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<document id="769F5D65EA38ED163A016C1CB86AAA78" ID-DOI="10.5281/zenodo.186352" ID-GBIF-Dataset="c87b0ab2-a35e-4069-8c94-cb29159d4ec0" ID-ISSN="1175-5326" ID-Zenodo-Dep="186352" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050678747" checkinUser="plazi" docAuthor="Raven, Robert J." docDate="2009" docId="774E87C4D66EFF82FF3BFD2BFDB9FDDA" docLanguage="en" docName="zt02035p040.pdf" docOrigin="Zootaxa 2035" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mitzoruga elapines Raven, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="8B77FFBCD678FF9BFFACFFF7FFD4FF81" masterDocTitle="Revisions of Australian ground-hunting spiders: IV. The spider subfamily Diaprograptinae subfam. nov. (Araneomorphae: Miturgidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="23" updateTime="1698578758242" updateUser="plazi">
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<mods:title id="578F3C7AB1232E5BF499D5E88442DC40">Revisions of Australian ground-hunting spiders: IV. The spider subfamily Diaprograptinae subfam. nov. (Araneomorphae: Miturgidae)</mods:title>
</mods:titleInfo>
<mods:name id="AE71F85568A946F4CB409CF01B5A0981" type="personal">
<mods:role id="06F8B2BE60A0E94CCB08BE42F7B9D63F">
<mods:roleTerm id="2A94BD9568F054EE54802EE7310A81BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="596FC56C1CE6A5158A1AB522109387FE">Raven, Robert J.</mods:namePart>
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<mods:title id="930060D28715901B6E93A4CC458606D3">Zootaxa</mods:title>
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<mods:part id="13E984307E9420BB428C88EE06D6F9D8">
<mods:date id="59844C40F0DB37D17BC272912F1D33D5">2009</mods:date>
<mods:detail id="165219E0463C456100BC1403C1B52E31" type="volume">
<mods:number id="7A07A89AAC51DEA41C4AAF2CA95313D3">2035</mods:number>
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<mods:classification id="5F10796038273127B2E97A8A04E9CAEA">journal article</mods:classification>
<mods:identifier id="85D61B026B2D158CAEF5A86EC49288BF" type="DOI">10.5281/zenodo.186352</mods:identifier>
<mods:identifier id="0127D3AA34A07172ADE68A9F83D2D9A5" type="GBIF-Dataset">c87b0ab2-a35e-4069-8c94-cb29159d4ec0</mods:identifier>
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<treatment id="774E87C4D66EFF82FF3BFD2BFDB9FDDA" ID-DOI="http://doi.org/10.5281/zenodo.6216638" ID-GBIF-Taxon="119612058" ID-Zenodo-Dep="6216638" LSID="urn:lsid:plazi:treatment:774E87C4D66EFF82FF3BFD2BFDB9FDDA" httpUri="http://treatment.plazi.org/id/774E87C4D66EFF82FF3BFD2BFDB9FDDA" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection id="B7FD6559D66EFF8DFF3BFD2BFDDEFC99" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="FF5836D2D66EFF8DFF3BFD2BFE30FD74" blockId="22.[151,522,731,792]" box="[151,484,731,758]" pageId="22" pageNumber="23">
<heading id="A41081BED66EFF8DFF3BFD2BFE30FD74" bold="true" box="[151,484,731,758]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="CD93EAC0D66EFF8DFF3BFD2BFE30FD74" bold="true" box="[151,484,731,758]" pageId="22" pageNumber="23">
<taxonomicName id="38E74D51D66EFF8DFF3BFD2BFEABFD77" ID-CoL="6RKZW" box="[151,383,732,758]" class="Arachnida" family="Miturgidae" genus="Mitzoruga" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="elapines" status="sp. nov.">
<emphasis id="CD93EAC0D66EFF8DFF3BFD2BFEABFD77" bold="true" box="[151,383,732,758]" italics="true" pageId="22" pageNumber="23">Mitzoruga elapines</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A057BBD66EFF8DFE2AFD2CFE30FD74" box="[390,484,731,757]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF5836D2D66EFF8DFF3BFD09FDDEFC99" blockId="22.[151,522,731,792]" box="[151,522,766,792]" pageId="22" pageNumber="23">
(
<figureCitation id="67DC2A57D66EFF8DFF33FD09FF23FC99" box="[159,247,766,792]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="22" pageNumber="23">Figs 14</figureCitation>
aj, 15ad, 16a, b, 25a)
</paragraph>
</subSubSection>
<subSubSection id="B7FD6559D66EFF8DFF3BFCBCFC27FBF5" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="FF5836D2D66EFF8DFF3BFCBCFCDFFC0D" blockId="22.[151,1438,843,1991]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF3BFCBCFDFAFCE4" bold="true" box="[151,558,843,870]" pageId="22" pageNumber="23">
Material examined.
<emphasis id="CD93EAC0D66EFF8DFE01FCBBFDF1FCE7" bold="true" box="[429,549,844,870]" italics="true" pageId="22" pageNumber="23">
<typeStatus id="205C8870D66EFF8DFE01FCBBFDF1FCE7" box="[429,549,844,870]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
Male, Cape River, Pentland
<geoCoordinate id="9AD35015D66EFF8DFC06FCBBFBE9FCE7" box="[938,1085,844,870]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="-20.441666">20°26'30&quot;S</geoCoordinate>
<geoCoordinate id="9AD35015D66EFF8DFBEAFCBBFB3AFCE7" box="[1094,1262,844,870]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="145.53055">145°31'50&quot;E</geoCoordinate>
, northeastern Queensland,
<date id="8B591012D66EFF8DFE9CFC85FE0BFC0D" box="[304,479,882,908]" pageId="22" pageNumber="23" value="1997-08-04">4 August 1997</date>
, R. Raven (QM
<accessionNumber id="E0B4AB31D66EFF8DFD0CFC85FD2AFC0D" box="[672,766,882,908]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S31422" pageId="22" pageNumber="23" type="EnaNcbi">S31422</accessionNumber>
).
</paragraph>
<paragraph id="FF5836D2D66EFF8DFF6AFC6EFC7CFC32" blockId="22.[151,1438,843,1991]" box="[198,936,921,947]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF6AFC6EFEECFC32" bold="true" box="[198,312,921,947]" italics="true" pageId="22" pageNumber="23">
<typeStatus id="205C8870D66EFF8DFF6AFC6EFEE0FC32" box="[198,308,921,947]" pageId="22" pageNumber="23" type="paratype">Paratype</typeStatus>
.
</emphasis>
Allotype female, taken with
<typeStatus id="205C8870D66EFF8DFD22FC6EFD20FC32" box="[654,756,921,947]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(QM
<accessionNumber id="E0B4AB31D66EFF8DFC91FC6EFC4FFC32" box="[829,923,921,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S31423" pageId="22" pageNumber="23" type="EnaNcbi">S31423</accessionNumber>
).
</paragraph>
<paragraph id="FF5836D2D66EFF8DFF6AFC48FC27FBF5" blockId="22.[151,1438,843,1991]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF6AFC48FCF3FC58" bold="true" box="[198,807,959,985]" pageId="22" pageNumber="23">
Other material examined.
<collectingCountry id="87F07642D66EFF8DFDBCFC48FD5EFC58" box="[528,650,959,985]" name="Australia" pageId="22" pageNumber="23">Australia</collectingCountry>
:
<emphasis id="CD93EAC0D66EFF8DFD3AFC48FCF3FC58" bold="true" box="[662,807,959,985]" italics="true" pageId="22" pageNumber="23">Queensland</emphasis>
</emphasis>
: Female, same data as
<typeStatus id="205C8870D66EFF8DFB99FC37FB4FFC5B" box="[1077,1179,960,986]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(QM
<accessionNumber id="E0B4AB31D66EFF8DFB4EFC37FA95FC5B" box="[1250,1345,960,986]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S31424" pageId="22" pageNumber="23" type="EnaNcbi">S31424</accessionNumber>
).
<emphasis id="CD93EAC0D66EFF8DFAF8FC48FED3FB81" bold="true" italics="true" pageId="22" pageNumber="23">
South
<collectingCountry id="87F07642D66EFF8DFF3BFC11FED3FB81" box="[151,263,998,1024]" name="Australia" pageId="22" pageNumber="23">Australia</collectingCountry>
</emphasis>
: male, Trinity Gardens, Adelaide,
<geoCoordinate id="9AD35015D66EFF8DFD33FC11FD2AFB81" box="[671,766,998,1024]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-34.933334">34°56'S</geoCoordinate>
,
<geoCoordinate id="9AD35015D66EFF8DFCA6FC11FCACFB81" box="[778,888,998,1024]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="138.61667">138°37'E</geoCoordinate>
,
<date id="8B591012D66EFF8DFC29FC11FB98FB81" box="[901,1100,998,1024]" pageId="22" pageNumber="23" value="1988-10-15">15 October 1988</date>
, D. Hirst (
<collectionCode id="99F6AE17D66EFF8DFB6BFC11FACAFB81" LSID="urn:lsid:biocol.org:col:34982" box="[1223,1310,998,1024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="22" pageNumber="23">SAMA</collectionCode>
<collectionCode id="99F6AE17D66EFF8DFA89FC11FA86FB81" box="[1317,1362,998,1024]" pageId="22" pageNumber="23">NN</collectionCode>
6192); female, same data but
<date id="8B591012D66EFF8DFE0DFBFAFD82FBA6" box="[417,598,1037,1063]" pageId="22" pageNumber="23" value="1988-03-12">12 March 1988</date>
(
<collectionCode id="99F6AE17D66EFF8DFDCAFBFAFD69FBA6" LSID="urn:lsid:biocol.org:col:34982" box="[614,701,1037,1063]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="22" pageNumber="23">SAMA</collectionCode>
<collectionCode id="99F6AE17D66EFF8DFD68FBFAFD25FBA6" box="[708,753,1037,1063]" pageId="22" pageNumber="23">NN</collectionCode>
61949); female, Nuriootpa,
<geoCoordinate id="9AD35015D66EFF8DFB94FBFAFB47FBA6" box="[1080,1171,1037,1063]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-34.483334">34°29'S</geoCoordinate>
<geoCoordinate id="9AD35015D66EFF8DFB30FBFAFADFFBA6" box="[1180,1291,1037,1063]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="139.0">139°00'E</geoCoordinate>
,
<date id="8B591012D66EFF8DFAB4FBFAFF0EFBCF" pageId="22" pageNumber="23" value="1991-10-27">27 October 1991</date>
, A. F. Longbottom (
<collectionCode id="99F6AE17D66EFF8DFE4CFBC3FDFBFBCF" LSID="urn:lsid:biocol.org:col:34981" box="[480,559,1076,1102]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="23">WAM</collectionCode>
98/1693); female, Kapunda,
<geoCoordinate id="9AD35015D66EFF8DFC07FBC3FBDBFBCF" box="[939,1039,1076,1102]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-34.35">34°21'S</geoCoordinate>
<geoCoordinate id="9AD35015D66EFF8DFBB4FBC3FB45FBCF" box="[1048,1169,1076,1102]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="138.91667">138°55'E</geoCoordinate>
,
<date id="8B591012D66EFF8DFB0CFBC3FA9BFBCF" box="[1184,1359,1076,1102]" pageId="22" pageNumber="23" value="1995-05-28">28 May 1995</date>
, A. F. Longbottom, under rocks at copper/barite mine, S.1786 (
<collectionCode id="99F6AE17D66EFF8DFC9CFBADFCAEFBF5" LSID="urn:lsid:biocol.org:col:34981" box="[816,890,1114,1140]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="23">WAM</collectionCode>
98/1697).
</paragraph>
</subSubSection>
<subSubSection id="B7FD6559D66EFF8DFF6AFB77FAB7FA8E" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="FF5836D2D66EFF8DFF6AFB77FAB7FA8E" blockId="22.[151,1438,843,1991]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF6AFB77FEEBFB1B" bold="true" box="[198,319,1152,1178]" pageId="22" pageNumber="23">Diagnosis</emphasis>
. Males differ from those of
<taxonomicName id="38E74D51D66EFF8DFD22FB76FCC8FB1B" box="[654,796,1153,1179]" class="Arachnida" family="Miturgidae" genus="Mitzoruga" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="CD93EAC0D66EFF8DFD22FB76FD72FB1B" box="[654,678,1153,1178]" italics="true" pageId="22" pageNumber="23">M</emphasis>
.
<emphasis id="CD93EAC0D66EFF8DFD1AFB76FCC8FB1B" box="[694,796,1153,1178]" italics="true" pageId="22" pageNumber="23">insularis</emphasis>
</taxonomicName>
in the much broader tripartite median apophysis (
<figureCitation id="67DC2A57D66EFF8DFAC1FB76FF61FB43" captionStart="FIGURE 15" captionStartId="24.[151,256,1694,1718]" captionTargetBox="[151,1436,387,1669]" captionTargetId="figure@24.[151,1436,387,1670]" captionTargetPageId="24" captionText="FIGURE 15. Mitzoruga elapines sp. nov., palpal tibia and cymbium, scanning electron micrographs: a, bulb, ventral view; b, cymbium and tibia, retrolateral view; c, RTA, retrolateral view; d, median apophysis, ventral view." httpUri="https://zenodo.org/record/186366/files/figure.png" pageId="22" pageNumber="23">Fig. 15</figureCitation>
d), the narrower base of embolus (
<figureCitation id="67DC2A57D66EFF8DFDE7FB5FFD75FB43" box="[587,673,1192,1218]" captionStart="FIGURE 15" captionStartId="24.[151,256,1694,1718]" captionTargetBox="[151,1436,387,1669]" captionTargetId="figure@24.[151,1436,387,1670]" captionTargetPageId="24" captionText="FIGURE 15. Mitzoruga elapines sp. nov., palpal tibia and cymbium, scanning electron micrographs: a, bulb, ventral view; b, cymbium and tibia, retrolateral view; c, RTA, retrolateral view; d, median apophysis, ventral view." httpUri="https://zenodo.org/record/186366/files/figure.png" pageId="22" pageNumber="23">Fig. 15</figureCitation>
a), the deeply incised closely juxtaposed apices of the RTA (
<figureCitation id="67DC2A57D66EFF8DFAC0FB5FFF60FB69" captionStart="FIGURE 15" captionStartId="24.[151,256,1694,1718]" captionTargetBox="[151,1436,387,1669]" captionTargetId="figure@24.[151,1436,387,1670]" captionTargetPageId="24" captionText="FIGURE 15. Mitzoruga elapines sp. nov., palpal tibia and cymbium, scanning electron micrographs: a, bulb, ventral view; b, cymbium and tibia, retrolateral view; c, RTA, retrolateral view; d, median apophysis, ventral view." httpUri="https://zenodo.org/record/186366/files/figure.png" pageId="22" pageNumber="23">Fig. 15</figureCitation>
c) and the presence of thick setae on coxae IIII as well as coxae IV; females differ in the relatively larger basal lobe of the spermathecae (
<figureCitation id="67DC2A57D66EFF8DFDA2FB02FDB6FA8E" box="[526,610,1269,1295]" captionStart="FIGURE 16" captionStartId="25.[151,255,1238,1262]" captionTargetBox="[151,1433,690,1200]" captionTargetId="figure@25.[151,1436,661,1213]" captionTargetPageId="25" captionText="FIGURE 16. Mitzoruga elapines sp. nov., female WAM 98 / 1697: a, b. epigyne, external (a) and internal (b) views." httpUri="https://zenodo.org/record/186367/files/figure.png" pageId="22" pageNumber="23">Fig. 16</figureCitation>
b) giving a more block-like appearance to the spermathecal folds.
</paragraph>
</subSubSection>
<subSubSection id="B7FD6559D66EFF8DFF6AFAECFE34FADC" pageId="22" pageNumber="23" type="etymology">
<paragraph id="FF5836D2D66EFF8DFF6AFAECFE34FADC" blockId="22.[151,1438,843,1991]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF6AFAECFE9FFAB4" bold="true" box="[198,331,1307,1333]" pageId="22" pageNumber="23">Etymology</emphasis>
. From elapine, alluding to the long pair of front fangs of the most venomous snakes of the family
<taxonomicName id="38E74D51D66EFF8DFF47FAB4FE08FADC" authority="Boie, 1827" authorityName="Boie" authorityYear="1827" box="[235,476,1347,1373]" class="Reptilia" family="Elapidae" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="family">Elapidae Boie, 1827</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B7FD6559D66EFF82FF6AFA9FFD33FE66" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" type="description">
<paragraph id="FF5836D2D66EFF8DFF6AFA9FFD6FFA02" blockId="22.[151,1438,843,1991]" box="[198,699,1384,1411]" pageId="22" pageNumber="23">
<emphasis id="CD93EAC0D66EFF8DFF6AFA9FFE83FA03" bold="true" box="[198,343,1384,1410]" pageId="22" pageNumber="23">Description</emphasis>
. Male (
<typeStatus id="205C8870D66EFF8DFE1DFA9EFDCDFA02" box="[433,537,1385,1411]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
QM
<accessionNumber id="E0B4AB31D66EFF8DFDFBFA9EFD67FA02" box="[599,691,1385,1411]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S31422" pageId="22" pageNumber="23" type="EnaNcbi">S31422</accessionNumber>
)
</paragraph>
<paragraph id="FF5836D2D66EFF83FF69FA67FE5BFECD" blockId="22.[151,1438,843,1991]" lastBlockId="24.[151,1436,152,332]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
Carapace 4.00 long, 2.48 wide. Abdomen 4.32 long, 2.64 wide.
<emphasis id="CD93EAC0D66EFF8DFC19FA67FBDCFA28" box="[949,1032,1424,1449]" italics="true" pageId="22" pageNumber="23">Colour</emphasis>
(
<figureCitation id="67DC2A57D66EFF8DFBB4FA67FBA5FA2B" box="[1048,1137,1424,1450]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="22" pageNumber="23">Figs 14</figureCitation>
a, b). (seemingly recently moulted) Carapace pallid with 2 narrow dark lateral bands and pair of wider dark bands from behind PLE more or less straight until they converge slightly at posterior declivity; eye region black. Abdomen dorsally pallid with narrow “V anteromedially breaking up into 3 irregular spots in posterior third; lateral of “V broader, irregular longitudinal bands. Legs concolorous with carapace, no bands. Sternum, maxillae and labium pallid; abdomen ventrally pallid with light black mottling strong laterally and posteriorly but weakening centrally.
<emphasis id="CD93EAC0D66EFF8DFE30F98FFDCFF910" box="[412,539,1656,1681]" italics="true" pageId="22" pageNumber="23">Carapace.</emphasis>
Long black hairs on margins; short fine on carapace; long setae between AME; clypeus rounded distinct.
<emphasis id="CD93EAC0D66EFF8DFDBFF968FD9EF939" box="[531,586,1695,1720]" italics="true" pageId="22" pageNumber="23">Eyes</emphasis>
. AME: ALE: PME: PLE, 10: 11: 10: 11. ALE much closer to AME than AME apart; PME as far apart as from PLE, ALE slightly larger and lower (in anterior view) than AMEall directed to front and side. Anterior row clearly recurved; posterior row strongly recurved or in two rows, with PLE directed to side and up.
<emphasis id="CD93EAC0D66EFF8DFE40F8E4FDA1F8AD" box="[492,629,1811,1836]" italics="true" pageId="22" pageNumber="23">Chelicerae.</emphasis>
Small, porrect; dentition 3P, 2R. Fangs short, transverse. Boss small but distinct. Maxillae rounded rectangular with triangular, shallow depression diagonally from corner to distal edge. Labium short, shield-shaped. Sternum (
<figureCitation id="67DC2A57D66EFF8DFD10F897FCD8F8FB" box="[700,780,1888,1914]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="22" pageNumber="23">Fig 14</figureCitation>
h) cordate; setal patterns fine radial, inwardly directed setae.
<emphasis id="CD93EAC0D66EFF8DFF4DF870FEF5F821" box="[225,289,1927,1952]" italics="true" pageId="22" pageNumber="23">Legs.</emphasis>
Coxal bases with slight triangular projections entally, 2030 short thorn-like spines ventrally on retroventral half of coxae IIV, more numerous on IV (
<figureCitation id="67DC2A57D66EFF8DFCB6F85AFCA7F846" box="[794,883,1965,1991]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="22" pageNumber="23">Figs 14</figureCitation>
f, h). Trichobothria in
<emphasis id="CD93EAC0D66EFF8DFBD4F85AFB40F847" box="[1144,1172,1965,1990]" italics="true" pageId="22" pageNumber="23">ca</emphasis>
. 2 rows each of 810 for length of tibiae, metatarsi and tarsi. Trochanteral notches shallow, symetrical, as wide as deep on IV. I: 3.23, 1.38, 3.00, 2.54, 2.00, 12.15. II: 3.00, 1.38, 2.46, 2.15, 1.77, 10.75. III: 2.61, 1.15, 2.00, 1.92, 1.38, 9.06. IV: 3.61, 1.31, 3.31, 3.15, 1.54, 12.92. Palp: 1.54, 0.77, 0.69, , 1.31, 4.31.
<taxonomicName id="38E74D51D66FFF8CFB89F9E8FB51F9B9" box="[1061,1157,1567,1592]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="CD93EAC0D66FFF8CFB89F9E8FB51F9B9" box="[1061,1157,1567,1592]" italics="true" pageId="23" pageNumber="24">Scopula</emphasis>
</taxonomicName>
. Tarsi III entire, hairs long, not dense, not obscuring cuticle; IV with line of emergent setae. Metatarsi III weak, in 2 lateral bands for distal two-thirds and also ventral for distal quarter; weak on distal quarter on III, IV and tibiae.
<emphasis id="CD93EAC0D66FFF8CFABEF99BFAB2F904" box="[1298,1382,1644,1669]" italics="true" pageId="23" pageNumber="24">Spines.</emphasis>
I: fe pv1p1d3r1; pa 0; ti p2r
<date id="8B591012D66FFF8CFE03F964FE19F92C" box="[431,461,1683,1709]" pageId="23" pageNumber="24">2v</date>
2.2.2; me p1r
<date id="8B591012D66FFF8CFDC1F964FD5DF92C" box="[621,649,1683,1709]" pageId="23" pageNumber="24">1v</date>
2.2. II: fe pv1p2d3r2; pa 0; ti p2d1 r
<date id="8B591012D66FFF8CFBE8F964FBB6F92C" box="[1092,1122,1683,1709]" pageId="23" pageNumber="24">2v</date>
2.2.2; me p1r
<date id="8B591012D66FFF8CFAAFF964FACAF92C" box="[1283,1310,1683,1709]" pageId="23" pageNumber="24">1v</date>
2.2. III: fe p4d3r4; pa p1; ti p2d2.1r
<date id="8B591012D66FFF8CFE75F94DFE2DF955" box="[473,505,1722,1748]" pageId="23" pageNumber="24">2v</date>
2.2.2; me p1.2.1r2.1.1 v2.2.1. IV: fe p4d3r2; pa 0; ti p2d2.1r
<date id="8B591012D66FFF8CFAA9F94DFAF1F955" box="[1285,1317,1722,1748]" pageId="23" pageNumber="24">2v</date>
2.2.2; me p2.2.1r2.2.2v2.2.1.1. Palp: fe p1d1.2; pa p1; ti p3.
<emphasis id="CD93EAC0D66FFF8CFD59F916FC93F97B" box="[757,839,1761,1786]" italics="true" pageId="23" pageNumber="24">Claws.</emphasis>
Long, with 3 long and one short basal teeth; tufts (
<figureCitation id="67DC2A57D66FFF8CFF33F8F0FF21F8A0" box="[159,245,1799,1825]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="23" pageNumber="24">
Fig.
<date id="8B591012D66FFF8CFF7BF8F0FF21F8A0" box="[215,245,1799,1825]" pageId="23" pageNumber="24" value="1914-01">14</date>
</figureCitation>
i) strong but small, paired and conical.
<emphasis id="CD93EAC0D66FFF8CFD68F8F0FCE7F8A1" box="[708,819,1799,1824]" italics="true" pageId="23" pageNumber="24">Abdomen</emphasis>
. Tracheal spiracle small, near spinnerets. ALS short with domed tips. PMS cylindrical. PLS small with slightly elongated apical segment, orange with large spigot.
<emphasis id="CD93EAC0D66FFF8CFF3BF8A3FF1AF8EC" box="[151,206,1876,1901]" italics="true" pageId="23" pageNumber="24">Palp</emphasis>
(
<figureCitation id="67DC2A57D66FFF8CFF72F8A3FEE9F8EF" box="[222,317,1876,1902]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="23" pageNumber="24">Figs. 14</figureCitation>
be, 15). Tibia short with long triangular prong on prolateral corner, dorsal depression across to large bifid process on retrolateral edge; RTA a large cone with unsclerotised distal face transversed by two keels and slender sinuous prong dorsally off face of median process (
<figureCitation id="67DC2A57D66FFF8CFC4AF855FBEDF83D" box="[998,1081,1954,1980]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="23" pageNumber="24">Fig 14</figureCitation>
e); retroventral collar of tibia sclerotised with distinct erect cone and narrow notch beside base of RTA. Subtegulum prolateral and roughly triangular rounded ridge distally with sclerotised collariform conductor. Median apophysis subapico-central, flat, shield-like, with small medial process distally and vertical claw-like process retrolateral of it. Embolus origin small, basal; embolus tapers quickly runs clockwise arising retrodistally. Cymbium asymetrical with short rounded apex and distinct wide sclerotised groove retrolaterally and with strong comb of curved setae ectally (
<figureCitation id="67DC2A57D660FF83FF59FEC5FE82FECD" box="[245,342,306,332]" captionStart="FIGURE 14" captionStartId="23.[151,255,1246,1270]" captionTargetBox="[151,1435,194,1221]" captionTargetId="figure@23.[151,1436,194,1222]" captionTargetPageId="23" captionText="FIGURE 14. Mitzoruga elapines sp. nov., male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98 / 1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot; thorns &quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2 mm for a, g (same scale); 0.5 mm for b, e (same scale); 2 mm for h, 1 mm for f, j." httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="24" pageNumber="25">Figs. 14</figureCitation>
c, d).
</paragraph>
<caption id="AB98665AD66FFF8CFF3BFB29FECAFA22" httpUri="https://zenodo.org/record/186365/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1435,194,1221]" targetPageId="23">
<paragraph id="FF5836D2D66FFF8CFF3BFB29FECAFA22" blockId="23.[151,1437,1246,1443]" pageId="23" pageNumber="24">
<emphasis id="CD93EAC0D66FFF8CFF3BFB29FEF3FB77" bold="true" box="[151,295,1246,1270]" pageId="23" pageNumber="24">FIGURE 14.</emphasis>
<taxonomicName id="38E74D51D66FFF8CFE82FB28FE29FB77" box="[302,509,1247,1270]" class="Arachnida" family="Miturgidae" genus="Mitzoruga" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elapines" status="sp. nov.">
<emphasis id="CD93EAC0D66FFF8CFE82FB28FE29FB77" box="[302,509,1247,1270]" italics="true" pageId="23" pageNumber="24">Mitzoruga elapines</emphasis>
</taxonomicName>
<emphasis id="CD93EAC0D66FFF8CFDA8FB29FD8FFB77" bold="true" box="[516,603,1246,1270]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="D6A057BBD66FFF8CFDA8FB29FD8FFB77" box="[516,603,1246,1270]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: a, carapace, dorsal view; inset shows grate-shaped tapetum of PME of WAM 98/1693; b, palpal tibia and bulb, ventral view; c, tips of enlarged cymbial setae; d, shafts of enlarged cymbial setae; e, palpal tibia and bulb, retrolateral view; f, coxae III, IV, showing &quot;thorns&quot;, ventral view; g, abdomen, dorsal view; h, sternum, maxillae and coxae, ventral view; I, claws and claw tufts; j, patella and tibia I, prolateral view, showing long bristles apically and distally on patella. Scale line: 2mm for a, g (same scale); 0.5 mm for b, e (same scale); 2mm for h, 1mm for f, j.
</paragraph>
</caption>
<caption id="AB98665AD660FF83FF3BF969FADAF959" httpUri="https://zenodo.org/record/186366/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1436,387,1669]" targetPageId="24">
<paragraph id="FF5836D2D660FF83FF3BF969FADAF959" blockId="24.[151,1437,1694,1752]" pageId="24" pageNumber="25">
<emphasis id="CD93EAC0D660FF83FF3BF969FEFDF937" bold="true" box="[151,297,1694,1718]" pageId="24" pageNumber="25">FIGURE 15.</emphasis>
<taxonomicName id="38E74D51D660FF83FE9DF969FDD1F934" box="[305,517,1694,1717]" class="Arachnida" family="Miturgidae" genus="Mitzoruga" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="elapines" status="sp. nov.">
<emphasis id="CD93EAC0D660FF83FE9DF969FDD1F934" box="[305,517,1694,1717]" italics="true" pageId="24" pageNumber="25">Mitzoruga elapines</emphasis>
</taxonomicName>
<emphasis id="CD93EAC0D660FF83FDA0F969FDB0F937" bold="true" box="[524,612,1694,1718]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="D6A057BBD660FF83FDA0F969FDB0F937" box="[524,612,1694,1718]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, palpal tibia and cymbium, scanning electron micrographs: a, bulb, ventral view; b, cymbium and tibia, retrolateral view; c, RTA, retrolateral view; d, median apophysis, ventral view.
</paragraph>
</caption>
<paragraph id="FF5836D2D660FF83FF69F8F1FDE7F8A1" blockId="24.[151,1437,1798,2018]" box="[197,563,1798,1824]" pageId="24" pageNumber="25">
Female (allotype, QM
<accessionNumber id="E0B4AB31D660FF83FE7CF8F1FDFFF8A1" box="[464,555,1798,1824]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S31423" pageId="24" pageNumber="25" type="EnaNcbi">S31423</accessionNumber>
)
</paragraph>
<paragraph id="FF5836D2D660FF82FF69F8DAFD33FE66" blockId="24.[151,1437,1798,2018]" lastBlockId="25.[151,1437,152,603]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Carapace 3.92 long, 3.04 wide. Abdomen 5.36 long, 3.20 wide.
<emphasis id="CD93EAC0D660FF83FC77F8DAFBE2F8C7" box="[987,1078,1837,1862]" italics="true" pageId="24" pageNumber="25">Colour.</emphasis>
Pattern like male but darker, stronger. Carapace margin with irregular &quot;intrusions&quot; and with narrow grey medial band through eyes and fovea. Abdomen dorsally dark with irregular pallid “V breaking up into dots posteriorly. Legs dark brown with orange brown metatarsi and tarsi.
<emphasis id="CD93EAC0D660FF83FDD2F856FD6CF83B" box="[638,696,1953,1978]" italics="true" pageId="24" pageNumber="25">Eyes</emphasis>
. AME:ALE:PME:PLE, 7:11:11:13. Posterior row recurved.
<emphasis id="CD93EAC0D660FF83FF3BF83FFEFCF860" box="[151,296,1992,2017]" italics="true" pageId="24" pageNumber="25">Chelicerae.</emphasis>
Like male but stout, same dentition with pallid region anteriorly above fang.
<emphasis id="CD93EAC0D660FF83FAAEF83FFAE9F860" box="[1282,1341,1992,2017]" italics="true" pageId="24" pageNumber="25">Legs</emphasis>
. Coxae ventrally without thorn-like spines.
<emphasis id="CD93EAC0D661FF82FD96FF6FFD57FF30" box="[570,643,152,177]" italics="true" pageId="25" pageNumber="26">Claws</emphasis>
. Tufts like male; palpal claws with 3 teeth but no tuft. I: 2.69, 1.23, 2.46, 1.92, 1.38, 9.68. II: 2.54, 1.23, 2.61, 2.00, 1.54, 9.92. III: 2.54, 1.00, 1.92, 1.85, 1.23, 8.54. IV: 3.23, 1.31, 3.15, 3.46, 1.54, 12.69. Palp: 1.00, 0.61, 0.69, 1.00, 3.30.
<emphasis id="CD93EAC0D661FF82FC29FF12FC05FF7F" box="[901,977,229,254]" italics="true" pageId="25" pageNumber="26">Spines</emphasis>
. I: fe pv1p1d2; pa0; ti v2.2.2w; me
<date id="8B591012D661FF82FAD2FF12FA4FFF7E" box="[1406,1435,229,255]" pageId="25" pageNumber="26">v2</date>
basal. II: fe p2d2r1; pa0; ti v1.2.2w; me
<date id="8B591012D661FF82FDD8FEFBFD45FEA7" box="[628,657,268,294]" pageId="25" pageNumber="26">v2</date>
basal. III: fe p4d3r2; pa0; ti p2d2r
<date id="8B591012D661FF82FB86FEFBFB93FEA7" box="[1066,1095,268,294]" pageId="25" pageNumber="26">2v</date>
2.2.2; me p2.1r1.2.1v2.1. IV: fe p2d3r1; pa0; ti p2d2.1r
<date id="8B591012D661FF82FE6AFEC5FE37FECD" box="[454,483,306,332]" pageId="25" pageNumber="26">2v</date>
3.2.2; me p1.2.2r1.2.2v2.2.1. Palp: fe p1d1.2; pa p1; ti p3d2; ta p3r
<date id="8B591012D661FF82FB51FEC5FACFFECD" box="[1277,1307,306,332]" pageId="25" pageNumber="26">3v</date>
2.
<taxonomicName id="38E74D51D661FF82FA9AFEC5FA40FECA" box="[1334,1428,306,331]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="CD93EAC0D661FF82FA9AFEC5FA40FECA" box="[1334,1428,306,331]" italics="true" pageId="25" pageNumber="26">Scopula</emphasis>
</taxonomicName>
. Dense, almost obscures cuticle on tarsi IIV, entire on IIII, divided by 12 lines of setae on IV. Metatarsi III full, dense, entire, except for space around spines; III, IV divided into 3 parts.
<taxonomicName id="38E74D51D661FF82FBECFE77FB4BFE1B" box="[1088,1183,384,410]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="37" phylum="Arthropoda" rank="genus">Scopula</taxonomicName>
of short hairs also in distal half of tibiae I, II, in 3 bands on III, and few scattered setae on IV.
<emphasis id="CD93EAC0D661FF82FB88FE50FB5CFE41" box="[1060,1160,423,448]" italics="true" pageId="25" pageNumber="26">Epigyne</emphasis>
(
<figureCitation id="67DC2A57D661FF82FB36FE51FB2DFE41" box="[1178,1273,422,448]" captionStart="FIGURE 16" captionStartId="25.[151,255,1238,1262]" captionTargetBox="[151,1433,690,1200]" captionTargetId="figure@25.[151,1436,661,1213]" captionTargetPageId="25" captionText="FIGURE 16. Mitzoruga elapines sp. nov., female WAM 98 / 1697: a, b. epigyne, external (a) and internal (b) views." httpUri="https://zenodo.org/record/186367/files/figure.png" pageId="25" pageNumber="26">Figs 16</figureCitation>
a, b). Narrow anterior medial ridge and wide septum posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="B7FD6559D661FF82FF69FE04FDB9FDDA" pageId="25" pageNumber="26" type="distribution">
<paragraph id="FF5836D2D661FF82FF69FE04FDB9FDDA" blockId="25.[151,1437,152,603]" pageId="25" pageNumber="26">
<emphasis id="CD93EAC0D661FF82FF69FE04FE2EFD8C" bold="true" box="[197,506,499,525]" pageId="25" pageNumber="26">Distribution and habitat</emphasis>
(
<figureCitation id="67DC2A57D661FF82FDA0FE03FDB1FD8F" box="[524,613,500,526]" captionStart="FIGURE 25" captionStartId="36.[151,255,1809,1833]" captionTargetBox="[158,1430,1193,1764]" captionTargetId="figure@36.[151,1436,1164,1784]" captionTargetPageId="36" captionText="FIGURE 25. Distribution records of Miturgidae in Australia: a, Diaprograpta, Eupograpta, Mitzoruga and Nuliodon; b, all records of Miturgidae for Australia." httpUri="https://zenodo.org/record/186376/files/figure.png" pageId="25" pageNumber="26">Fig. 25</figureCitation>
a). Known only from Cape River, west of Charters Towers, northeast Queensland, and near Adelaide, South
<collectingCountry id="87F07642D661FF82FDC0FDEDFD0AFDB5" box="[620,734,538,564]" name="Australia" pageId="25" pageNumber="26">Australia</collectingCountry>
. At Cape River, the spiders were taken in litter around the bases of grasses in open eucalypt forest.
</paragraph>
</subSubSection>
</treatment>
</document>