concatenated sequence data and using two age constraints (for details see text). Black circles and stars at branch nodes refer to posterior probabilities ≥ 0.98 and bootstrap values> 75, respectively. Grey bars specify the 95 % HPD of the respective node age. Collapsed, colored trechine clades refer to previous results of Faille et al. (2013). The southern hemisphere Duvaliomimus - Paratrechus clade with the Ethiopian endemic Baehria gen. nov. is highlighted ochre." figureDoi="10.3897/dez.70.107425.figure2" httpUri="https://binary.pensoft.net/fig/913112" pageId="0" pageNumber="311">2</figureCitation>
|
||
). Apart from a general "
|
||
<taxonomicName id="A51456F61F603E0D98BF01FD371E4A8E" authorityName="Delarouzee" authorityYear="1859" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Duvalius" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F41C53ABD2494035DDE9D84FCCF1949F" italics="true" pageId="0" pageNumber="311">Duvalius</emphasis>
|
||
</taxonomicName>
|
||
-like" appearance, the three genera share some common features like the large size, cordate pronotum, rounded head with salient temples, and pubescent protibiae.
|
||
<taxonomicName id="C16FB87309ED453826312414BA6EF895" class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="16AD644E22546EB4031A220BBEB1E276" italics="true" pageId="0" pageNumber="311">Baehria</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. differs from both these genera by the presence of three or four setae instead of two on each side of the clypeus, and by the position of the anterior discal seta on elytra: in
|
||
<taxonomicName id="F125814A8437A883774C23D0FC51286D" class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DA78CEB2CA68CDAD0084461FCF052D51" italics="true" pageId="0" pageNumber="311">Baehria</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., this seta inserts in the 4th interval instead on the 3rd stria in
|
||
<taxonomicName id="308B00E47B8ECDC6418018923A94CB79" authorityName="Jeannel" authorityYear="1928" class="Insecta" family="Carabidae" genus="Duvaliomimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Duvaliomimus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFA6F7FC5326DFA56FFE2CB6D0712E57" italics="true" pageId="0" pageNumber="311">Duvaliomimus</emphasis>
|
||
</taxonomicName>
|
||
and on the 5th stria in
|
||
<taxonomicName id="38D2268F88C7FBA564EB53FE66E02F1F" authorityName="Jeannel" authorityYear="1920" class="Insecta" family="Carabidae" genus="Paratrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratrechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F57CE89FAA829675FA28CD6FDDE815D3" italics="true" pageId="0" pageNumber="311">Paratrechus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="E65FFA9249398863843BBACA202CFA99" class="Insecta" family="Carabidae" genus="Baehria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baehria" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7DBF646AD3376973749F85C40AE6FF8" italics="true" pageId="0" pageNumber="311">Baehria</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. differs additionally from
|
||
<taxonomicName id="B725574D92F35E2B1246E1A008A85735" authorityName="Jeannel" authorityYear="1920" class="Insecta" family="Carabidae" genus="Paratrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratrechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B848E1C64950F5B68BCE89FD9A0DF7B0" italics="true" pageId="0" pageNumber="311">Paratrechus</emphasis>
|
||
</taxonomicName>
|
||
by the simple apex of the aedeagal median lobe, which is button-like shaped in
|
||
<taxonomicName id="45F32FA4B5DBE9135B26FAC365A695B5" authorityName="Jeannel" authorityYear="1920" class="Insecta" family="Carabidae" genus="Paratrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratrechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FA979B40E0CBDED9272CE3AC49D5366" italics="true" pageId="0" pageNumber="311">Paratrechus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="05241082FD9E8795A95DB880B61B03EA" author="Jeannel, R" journalOrPublisher="Journal d'Entomologie" pageId="0" pageNumber="311" pagination="1 - 808" refId="B22" refString="Jeannel, R, 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Troisieme livraison). L'Abeille. Journal d'Entomologie 35: 1 - 808" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Troisieme livraison). L'Abeille." volume="35" year="1928">Jeannel 1928</bibRefCitation>
|
||
; Barr 1982;
|
||
<bibRefCitation id="9ED77669E482536839E88B78343C37A1" author="Townsend, JI" journalOrPublisher="Manaaki Whenua Press, Lincoln" pageId="0" pageNumber="311" refId="B52" refString="Townsend, JI, 2010. Trechini (Insecta: Coleoptera: Carabidae: Trechinae). Fauna of New Zealand 62. Manaaki Whenua Press, Lincoln" title="Trechini (Insecta: Coleoptera: Carabidae: Trechinae). Fauna of New Zealand 62." year="2010">Townsend 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |