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<document ID-DOI="10.11646/zootaxa.5258.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7784364" ID-ZooBank="633CA10E-D3FA-4548-9316-79E97BB0FA6B" approvalRequired="5" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="1" checkinTime="1680164730542" checkinUser="plazi" docAuthor="Zhu, Xiaoju, Zhang, Li &amp; Wang, Shuxia" docDate="2023" docId="781087D4FFEC5873A483D3538BC2AF47" docLanguage="en" docName="zootaxa.5258.4.2.pdf" docOrigin="Zootaxa 5258 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agonopterix introrsa Zhu &amp; Zhang &amp; Wang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="386" masterDocId="8429FFACFFEA5874A414D04C8B2FAE5A" masterDocTitle="New species and newly recorded species of the genus Agonopterix Hübner, [1825] (Lepidoptera: Depressariidae) from China" masterLastPageNumber="404" masterPageNumber="379" pageNumber="385" updateTime="1680165344539" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and newly recorded species of the genus Agonopterix Hübner, [1825] (Lepidoptera: Depressariidae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhu, Xiaoju</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-4590-8480</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Nankai University, Tianjin 300071, China. &amp; https: // orcid. org / 0000 - 0002 - 4590 - 8480</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Li</mods:namePart>
<mods:affiliation>College of Life Sciences, Nankai University, Tianjin 300071, China.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Shuxia</mods:namePart>
<mods:affiliation>College of Life Sciences, Nankai University, Tianjin 300071, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5258</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:start>379</mods:start>
<mods:end>404</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5258.4.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7784364</mods:identifier>
<mods:identifier type="ZooBank">633CA10E-D3FA-4548-9316-79E97BB0FA6B</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7784396" ID-Zenodo-Dep="7784396" LSID="urn:lsid:plazi:treatment:781087D4FFEC5873A483D3538BC2AF47" httpUri="http://treatment.plazi.org/id/781087D4FFEC5873A483D3538BC2AF47" lastPageId="7" lastPageNumber="386" pageId="6" pageNumber="385">
<subSubSection box="[151,684,798,825]" pageId="6" pageNumber="385" type="nomenclature">
<paragraph blockId="6.[151,684,798,861]" box="[151,684,798,825]" pageId="6" pageNumber="385">
<heading bold="true" box="[151,684,798,825]" fontSize="11" level="1" pageId="6" pageNumber="385" reason="1">
<emphasis bold="true" box="[151,684,798,825]" pageId="6" pageNumber="385">
<taxonomicName authority="Zhu &amp; Wang" authorityName="Zhu &amp; Zhang &amp; Wang" authorityYear="2023" box="[151,578,798,825]" class="Insecta" family="Oecophoridae" genus="Agonopterix" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="introrsa" status="sp. nov.">
<emphasis bold="true" box="[151,404,799,825]" italics="true" pageId="6" pageNumber="385">Agonopterix introrsa</emphasis>
Zhu &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel box="[591,684,799,825]" pageId="6" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,281,835,861]" pageId="6" pageNumber="385" type="description">
<paragraph blockId="6.[151,684,798,861]" box="[151,281,835,861]" pageId="6" pageNumber="385">
(
<figureCitation box="[159,232,835,861]" captionStart="FIGURES 18" captionStartId="8.[151,264,1609,1633]" captionTargetBox="[171,1407,181,1566]" captionTargetId="figure-22@8.[151,1435,181,1585]" captionTargetPageId="8" captionText="FIGURES 18. Adults of Agonopterix spp. 1, A. basinigra sp. nov., ♁; 2, A. cochleata sp. nov., ♁; 3, A. epunctata sp. nov., ♁; 4, A. foliiformis sp. nov., ♁; 5, A. gibbosa sp. nov., ♁; 6, A. hypodroma sp. nov., ♁; 7, A. introrsa sp. nov., ♁; 8, A. magnimacularis sp. nov., ♁. All holotypes. Scales = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7784366" httpUri="https://zenodo.org/record/7784366/files/figure.png" pageId="6" pageNumber="385">Figs 7</figureCitation>
,
<figureCitation box="[244,273,835,861]" captionStart="FIGURES 3035" captionStartId="18.[151,264,1903,1927]" captionTargetBox="[180,1407,181,1866]" captionTargetId="figure-22@18.[180,1407,181,1880]" captionTargetPageId="18" captionText="FIGURES 3035. Male genitalia of Agonopterix spp. 30, A. introrsa sp. nov., holotype, slide No. ZXJ20453; 31, A. magnimacularis sp. nov., holotype, slide No. ZXJ20395; 32, A. nephimacula sp. nov., holotype, slide No. ZXJ20473; 33, A. nivimacularis sp. nov., paratype, slide No. ZXJ20404, 33a, enlarged uncus, slide No. ZXJ20436; 34, A. parallela sp. nov., holotype, slide No. ZXJ20557; 35, A. simipullella sp. nov., holotype, ZXJ20486. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7784380" httpUri="https://zenodo.org/record/7784380/files/figure.png" pageId="6" pageNumber="385">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="385" type="materials_examined">
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">
<emphasis bold="true" box="[151,332,906,933]" pageId="6" pageNumber="385">Type material.</emphasis>
<materialsCitation collectingDate="2014-06-17" collectorName="SR Liu &amp; No." country="China" county="Xinglong County" elevation="1000" latitude="40.63" location="Mt. Wuling" longLatPrecision="697" longitude="117.43" pageId="6" pageNumber="385" specimenCount="1" stateProvince="Hebei" typeStatus="holotype">
<emphasis bold="true" box="[346,539,906,932]" pageId="6" pageNumber="385">
<collectingCountry box="[346,441,906,932]" name="China" pageId="6" pageNumber="385">CHINA</collectingCountry>
,
<collectingRegion box="[460,534,906,932]" country="China" name="Hebei" pageId="6" pageNumber="385">Hebei</collectingRegion>
:
</emphasis>
<typeStatus box="[553,657,907,933]" pageId="6" pageNumber="385">Holotype</typeStatus>
♁,
<location LSID="urn:lsid:plazi:treatment:781087D4FFEC5873A483D3538BC2AF47:F5666019FFEC5872A6DDD3C7887EADFF" box="[713,849,907,933]" country="China" county="Xinglong County" latitude="40.63" longLatPrecision="697" longitude="117.43" name="Mt. Wuling" pageId="6" pageNumber="385" stateProvince="Hebei">Mt. Wuling</location>
(
<geoCoordinate box="[870,967,907,933]" degrees="40.63" direction="north" orientation="latitude" pageId="6" pageNumber="385" precision="555" value="40.63">40.63ºN</geoCoordinate>
,
<geoCoordinate box="[985,1091,907,933]" degrees="117.43" direction="east" orientation="longitude" pageId="6" pageNumber="385" precision="555" value="117.43">117.43ºE</geoCoordinate>
),
<collectingCounty box="[1116,1320,907,933]" pageId="6" pageNumber="385">Xinglong County</collectingCounty>
,
<quantity box="[1338,1431,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="385" unit="m" value="1000.0">
<elevation box="[1338,1431,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="385" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date box="[151,282,943,969]" pageId="6" pageNumber="385" value="2014-06-17">
<collectingDate box="[151,282,943,969]" pageId="6" pageNumber="385" value="2014-06-17">17.VI.2014</collectingDate>
</date>
. leg.
<collectorName box="[341,424,943,969]" pageId="6" pageNumber="385">SR Liu</collectorName>
, slide
<collectorName box="[495,537,943,969]" pageId="6" pageNumber="385">No.</collectorName>
ZXJ20453
</materialsCitation>
.
<materialsCitation collectingDate="2000-05-29" collectorName="HL Yu &amp; No. ZL" country="China" elevation="1500" location="Lushi" pageId="6" pageNumber="385" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
<typeStatus box="[676,789,943,969]" pageId="6" pageNumber="385">Paratypes</typeStatus>
:
<emphasis bold="true" box="[799,878,943,969]" pageId="6" pageNumber="385">
<collectingRegion box="[799,878,943,969]" country="China" name="Henan" pageId="6" pageNumber="385">Henan</collectingRegion>
</emphasis>
: 1♁,
<location LSID="urn:lsid:plazi:treatment:781087D4FFEC5873A483D3538BC2AF47:F5666019FFEC5872A7BAD3E388DEAD93" box="[942,1009,943,969]" country="China" name="Lushi" pageId="6" pageNumber="385" stateProvince="Henan">Lushi</location>
,
<quantity box="[1020,1107,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="385" unit="m" value="1500.0">
<elevation box="[1020,1107,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="385" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date box="[1119,1236,943,969]" pageId="6" pageNumber="385" value="2000-05-29">
<collectingDate box="[1119,1236,943,969]" pageId="6" pageNumber="385" value="2000-05-29">29.V.2000</collectingDate>
</date>
, leg.
<collectorName box="[1296,1372,943,969]" pageId="6" pageNumber="385">HL Yu</collectorName>
, slide
<collectorName box="[151,235,979,1005]" pageId="6" pageNumber="385">No. ZL</collectorName>
09004
</materialsCitation>
;
<materialsCitation box="[318,1315,978,1007]" collectingDate="1987-06-15" collectorName="HH Li &amp; No. W" country="China" elevation="1620" location="Ningshan" municipality="Huoditang" pageId="6" pageNumber="385" specimenCount="1" stateProvince="Shaanxi" typeStatus="paratype">
<emphasis bold="true" box="[318,414,978,1004]" pageId="6" pageNumber="385">
<collectingRegion box="[318,414,978,1004]" country="China" name="Shaanxi" pageId="6" pageNumber="385">Shaanxi</collectingRegion>
</emphasis>
: 1♁,
<collectingMunicipality box="[479,600,979,1005]" pageId="6" pageNumber="385">Huoditang</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:781087D4FFEC5873A483D3538BC2AF47:F5666019FFEC5872A677D39F89FEADB7" box="[611,721,979,1005]" country="China" municipality="Huoditang" name="Ningshan" pageId="6" pageNumber="385" stateProvince="Shaanxi">Ningshan</location>
,
<quantity box="[732,819,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="6" pageNumber="385" unit="m" value="1620.0">
<elevation box="[732,819,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="6" pageNumber="385" unit="m" value="1620.0">1620 m</elevation>
</quantity>
,
<date box="[831,961,978,1005]" pageId="6" pageNumber="385" value="1987-06-15">
<collectingDate box="[831,961,978,1005]" pageId="6" pageNumber="385" value="1987-06-15">15.VI.1987</collectingDate>
</date>
, leg.
<collectorName box="[1021,1095,979,1005]" pageId="6" pageNumber="385">HH Li</collectorName>
, slide
<collectorName box="[1167,1243,979,1005]" pageId="6" pageNumber="385">No. W</collectorName>
98014
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="385" type="diagnosis">
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">
<emphasis bold="true" box="[199,322,1014,1040]" pageId="6" pageNumber="385">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName authority="Wang, 2007" authorityName="Wang" authorityYear="2007" box="[677,995,1015,1041]" class="Insecta" family="Oecophoridae" genus="Agonopterix" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="acutivalvula">
<emphasis box="[677,848,1015,1041]" italics="true" pageId="6" pageNumber="385">A. acutivalvula</emphasis>
Wang, 2007
</taxonomicName>
(
<bibRefCitation author="Fan, X. M. &amp; Wang, S. X." box="[1012,1228,1015,1041]" pageId="6" pageNumber="385" pagination="215 - 222" refId="ref15068" refString="Fan, X. M. &amp; Wang, S. X. (2007) Genus Agonopterix Hubner (Lepidoptera: Elachistidae: Depressariinae) from China with descriptions of four new species. Entomotaxonomia, 29 (3), 215 - 222." type="journal article" year="2007">Fan &amp; Wang 2007</bibRefCitation>
) in both forewing pattern and male genitalia. It can be distinguished in the male genitalia by the reduced uncus, the juxta shallowly concave medially on the posterior margin and the cuiller reaching anterior 1/5 from the costa. In
<taxonomicName authorityName="Wang" authorityYear="2007" box="[1220,1389,1087,1113]" class="Insecta" family="Oecophoridae" genus="Agonopterix" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="acutivalvula">
<emphasis box="[1220,1389,1087,1113]" italics="true" pageId="6" pageNumber="385">A. acutivalvula</emphasis>
</taxonomicName>
, the uncus is papilliform, the juxta is deeply concave in broad V shape on the posterior margin, and the cuiller almost exceeds the costa (
<bibRefCitation author="Fan, X. M. &amp; Wang, S. X." box="[358,620,1159,1185]" pageId="6" pageNumber="385" pagination="215 - 222" refId="ref15068" refString="Fan, X. M. &amp; Wang, S. X. (2007) Genus Agonopterix Hubner (Lepidoptera: Elachistidae: Depressariinae) from China with descriptions of four new species. Entomotaxonomia, 29 (3), 215 - 222." type="journal article" year="2007">Fan &amp; Wang 2007: 217</bibRefCitation>
, fig. 8). It is similar to
<taxonomicName authority="Zhu &amp; Zhang &amp; Wang, 2023" authorityName="Zhu &amp; Zhang &amp; Wang" authorityYear="2023" box="[871,1018,1159,1185]" class="Insecta" family="Oecophoridae" genus="Agonopterix" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="foliiformis" status="sp. nov.">
<emphasis box="[871,1018,1159,1185]" italics="true" pageId="6" pageNumber="385">A. foliiformis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1024,1113,1159,1185]" pageId="6" pageNumber="385">
<taxonomicNameLabel box="[1024,1113,1159,1185]" pageId="6" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and the differences between them are stated in the diagnosis of the latter species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="386" pageId="6" pageNumber="385" type="description">
<paragraph blockId="6.[151,1437,906,2013]" box="[199,808,1230,1257]" pageId="6" pageNumber="385">
<emphasis bold="true" box="[199,345,1230,1256]" pageId="6" pageNumber="385">Description.</emphasis>
Adult (
<figureCitation box="[429,498,1231,1257]" captionStart="FIGURES 18" captionStartId="8.[151,264,1609,1633]" captionTargetBox="[171,1407,181,1566]" captionTargetId="figure-22@8.[151,1435,181,1585]" captionTargetPageId="8" captionText="FIGURES 18. Adults of Agonopterix spp. 1, A. basinigra sp. nov., ♁; 2, A. cochleata sp. nov., ♁; 3, A. epunctata sp. nov., ♁; 4, A. foliiformis sp. nov., ♁; 5, A. gibbosa sp. nov., ♁; 6, A. hypodroma sp. nov., ♁; 7, A. introrsa sp. nov., ♁; 8, A. magnimacularis sp. nov., ♁. All holotypes. Scales = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7784366" httpUri="https://zenodo.org/record/7784366/files/figure.png" pageId="6" pageNumber="385">Fig. 7</figureCitation>
). Wingspan
<quantity box="[638,804,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.15" metricValueMin="1.95" pageId="6" pageNumber="385" unit="mm" value="20.5" valueMax="21.5" valueMin="19.5">19.521.5 mm</quantity>
.
</paragraph>
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">
<emphasis box="[199,260,1267,1293]" italics="true" pageId="6" pageNumber="385">Head</emphasis>
white, with sparse brown scales, neck with erect white scales basally, with white-tipped brown distally. Labial palpus yellowish white; basal segment black on outside; second segment mixed with black and greyish brown scales on outside and on dorsal surface, with protruding scales in distal half; third segment black near base, with a wide black ring beyond middle, black at apex. Antenna with scape greyish brown mixed with black, white on anterior margin and at terminal; pecten white; flagellum dark brown alternated with yellowish brown.
</paragraph>
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">
<emphasis box="[199,330,1447,1472]" italics="true" pageId="6" pageNumber="385">Mesonotum</emphasis>
and tegula greyish brown. Foreleg with coxa yellowish white, femur, tibia and tarsus greyish brown mixed with black scales, tarsus yellowish white at apex of each tarsomere; midleg with femur grey mixed with black, tibia grey mixed with sparse black scales, ochreous at apex, tarsus grey mixed with black, yellowish white at apex of each tarsomere; hindleg with femur grey mixed with black, tibia yellowish white, with sparse black scales dorsally, tarsus yellowish white, with sparse black scales on outside.
</paragraph>
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">Forewing ground colour dusty grey, with sparse black scales; costal margin with irregularly arranged black spots; cell with two black spots obliquely placed near middle, with a white spot on outer margin surrounded by black scales; diffused black speckle above preceding white spot; fringe concolourous with forewing. Hindwing and fringe grey.</paragraph>
<paragraph blockId="6.[151,1437,906,2013]" pageId="6" pageNumber="385">
Male genitalia (
<figureCitation box="[380,466,1735,1761]" captionStart="FIGURES 3035" captionStartId="18.[151,264,1903,1927]" captionTargetBox="[180,1407,181,1866]" captionTargetId="figure-22@18.[180,1407,181,1880]" captionTargetPageId="18" captionText="FIGURES 3035. Male genitalia of Agonopterix spp. 30, A. introrsa sp. nov., holotype, slide No. ZXJ20453; 31, A. magnimacularis sp. nov., holotype, slide No. ZXJ20395; 32, A. nephimacula sp. nov., holotype, slide No. ZXJ20473; 33, A. nivimacularis sp. nov., paratype, slide No. ZXJ20404, 33a, enlarged uncus, slide No. ZXJ20436; 34, A. parallela sp. nov., holotype, slide No. ZXJ20557; 35, A. simipullella sp. nov., holotype, ZXJ20486. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7784380" httpUri="https://zenodo.org/record/7784380/files/figure.png" pageId="6" pageNumber="385">Fig. 30</figureCitation>
). Uncus reduced. Socii widened from narrow base to obtusely rounded apex, setose, widely apart at base, distance between base almost as wide as posterior margin of tegumen. Gnathos arrow-like, with fine spines. Valva wide at base, gradually narrowed to basal 3/4, sharply narrowed from distal 1/4 to narrowly rounded apex. Transtilla a narrow band; transtilla lobes dilated, setose. Sacculus reaching beyond 1/3 length of valva; cuiller narrowly clavate, extending obliquely inward, reaching anterior 1/5 from costa. Juxta shallowly concave medially on posterior margin, obtusely produced laterally, shrunken laterally near anterior margin, obtusely rounded on anterior margin; lateral lobes stout, semiovate, reaching posterior margin of juxta. Phallus wide in basal 1/4, uniformly slender medially, distal 1/4 tapered to apex; cornuti being numerous tiny spinules.
</paragraph>
<paragraph blockId="7.[151,1436,151,285]" box="[199,401,151,177]" pageId="7" pageNumber="386">Female unknown.</paragraph>
</subSubSection>
<subSubSection box="[199,718,186,213]" pageId="7" pageNumber="386" type="distribution">
<paragraph blockId="7.[151,1436,151,285]" box="[199,718,186,213]" pageId="7" pageNumber="386">
<emphasis bold="true" box="[199,353,186,212]" pageId="7" pageNumber="386">Distribution.</emphasis>
<collectingCountry box="[359,426,187,213]" name="China" pageId="7" pageNumber="386">China</collectingCountry>
(
<collectingRegion box="[441,512,187,213]" country="China" name="Hebei" pageId="7" pageNumber="386">Hebei</collectingRegion>
,
<collectingRegion box="[524,600,187,213]" country="China" name="Henan" pageId="7" pageNumber="386">Henan</collectingRegion>
,
<collectingRegion box="[611,706,187,213]" country="China" name="Shaanxi" pageId="7" pageNumber="386">Shaanxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="386" type="etymology">
<paragraph blockId="7.[151,1436,151,285]" pageId="7" pageNumber="386">
<emphasis bold="true" box="[199,334,223,249]" pageId="7" pageNumber="386">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis box="[849,949,223,248]" italics="true" pageId="7" pageNumber="386">introrsus</emphasis>
, referring to the cuiller extending obliquely inward.
</paragraph>
</subSubSection>
</treatment>
</document>