447 lines
64 KiB
XML
447 lines
64 KiB
XML
<document id="1309D4133E25EF3959CA1C613D192917" ID="10.11646/zootaxa.4138.3.2" ID-DOI="10.11646/zootaxa.4138.3.2" ID-GBIF-Dataset="c785c9ac-b71c-4487-b770-35ebfe797c8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="258464" ID-ZooBank="DDF68118-EA96-49A5-B316-47814E19ED3D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468823189128" checkinUser="plazi" docAuthor="Leschen, Richard A. B., Escalona, Hermes E. & Elgueta, Mario" docDate="2016" docId="746C8797FFF1D000ACDDFD6AFDF6972A" docLanguage="en" docName="zootaxa.4138.3.2.pdf" docOrigin="Zootaxa 4138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ulodidae Pascoe 1869" docType="treatment" docVersion="9" lastPageNumber="449" masterDocId="8855FFEFFFF6D009AC4AFF96FF8B9636" masterDocTitle="Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)" masterLastPageNumber="473" masterPageNumber="441" pageNumber="448" updateTime="1698365200336" updateUser="plazi">
|
||
<mods:mods id="0353FC91156FD540E11074DA90AF417B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6A6B60266BE77F6D2314F5224ADCF344">
|
||
<mods:title id="2121EBDB62DC045E8305BCB15761B2F1">Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="17677F383B547F8C52AD2E4C1A87473F" type="personal">
|
||
<mods:role id="08238B5876D9A5CCF05F94D15DE308F5">
|
||
<mods:roleTerm id="D023D7DC858CF32AA4832AFF4B6DAA92">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F72333DC8B2F520CE1D29CB03EA1E6A6">Leschen, Richard A. B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2054F3C76E3BF3053B42BF81857DBE54" type="personal">
|
||
<mods:role id="8D2F8951FB49F2BFEC0E314FF618FB85">
|
||
<mods:roleTerm id="BA7D37002656D9D93F7743123122BCB9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="64D9DCEB5AAFF8967F0316EA84A4EA9F">Escalona, Hermes E.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EF1F20B2598F1ECD4F6A206BA5AB99C5" type="personal">
|
||
<mods:role id="3A87BEDB5EB4366C8D834680E2CB8513">
|
||
<mods:roleTerm id="7D6AA801217FD57ACDC2BE7A04A9FFE8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8AD8A88CD05AF27F9A35580ECCEBF900">Elgueta, Mario</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4A96BDE85380849B6C045A6AE740E980">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D61FE85FFB64FED426DD429C2B6AD5D4" type="host">
|
||
<mods:titleInfo id="CCC13D1A3E40445A8DE2ABED426DD837">
|
||
<mods:title id="F5F237EC4FA4E9BAC2F16B021C570706">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="366CD0E32C00554E62626CC4273FEB4F">
|
||
<mods:date id="31C8A5C82979976E36913A7AE3782629">2016</mods:date>
|
||
<mods:detail id="AF819C0E3E95A10ECFD71C91440455CA" type="volume">
|
||
<mods:number id="FFB3BBCEF681D5ED86C2B5F145D5C56D">4138</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="28F7A8D67A0AD5738FAB99EE4B69F57A" type="issue">
|
||
<mods:number id="9E280AEE2B7DBBE1D008CF2015B5937F">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="98036AD0130A0B78170369C3A6930BF4" unit="page">
|
||
<mods:start id="F3DB857DC06EE3B4344F6862F40979CF">441</mods:start>
|
||
<mods:end id="A5E3DBE355ED2AC22931C5E617255A68">473</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="62B7510C27E5D3CB592F44588C501636">journal article</mods:classification>
|
||
<mods:identifier id="2137FD3DBB867AC830FECBF52E3F215A" type="DOI">10.11646/zootaxa.4138.3.2</mods:identifier>
|
||
<mods:identifier id="7ADAF8C1F67E24E133795A47CFF33107" type="GBIF-Dataset">c785c9ac-b71c-4487-b770-35ebfe797c8a</mods:identifier>
|
||
<mods:identifier id="BDF3C54E009812A3BDCA6581BD6EC542" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="BAB82934AFA037841D1E4841F609107E" type="Zenodo-Dep">258464</mods:identifier>
|
||
<mods:identifier id="F0E1C3CA5510187EC175C274A6921911" type="ZooBank">DDF68118-EA96-49A5-B316-47814E19ED3D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="746C8797FFF1D000ACDDFD6AFDF6972A" ID-DOI="http://doi.org/10.5281/zenodo.5668193" ID-GBIF-Taxon="123195951" ID-Zenodo-Dep="5668193" LSID="urn:lsid:plazi:treatment:746C8797FFF1D000ACDDFD6AFDF6972A" httpUri="http://treatment.plazi.org/id/746C8797FFF1D000ACDDFD6AFDF6972A" lastPageId="9" lastPageNumber="449" pageId="7" pageNumber="448">
|
||
<subSubSection id="B4DF650AFFF1D00EACDDFD6AFE269520" box="[151,429,764,790]" pageId="7" pageNumber="448" type="nomenclature">
|
||
<paragraph id="FC7A3681FFF1D00EACDDFD6AFE269520" blockId="7.[151,429,764,790]" box="[151,429,764,790]" pageId="7" pageNumber="448">
|
||
<heading id="A73281EDFFF1D00EACDDFD6AFE269520" bold="true" box="[151,429,764,790]" fontSize="11" level="1" pageId="7" pageNumber="448" reason="1">
|
||
<taxonomicName id="3BC54D02FFF1D00EACDDFD6AFE269520" ID-CoL="HNF" ID-ENA="219433" authority="Pascoe, 1869" authorityName="Pascoe" authorityYear="1869" box="[151,429,764,790]" class="Insecta" family="Ulodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="family">
|
||
<emphasis id="CEB1EA93FFF1D00EACDDFD6AFE269520" bold="true" box="[151,429,764,790]" pageId="7" pageNumber="448">
|
||
Ulodidae
|
||
<bibRefCitation id="98544B70FFF1D00EAD44FD6AFE269520" author="Pascoe" box="[270,429,764,790]" pageId="7" pageNumber="448" refString="Pascoe, F. P. (1869) Descriptions of new genera and species of Tenebrionidae from Australia and Tasmania. Annals and Magazine of Natural History, (4) 3, 29 - 45, plate X." type="journal article" year="1869">Pascoe, 1869</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4DF650AFFF1D001ACDDFCD4FC5391A3" lastPageId="8" lastPageNumber="449" pageId="7" pageNumber="448" type="diagnosis">
|
||
<paragraph id="FC7A3681FFF1D00EACDDFCD4FD7995F0" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="448">
|
||
<emphasis id="CEB1EA93FFF1D00EACDDFCD4FED8956D" bold="true" box="[151,339,834,859]" pageId="7" pageNumber="448">Adult diagnosis</emphasis>
|
||
. Antennal insertions visible in dorsal view (
|
||
<figureCitation id="64FE2A04FFF1D00EAF13FCD4FC3D956D" box="[857,950,834,859]" captionStart="FIGURES 9 A – F" captionStartId="22.[151,264,1802,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@22.[151,1436,193,1780]" captionTargetPageId="22" captionText="FIGURES 9 A – F. Ulodes hispidus Pascoe: A – C, head, dorsal and ventral, C, detail; D – F, antenna: D, dorsal view; E, antennal club detail; F, antennal club." httpUri="https://zenodo.org/record/258474/files/figure.png" pageId="7" pageNumber="448">Fig. 9A</figureCitation>
|
||
). Prosternal process strongly and abruptly expanded at apex and meeting the hypomeron, procoxal cavities externally closed (
|
||
<figureCitation id="64FE2A04FFF1D00EA809FCF0FB299549" box="[1091,1186,870,895]" captionStart="FIGURES 10 A – D" captionStartId="23.[151,264,1767,1789]" captionTargetBox="[151,1436,193,1745]" captionTargetId="figure@23.[151,1436,193,1746]" captionTargetPageId="23" captionText="FIGURES 10 A – D. Ulodes hispidus Pascoe: A, pronotum; B, prothorax ventral; C, pterothorax and abdomen; D, abdomen." httpUri="https://zenodo.org/record/258475/files/figure.png" pageId="7" pageNumber="448">Fig.10B</figureCitation>
|
||
). Abdominal ventrites 1 and 2 connate (
|
||
<figureCitation id="64FE2A04FFF1D00EAD29FC1CFE4B9595" box="[355,448,906,931]" captionStart="FIGURES 7 A – N. A" captionStartId="20.[151,264,1802,1824]" captionTargetBox="[158,1422,199,1773]" captionTargetId="figure@20.[151,1436,193,1780]" captionTargetPageId="20" captionText="FIGURES 7 A – N. A, Pteroderes tuberosus Germain, head dorsal; B, Trachyderastes dipsaconiae Kaszab, maxilla; C, Brouniphylax sp., maxilla; D, Arthopus brouni Sharp, maxilla; E, Pteroderes tuberosus Germain, labium; F, Brouniphylax exiquus (Broun), antenna; G, Ganyme sp., prothorax frontal view. H – J, Metendosternite: H, Pteroderes tuberosus Germain; I, Brouniphylax exiquus (Broun); J, Trachyderas costulatum Fairmaire & Germain. K – L, Ulobostrichus monteithi sp. n., abdomen and detail; M, Pteroderes tuberosus Germain, abdomen; N, Brouniphylax sp., female ovipositor." httpUri="https://zenodo.org/record/258472/files/figure.png" pageId="7" pageNumber="448">Fig. 7K</figureCitation>
|
||
). Tarsal formula 5-5-4 or 4-4-4 (
|
||
<taxonomicName id="3BC54D02FFF1D00EAF0AFC1AFC039595" box="[832,904,908,931]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF0AFC1AFC039595" box="[832,904,908,931]" italics="true" pageId="7" pageNumber="448">Meryx</emphasis>
|
||
</taxonomicName>
|
||
). Aedeagus tenebrionoid (
|
||
<figureCitation id="64FE2A04FFF1D00EA8F0FC1CFAB99595" box="[1210,1330,906,931]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="7" pageNumber="448">Figs 8J–L</figureCitation>
|
||
); tegmen without alae and anterior edge of penis without struts.
|
||
</paragraph>
|
||
<paragraph id="FC7A3681FFF1D00EAC8DFC44FAA5924D" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="448">
|
||
<emphasis id="CEB1EA93FFF1D00EAC8DFC44FE1795DD" bold="true" box="[199,412,978,1003]" pageId="7" pageNumber="448">Adult description</emphasis>
|
||
. Length
|
||
<quantity id="3B3D9B64FFF1D00EAE40FC44FD2D95DC" box="[522,678,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="1.415" metricValueMax="2.45" metricValueMin="0.38" pageId="7" pageNumber="448" unit="mm" value="14.15" valueMax="24.5" valueMin="3.8">3.8–24.5 mm</quantity>
|
||
. Body about 1.3–2.5 times as long as wide, slightly flattened to strongly convex; sides oval to parallel. Surfaces glabrous (
|
||
<taxonomicName id="3BC54D02FFF1D00EAF69FC60FC019239" box="[803,906,1014,1039]" class="Insecta" family="Ulodidae" genus="Arthopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF69FC60FC019239" box="[803,906,1014,1039]" italics="true" pageId="7" pageNumber="448">Arthopus</emphasis>
|
||
</taxonomicName>
|
||
) or subglabrous (
|
||
<taxonomicName id="3BC54D02FFF1D00EA805FC60FB629239" box="[1103,1257,1014,1039]" class="Insecta" family="Ulodidae" genus="Ulocyphaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFF1D00EA805FC60FB629239" box="[1103,1257,1014,1039]" italics="true" pageId="7" pageNumber="448">Ulocyphaleus</emphasis>
|
||
</taxonomicName>
|
||
), setose (
|
||
<taxonomicName id="3BC54D02FFF1D00EA91EFC61FA179238" box="[1364,1436,1015,1038]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA91EFC61FA179238" box="[1364,1436,1015,1038]" italics="true" pageId="7" pageNumber="448">Meryx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EAC82FB8CFEF89205" box="[200,371,1050,1075]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAC82FB8CFEF89205" box="[200,371,1050,1075]" italics="true" pageId="7" pageNumber="448">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
) or clothed with distinct bristles or scale-like setae that are erect or adpressed to the cuticle; wax-debris encrustations present on specimens of
|
||
<taxonomicName id="3BC54D02FFF1D00EAF5AFBA8FBB19261" authority="Exohadrus" authorityName="Exohadrus" box="[784,1082,1086,1111]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF5AFBA8FC2C9261" box="[784,935,1086,1111]" italics="true" pageId="7" pageNumber="448">Brouniphylax</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFF1D00EAF8BFBA8FBB19261" box="[961,1082,1086,1111]" italics="true" pageId="7" pageNumber="448">Exohadrus</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<taxonomicName id="3BC54D02FFF1D00EA8E8FBA9FE9B924D" authority="Notocerastes, Pteroderes" authorityName="Notocerastes, Pteroderes" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA8E8FBA9FB629260" box="[1186,1257,1087,1110]" italics="true" pageId="7" pageNumber="448">Meryx</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFF1D00EA949FBA9FA1E9260" box="[1283,1429,1087,1110]" italics="true" pageId="7" pageNumber="448">Notocerastes</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFF1D00EACDDFBF4FE9B924D" box="[151,272,1122,1147]" italics="true" pageId="7" pageNumber="448">Pteroderes</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<emphasis id="CEB1EA93FFF1D00EAD2AFBF4FE64924D" box="[352,495,1122,1147]" italics="true" pageId="7" pageNumber="448">
|
||
<taxonomicName id="3BC54D02FFF1D00EAD2AFBF4FE67924D" box="[352,492,1122,1147]" class="Insecta" family="Tenebrionidae" genus="Syrphetodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">Syrphetodes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EAE6CFBF4FD4F924D" box="[550,708,1122,1147]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFF1D00EAE6CFBF4FD4F924D" box="[550,708,1122,1147]" italics="true" pageId="7" pageNumber="448">Ulobostrichus</emphasis>
|
||
</taxonomicName>
|
||
. Color mottled to unicolorous and sometimes metallic.
|
||
</paragraph>
|
||
<paragraph id="FC7A3681FFF1D00EAC8DFB10FD589159" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="448">
|
||
Head prognathous, often deeply imbedded into prothorax, not abruptly constricted posteriorly to form a neck, but a ridge behind the eye may be present; temples short or absent; frons impressed or not. Eyes (
|
||
<figureCitation id="64FE2A04FFF1D00EA95DFB3CFA1F92F5" box="[1303,1428,1194,1219]" captionStart="FIGURES 6 A – O. A – C" captionStartId="19.[151,264,1702,1724]" captionTargetBox="[159,1433,201,1671]" captionTargetId="figure@19.[151,1436,193,1680]" captionTargetPageId="19" captionText="FIGURES 6 A – O. A – C head dorsal: A, Syrphetodes marginatus Pascoe; B, Pteroderes tuberosus Germain; C, Ulocyphaleus valdivianus (Philippi & Philippi). D – G, head lateral: D, Meryx aequalis Blackburn; E, Pteroderes tuberosus Germain; F, Ganyme tuberculata Germain; G, Notocerastes blackburni Carter. H, Ulobostrichus monteithi sp. n., mid leg. I, Arthopus brouni Sharp, mid-claw. J – O, Maxillary palps: J, Ulocyphaleus valdivianus (Philippi & Philippi); K, Syrphetodes marginatus Pascoe; L, Ganyme sapphiro (Newman); M, Trachyderastes dipsaconiae Kaszab; N, Meryx rugosa Latreille, O, Pteroderes tuberosus Germain." httpUri="https://zenodo.org/record/258471/files/figure.png" pageId="7" pageNumber="448">Figs 6A–F</figureCitation>
|
||
) moderately to strongly (
|
||
<taxonomicName id="3BC54D02FFF1D00EAD8BFB58FDA292D1" box="[449,553,1230,1255]" class="Insecta" family="Zopheridae" genus="Phaennis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAD8BFB58FDA292D1" box="[449,553,1230,1255]" italics="true" pageId="7" pageNumber="448">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EAE39FB58FD6792D1" box="[627,748,1230,1255]" class="Insecta" family="Ulodidae" genus="Pteroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAE39FB58FD6792D1" box="[627,748,1230,1255]" italics="true" pageId="7" pageNumber="448">Pteroderes</emphasis>
|
||
</taxonomicName>
|
||
) protuberant, finely to coarsely facetted, usually entire, sometimes emarginate, rarely by a canthus or antennal insertion (
|
||
<taxonomicName id="3BC54D02FFF1D00EAFE6FB64FBDC933D" box="[940,1111,1266,1291]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAFE6FB64FBDC933D" box="[940,1111,1266,1291]" italics="true" pageId="7" pageNumber="448">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
), posteriorly emarginate in
|
||
<taxonomicName id="3BC54D02FFF1D00EACDDFA80FEA89319" box="[151,291,1302,1327]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EACDDFA80FEA89319" box="[151,291,1302,1327]" italics="true" pageId="7" pageNumber="448">Trachyderas</emphasis>
|
||
</taxonomicName>
|
||
, narrow in
|
||
<taxonomicName id="3BC54D02FFF1D00EADE7FA81FD819318" box="[429,522,1303,1326]" class="Insecta" family="Tenebrionidae" genus="Ganyme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EADE7FA81FD819318" box="[429,522,1303,1326]" italics="true" pageId="7" pageNumber="448">Ganyme</emphasis>
|
||
</taxonomicName>
|
||
and also strongly c-shaped in
|
||
<taxonomicName id="3BC54D02FFF1D00EAF3BFA80FB959319" box="[881,1054,1302,1327]" class="Insecta" family="Ulodidae" genus="Waitomophylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF3BFA80FB959319" box="[881,1054,1302,1327]" italics="true" pageId="7" pageNumber="448">Waitomophylax</emphasis>
|
||
</taxonomicName>
|
||
; interfacetal setae usually absent (present in
|
||
<taxonomicName id="3BC54D02FFF1D00EAD5FFAACFEF69365" box="[277,381,1338,1363]" class="Insecta" family="Zopheridae" genus="Phaennis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAD5FFAACFEF69365" box="[277,381,1338,1363]" italics="true" pageId="7" pageNumber="448">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
). Antennal insertions exposed from above, or slightly concealed and inserted sublaterally and hidden by a marginal deflexed ridge or cephalic horns (
|
||
<taxonomicName id="3BC54D02FFF1D00EAF4DFAC9FC129340" box="[775,921,1375,1398]" class="Insecta" family="Zopheridae" genus="Notocerastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF4DFAC9FC129340" box="[775,921,1375,1398]" italics="true" pageId="7" pageNumber="448">Notocerastes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EAF9BFAC8FBB29341" box="[977,1081,1374,1399]" class="Insecta" family="Zopheridae" genus="Phaennis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF9BFAC8FBB29341" box="[977,1081,1374,1399]" italics="true" pageId="7" pageNumber="448">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
); two anterolateral horns (male
|
||
<taxonomicName id="3BC54D02FFF1D00EACDDFA14FF7493AD" box="[151,255,1410,1435]" class="Insecta" family="Zopheridae" genus="Phaennis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EACDDFA14FF7493AD" box="[151,255,1410,1435]" italics="true" pageId="7" pageNumber="448">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="64FE2A04FFF1D00EAD45FA14FEE393AD" box="[271,360,1410,1435]" captionStart="FIGURES 6 A – O. A – C" captionStartId="19.[151,264,1702,1724]" captionTargetBox="[159,1433,201,1671]" captionTargetId="figure@19.[151,1436,193,1680]" captionTargetPageId="19" captionText="FIGURES 6 A – O. A – C head dorsal: A, Syrphetodes marginatus Pascoe; B, Pteroderes tuberosus Germain; C, Ulocyphaleus valdivianus (Philippi & Philippi). D – G, head lateral: D, Meryx aequalis Blackburn; E, Pteroderes tuberosus Germain; F, Ganyme tuberculata Germain; G, Notocerastes blackburni Carter. H, Ulobostrichus monteithi sp. n., mid leg. I, Arthopus brouni Sharp, mid-claw. J – O, Maxillary palps: J, Ulocyphaleus valdivianus (Philippi & Philippi); K, Syrphetodes marginatus Pascoe; L, Ganyme sapphiro (Newman); M, Trachyderastes dipsaconiae Kaszab; N, Meryx rugosa Latreille, O, Pteroderes tuberosus Germain." httpUri="https://zenodo.org/record/258471/files/figure.png" pageId="7" pageNumber="448">Fig. 6G</figureCitation>
|
||
) or one short median (
|
||
<taxonomicName id="3BC54D02FFF1D00EAE25FA12FC8A93AD" box="[623,769,1412,1435]" class="Insecta" family="Zopheridae" genus="Notocerastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAE25FA12FC8A93AD" box="[623,769,1412,1435]" italics="true" pageId="7" pageNumber="448">Notocerastes</emphasis>
|
||
</taxonomicName>
|
||
) horn present in the male. Frontoclypeal suture present or absent. Antennae usually extending posteriorly beyond basal margin of elytra (barely so in
|
||
<taxonomicName id="3BC54D02FFF1D00EA8C7FA30FA8D9389" box="[1165,1286,1446,1471]" class="Insecta" family="Ulodidae" genus="Pteroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA8C7FA30FA8D9389" box="[1165,1286,1446,1471]" italics="true" pageId="7" pageNumber="448">Pteroderes</emphasis>
|
||
</taxonomicName>
|
||
and not at all in
|
||
<taxonomicName id="3BC54D02FFF1D00EACFDFA5CFEDD93D5" box="[183,342,1482,1507]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFF1D00EACFDFA5CFEDD93D5" box="[183,342,1482,1507]" italics="true" pageId="7" pageNumber="448">Ulobostrichus</emphasis>
|
||
</taxonomicName>
|
||
); 11-segmented, usually capitate with 3-segmented club (4-segmented in
|
||
<taxonomicName id="3BC54D02FFF1D00EA8E6FA5CFADC93D5" box="[1196,1367,1482,1507]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA8E6FA5CFADC93D5" box="[1196,1367,1482,1507]" italics="true" pageId="7" pageNumber="448">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
), that may be weakly clubbed, rarely filiform (
|
||
<taxonomicName id="3BC54D02FFF1D00EAE22FA78FC369031" authority="Ulocyphaleus" authorityName="Ulocyphaleus" box="[616,957,1518,1543]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAE22FA78FC989031" box="[616,787,1518,1543]" italics="true" pageId="7" pageNumber="448">Trachyderastes</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFF1D00EAF69FA78FC369031" box="[803,957,1518,1543]" italics="true" pageId="7" pageNumber="448">Ulocyphaleus</emphasis>
|
||
</taxonomicName>
|
||
), a 10-segmented antenna with a weak 2- segmented clubbed is present in
|
||
<taxonomicName id="3BC54D02FFF1D00EAE5EF984FD38901D" box="[532,691,1554,1579]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFF1D00EAE5EF984FD38901D" box="[532,691,1554,1579]" italics="true" pageId="7" pageNumber="448">Ulobostrichus</emphasis>
|
||
</taxonomicName>
|
||
; sensory areas of antennomeres (usually 9–11) demarcated by a distinct rim in some taxa. Labrum visible and free, transverse to subquadrate and not concealed beneath clypeus. Mandible short and broad, bidentate; incisor edge simple or with a single tooth with well-developed mola and prostheca. Maxilla (
|
||
<figureCitation id="64FE2A04FFF1D00EAD30F9E8FE7390A1" box="[378,504,1662,1687]" captionStart="FIGURES 7 A – N. A" captionStartId="20.[151,264,1802,1824]" captionTargetBox="[158,1422,199,1773]" captionTargetId="figure@20.[151,1436,193,1780]" captionTargetPageId="20" captionText="FIGURES 7 A – N. A, Pteroderes tuberosus Germain, head dorsal; B, Trachyderastes dipsaconiae Kaszab, maxilla; C, Brouniphylax sp., maxilla; D, Arthopus brouni Sharp, maxilla; E, Pteroderes tuberosus Germain, labium; F, Brouniphylax exiquus (Broun), antenna; G, Ganyme sp., prothorax frontal view. H – J, Metendosternite: H, Pteroderes tuberosus Germain; I, Brouniphylax exiquus (Broun); J, Trachyderas costulatum Fairmaire & Germain. K – L, Ulobostrichus monteithi sp. n., abdomen and detail; M, Pteroderes tuberosus Germain, abdomen; N, Brouniphylax sp., female ovipositor." httpUri="https://zenodo.org/record/258472/files/figure.png" pageId="7" pageNumber="448">Figs 7B–D</figureCitation>
|
||
) with galea setose and lacinia with or without (
|
||
<taxonomicName id="3BC54D02FFF1D00EA851F9E9FBE890A0" box="[1051,1123,1663,1686]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA851F9E9FBE890A0" box="[1051,1123,1663,1686]" italics="true" pageId="7" pageNumber="448">Meryx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EA8D6F9E8FACC90A1" box="[1180,1351,1662,1687]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EA8D6F9E8FACC90A1" box="[1180,1351,1662,1687]" italics="true" pageId="7" pageNumber="448">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
) apical spines or uncus; apical palpomere cylindrical to fusiform, slightly expanded and truncate to subtriangulate and subcylindrical-curved (
|
||
<taxonomicName id="3BC54D02FFF1D00EADEBF950FD8390E9" box="[417,520,1734,1759]" class="Insecta" family="Ulodidae" genus="Arthopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EADEBF950FD8390E9" box="[417,520,1734,1759]" italics="true" pageId="7" pageNumber="448">Arthopus</emphasis>
|
||
</taxonomicName>
|
||
). Submentum without setose pit; ligula undivided to moderately emarginate; prementum sclerotized or not; apical labial palpomeres cylindrical to fusiform with variable widths. Genal ridges or projections absent; ventral part of the head expanded laterally below eye and forming an ocular shelf in
|
||
<taxonomicName id="3BC54D02FFF1D00EACDDF8A4FEA5917D" box="[151,302,1842,1867]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EACDDF8A4FEA5917D" box="[151,302,1842,1867]" italics="true" pageId="7" pageNumber="448">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
and subocular tubercles present in
|
||
<taxonomicName id="3BC54D02FFF1D00EAE8FF8A4FCF9917D" box="[709,882,1842,1867]" class="Insecta" family="Ulodidae" genus="Waitomophylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAE8FF8A4FCF9917D" box="[709,882,1842,1867]" italics="true" pageId="7" pageNumber="448">Waitomophylax</emphasis>
|
||
</taxonomicName>
|
||
. Gular sutures short or absent, widely separated and convergent, or well developed (
|
||
<taxonomicName id="3BC54D02FFF1D00EAE63F8C0FD489159" box="[553,707,1878,1903]" class="Insecta" family="Ulodidae" genus="Ulocyphaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFF1D00EAE63F8C0FD489159" box="[553,707,1878,1903]" italics="true" pageId="7" pageNumber="448">Ulocyphaleus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC7A3681FFF1D001AC8DF8EDFD4993F6" blockId="7.[151,1437,834,2011]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="449" pageId="7" pageNumber="448">
|
||
Pronotum 0.4–0.8 times as long as wide; widest anteriorly, at middle, or posteriorly with sides more or less straight to strongly curved, sometimes variously lobed (e.g.,
|
||
<taxonomicName id="3BC54D02FFF1D00EAF39F808FC679181" box="[883,1004,1950,1975]" class="Insecta" family="Ulodidae" genus="Pteroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF39F808FC679181" box="[883,1004,1950,1975]" italics="true" pageId="7" pageNumber="448">Pteroderes</emphasis>
|
||
</taxonomicName>
|
||
) and often explanate; disc variable, sometimes with tubercles, may be vaulted,
|
||
<taxonomicName id="3BC54D02FFF1D00EAECBF854FD6291ED" box="[641,745,1986,2011]" class="Insecta" family="Zopheridae" genus="Phaennis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAECBF854FD6291ED" box="[641,745,1986,2011]" italics="true" pageId="7" pageNumber="448">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFF1D00EAF6BF852FC3891ED" box="[801,947,1988,2011]" class="Insecta" family="Zopheridae" genus="Notocerastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFF1D00EAF6BF852FC3891ED" box="[801,947,1988,2011]" italics="true" pageId="7" pageNumber="448">Notocerastes</emphasis>
|
||
</taxonomicName>
|
||
with paired basal impressions or pits; base narrower than or equal to elytral bases. Lateral pronotal carinae complete, edges sharp or rounded, simple to coarsely dentate, with or without raised margin or bead; anterior and posterior angles acute or rounded; posterior edge straight, evenly rounded or sinuate. Antero- or ventrolateral portions of prothorax often without cavities or grooves, present in
|
||
<taxonomicName id="3BC54D02FFFED001AD3DFE92FE7B972B" box="[375,496,260,285]" class="Insecta" family="Ulodidae" genus="Exohadrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AD3DFE92FE7B972B" box="[375,496,260,285]" italics="true" pageId="8" pageNumber="449">Exohadrus</emphasis>
|
||
</taxonomicName>
|
||
. Prosternum in front of coxae shorter than or as long as prosternal process, flat to strongly convex. Prosternal process strongly and abruptly expanded at apex and meeting the hypomeron; flat, concave, or only slightly elevated or curved behind coxae. Notosternal sutures present, incomplete or absent. Procoxae not projecting below prosternum, lateral extensions short, rarely long and concealed (
|
||
<taxonomicName id="3BC54D02FFFED001A894FEF9FA0097BE" box="[1246,1419,367,392]" class="Insecta" family="Ulodidae" genus="Waitomophylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A894FEF9FA0097BE" box="[1246,1419,367,392]" italics="true" pageId="8" pageNumber="449">Waitomophylax</emphasis>
|
||
</taxonomicName>
|
||
). Procoxal cavities externally closed with postcoxal projection meeting prosternal process (
|
||
<figureCitation id="64FE2A04FFFED001A8E7FE02FA93979A" box="[1197,1304,404,429]" captionStart="FIGURES 10 A – D" captionStartId="23.[151,264,1767,1789]" captionTargetBox="[151,1436,193,1745]" captionTargetId="figure@23.[151,1436,193,1746]" captionTargetPageId="23" captionText="FIGURES 10 A – D. Ulodes hispidus Pascoe: A, pronotum; B, prothorax ventral; C, pterothorax and abdomen; D, abdomen." httpUri="https://zenodo.org/record/258475/files/figure.png" pageId="8" pageNumber="449">Fig. 10B</figureCitation>
|
||
); internally open; more or less circular or slightly transverse, narrowly to widely separated; protrochanteral notch absent or weakly indicated. Scutellar shield fully developed and abruptly elevated (
|
||
<figureCitation id="64FE2A04FFFED001AFA6FE4AFBDD97C2" box="[1004,1110,476,501]" captionStart="FIGURES 11 A – E" captionStartId="24.[151,264,1802,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 11 A – E. Ganyme tuberculatus (Carter): A-B, head dorsal and ventral; C, labium; D, scutellum; E, antenna." httpUri="https://zenodo.org/record/258476/files/figure.png" pageId="8" pageNumber="449">Fig. 11D</figureCitation>
|
||
), reduced (
|
||
<taxonomicName id="3BC54D02FFFED001A890FE4AFAFA97C3" box="[1242,1393,476,501]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A890FE4AFAFA97C3" box="[1242,1393,476,501]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
) or absent (
|
||
<taxonomicName id="3BC54D02FFFED001ACA4FE69FEEC942E" box="[238,359,511,536]" class="Insecta" family="Ulodidae" genus="Exohadrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001ACA4FE69FEEC942E" box="[238,359,511,536]" italics="true" pageId="8" pageNumber="449">Exohadrus</emphasis>
|
||
</taxonomicName>
|
||
). Elytra 0.85–2.1 times as long as combined width and 1.7–3.2 times as long as pronotum; sides explanate or convex and rarely declivous posteriorly; punctures penetrating deeply into the elytra forming a “window” or not, serially punctate or irregularly punctate; striole present or absent; surfaces smooth, carinate, or tuberculate; apices meeting or almost meeting at suture which is not deflected at apex; epipleuron usually complete, but vaguely defined (
|
||
<taxonomicName id="3BC54D02FFFED001AE5DFD19FD25949E" box="[535,686,655,680]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AE5DFD19FD25949E" box="[535,686,655,680]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
) and variously formed in
|
||
<taxonomicName id="3BC54D02FFFED001AFBCFD19FBF4949E" box="[1014,1151,655,680]" class="Insecta" family="Tenebrionidae" genus="Syrphetodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AFBCFD19FBF4949E" box="[1014,1151,655,680]" italics="true" pageId="8" pageNumber="449">Syrphetodes</emphasis>
|
||
</taxonomicName>
|
||
; lateral edge simple or microtuberculate. Mesoventrite usually separated by complete sutures from mesanepisterna (not in
|
||
<taxonomicName id="3BC54D02FFFED001A8B9FD22FA0194FB" box="[1267,1418,692,717]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A8B9FD22FA0194FB" box="[1267,1418,692,717]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
); anterior edge on same plane as metaventrite and without paired procoxal rests; anterior edge width variable, with or without median prosternal rest or with broadly transverse prothoracic rest. Mesanepisterna almost meeting at midline (
|
||
<taxonomicName id="3BC54D02FFFED001ACB7FCB7FECE950E" box="[253,325,801,824]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001ACB7FCB7FECE950E" box="[253,325,801,824]" italics="true" pageId="8" pageNumber="449">Meryx</emphasis>
|
||
</taxonomicName>
|
||
) or broadly separated (measured by its width to mesoventral process). Mesocoxal cavities circular to slightly transverse (
|
||
<figureCitation id="64FE2A04FFFED001ADEDFCD2FD9E956A" box="[423,533,836,861]" captionStart="FIGURES 10 A – D" captionStartId="23.[151,264,1767,1789]" captionTargetBox="[151,1436,193,1745]" captionTargetId="figure@23.[151,1436,193,1746]" captionTargetPageId="23" captionText="FIGURES 10 A – D. Ulodes hispidus Pascoe: A, pronotum; B, prothorax ventral; C, pterothorax and abdomen; D, abdomen." httpUri="https://zenodo.org/record/258475/files/figure.png" pageId="8" pageNumber="449">Fig. 10C</figureCitation>
|
||
); narrowly to widely separated at middle; open laterally, partly closed by mesepimeron; mesotrochantin always exposed; mesocoxa not conical and projecting. Mesometaventral junction simple or monocondylic. Metaventrite well-developed (longer than wide or quadrate) or reduced and transverse in apterous taxa; with discrimen present (long or relatively shortened) or absent, in the form of a groove in some taxa; postcoxal lines absent; exposed portion of metanepisternum elongate, triangulate in
|
||
<taxonomicName id="3BC54D02FFFED001A83DFC42FA8595DB" box="[1143,1294,980,1005]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A83DFC42FA8595DB" box="[1143,1294,980,1005]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
. Metacoxae usually contiguous or narrowly separated, moderately (
|
||
<taxonomicName id="3BC54D02FFFED001AF62FC61FB169226" authority="Ganyme, Syrphetodes" authorityName="Ganyme, Syrphetodes" box="[808,1181,1015,1040]" class="Insecta" family="Ulodidae" genus="Arthopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AF62FC61FC059226" box="[808,910,1015,1040]" italics="true" pageId="8" pageNumber="449">Arthopus</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001AFEEFC6EFB8A9239" box="[932,1025,1016,1039]" italics="true" pageId="8" pageNumber="449">Ganyme</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001A85FFC61FB169226" box="[1045,1181,1015,1040]" italics="true" pageId="8" pageNumber="449">Syrphetodes</emphasis>
|
||
</taxonomicName>
|
||
) to widely separated (
|
||
<taxonomicName id="3BC54D02FFFED001ACEAFB8AFE489203" authority="Exohadrus" authorityName="Exohadrus" box="[160,451,1052,1077]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001ACEAFB8AFEBC9203" box="[160,311,1052,1077]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001AD00FB8AFE489203" box="[330,451,1052,1077]" italics="true" pageId="8" pageNumber="449">Exohadrus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3BC54D02FFFED001AE41FB8AFD0F9203" box="[523,644,1052,1077]" class="Insecta" family="Ulodidae" genus="Pteroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AE41FB8AFD0F9203" box="[523,644,1052,1077]" italics="true" pageId="8" pageNumber="449">Pteroderes</emphasis>
|
||
</taxonomicName>
|
||
) and extending laterally to meet elytra. Metendosternite with arms moderately to very long (
|
||
<figureCitation id="64FE2A04FFFED001AD98FBA9FDDA926E" box="[466,593,1087,1112]" captionStart="FIGURES 7 A – N. A" captionStartId="20.[151,264,1802,1824]" captionTargetBox="[158,1422,199,1773]" captionTargetId="figure@20.[151,1436,193,1780]" captionTargetPageId="20" captionText="FIGURES 7 A – N. A, Pteroderes tuberosus Germain, head dorsal; B, Trachyderastes dipsaconiae Kaszab, maxilla; C, Brouniphylax sp., maxilla; D, Arthopus brouni Sharp, maxilla; E, Pteroderes tuberosus Germain, labium; F, Brouniphylax exiquus (Broun), antenna; G, Ganyme sp., prothorax frontal view. H – J, Metendosternite: H, Pteroderes tuberosus Germain; I, Brouniphylax exiquus (Broun); J, Trachyderas costulatum Fairmaire & Germain. K – L, Ulobostrichus monteithi sp. n., abdomen and detail; M, Pteroderes tuberosus Germain, abdomen; N, Brouniphylax sp., female ovipositor." httpUri="https://zenodo.org/record/258472/files/figure.png" pageId="8" pageNumber="449">Figs 7H–I</figureCitation>
|
||
), laminae present (
|
||
<emphasis id="CEB1EA93FFFED001AF77FBD7FBA6926E" box="[829,1069,1087,1112]" italics="true" pageId="8" pageNumber="449">
|
||
<taxonomicName id="3BC54D02FFFED001AF77FBD7FBA2926E" authority="Trachyderas" authorityName="Trachyderas" box="[829,1065,1087,1112]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">Meryx, Trachyderas</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3BC54D02FFFED001A83EFBA9FA85926E" box="[1140,1294,1087,1112]" class="Insecta" family="Ulodidae" genus="Ulocyphaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CEB1EA93FFFED001A83EFBA9FA85926E" box="[1140,1294,1087,1112]" italics="true" pageId="8" pageNumber="449">Ulocyphaleus</emphasis>
|
||
</taxonomicName>
|
||
) or absent; ventrolateral processes absent or weakly developed; anterior tendons present, poorly developed and moderately to widely separated, or absent. Hind wings present (
|
||
<figureCitation id="64FE2A04FFFED001AE9DFB11FCA39296" box="[727,808,1159,1184]" captionStart="FIGURES 5 A – J. A – D" captionStartId="18.[151,264,1725,1747]" captionTargetBox="[193,1397,210,1689]" captionTargetId="figure@18.[180,1408,193,1703]" captionTargetPageId="18" captionText="FIGURES 5 A – J. A – D, Head and prothorax, ventral view: A, Ulocyphaleus valdivianus (Philippi & Philippi); B, Trachyderas costulatum Fairmaire & Germain; C, Syrphetodes marginatus Pascoe; D, Arthopus brouni Sharp. E, Trachyderastes dipsaconiae Kaszab, antenna detail; F – I, Ulobostrichus monteithi sp. n. F, antenna, G, elytral apex, H, body lateral view, I, body latero-ventral view; J, Meryx rugosa Latreille, wing." httpUri="https://zenodo.org/record/258470/files/figure.png" pageId="8" pageNumber="449">Fig. 5J</figureCitation>
|
||
) or absent; with moderately to very long apical field; radial cell well developed, with inner posterobasal angle right; cross-vein r3 absent; medial field with four free veins, medial fleck present; wedge cell long and narrow; medial and anal embayments absent. Femoral attachment of mid trochanter usually strongly oblique with base of femur abutting coxa. Tibiae simple; two apical spurs distinct. Tarsi 5-5-4 (4-4-
|
||
<quantity id="3B3D9B64FFFED001ADF8FA8EFE549306" box="[434,479,1303,1328]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="449" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="3BC54D02FFFED001ADACFA8FFDA69306" box="[486,557,1305,1328]" class="Insecta" family="Ulodidae" genus="Meryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001ADACFA8FFDA69306" box="[486,557,1305,1328]" italics="true" pageId="8" pageNumber="449">Meryx</emphasis>
|
||
</taxonomicName>
|
||
); tarsomeres (
|
||
<figureCitation id="64FE2A04FFFED001AE85FA81FCC89306" box="[719,835,1303,1328]" captionStart="FIGURES 6 A – O. A – C" captionStartId="19.[151,264,1702,1724]" captionTargetBox="[159,1433,201,1671]" captionTargetId="figure@19.[151,1436,193,1680]" captionTargetPageId="19" captionText="FIGURES 6 A – O. A – C head dorsal: A, Syrphetodes marginatus Pascoe; B, Pteroderes tuberosus Germain; C, Ulocyphaleus valdivianus (Philippi & Philippi). D – G, head lateral: D, Meryx aequalis Blackburn; E, Pteroderes tuberosus Germain; F, Ganyme tuberculata Germain; G, Notocerastes blackburni Carter. H, Ulobostrichus monteithi sp. n., mid leg. I, Arthopus brouni Sharp, mid-claw. J – O, Maxillary palps: J, Ulocyphaleus valdivianus (Philippi & Philippi); K, Syrphetodes marginatus Pascoe; L, Ganyme sapphiro (Newman); M, Trachyderastes dipsaconiae Kaszab; N, Meryx rugosa Latreille, O, Pteroderes tuberosus Germain." httpUri="https://zenodo.org/record/258471/files/figure.png" pageId="8" pageNumber="449">Figs 6H, I</figureCitation>
|
||
) simple and similar in form (protarsomere 2 variable in length), tarsomeres 1–4 usually small and with their combined length equal or subequal to tarsomere 5, combined length of tarsomeres 1–4 significantly greater in length than tarsomere
|
||
<quantity id="3B3D9B64FFFED001A86FFAC9FBDB934E" box="[1061,1104,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="449" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="3BC54D02FFFED001A81CFAC9FA8A934E" box="[1110,1281,1375,1400]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A81CFAC9FA8A934E" box="[1110,1281,1375,1400]" italics="true" pageId="8" pageNumber="449">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
, metatarsus 1 of
|
||
<taxonomicName id="3BC54D02FFFED001ACF2FA12FECF93AB" box="[184,324,1412,1437]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001ACF2FA12FECF93AB" box="[184,324,1412,1437]" italics="true" pageId="8" pageNumber="449">Trachyderas</emphasis>
|
||
</taxonomicName>
|
||
elongate and equal to 4; tarsal pads usually absent or poorly developed; pretarsal claws simple; empodium bisetose, setal insertions often ventral.
|
||
</paragraph>
|
||
<paragraph id="FC7A3681FFFED001AC8DFA5AFC5391A3" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="449">
|
||
Abdomen (
|
||
<figureCitation id="64FE2A04FFFED001AD0DFA5AFE2C93D2" box="[327,423,1484,1509]" captionStart="FIGURES 7 A – N. A" captionStartId="20.[151,264,1802,1824]" captionTargetBox="[158,1422,199,1773]" captionTargetId="figure@20.[151,1436,193,1780]" captionTargetPageId="20" captionText="FIGURES 7 A – N. A, Pteroderes tuberosus Germain, head dorsal; B, Trachyderastes dipsaconiae Kaszab, maxilla; C, Brouniphylax sp., maxilla; D, Arthopus brouni Sharp, maxilla; E, Pteroderes tuberosus Germain, labium; F, Brouniphylax exiquus (Broun), antenna; G, Ganyme sp., prothorax frontal view. H – J, Metendosternite: H, Pteroderes tuberosus Germain; I, Brouniphylax exiquus (Broun); J, Trachyderas costulatum Fairmaire & Germain. K – L, Ulobostrichus monteithi sp. n., abdomen and detail; M, Pteroderes tuberosus Germain, abdomen; N, Brouniphylax sp., female ovipositor." httpUri="https://zenodo.org/record/258472/files/figure.png" pageId="8" pageNumber="449">Figs 7K</figureCitation>
|
||
,
|
||
<figureCitation id="64FE2A04FFFED001ADFFFA5AFE6C93D2" box="[437,487,1484,1508]" captionStart="FIGURES 10 A – D" captionStartId="23.[151,264,1767,1789]" captionTargetBox="[151,1436,193,1745]" captionTargetId="figure@23.[151,1436,193,1746]" captionTargetPageId="23" captionText="FIGURES 10 A – D. Ulodes hispidus Pascoe: A, pronotum; B, prothorax ventral; C, pterothorax and abdomen; D, abdomen." httpUri="https://zenodo.org/record/258475/files/figure.png" pageId="8" pageNumber="449">10D</figureCitation>
|
||
) with five ventrites of relatively equal lengths, with ventrite 1 equal to or slightly shorter than ventrite 2, surface flattened to weakly convex in lateral view, postcoxal lines absent and intercoxal process usually acute or subacute, broadly truncate in
|
||
<taxonomicName id="3BC54D02FFFED001AEBEF982FC00901B" box="[756,907,1556,1581]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AEBEF982FC00901B" box="[756,907,1556,1581]" italics="true" pageId="8" pageNumber="449">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
; ventrites 1 and 2 connate; membranes 3 and 4 externally visible in
|
||
<emphasis id="CEB1EA93FFFED001ADC4F9A1FE759066" box="[398,510,1591,1616]" italics="true" pageId="8" pageNumber="449">Arthropus</emphasis>
|
||
,
|
||
<taxonomicName id="3BC54D02FFFED001AE59F9A1FB149066" authority="Dipsaconia, Meryx, Notocerastes, Phaennis" authorityName="Dipsaconia, Meryx, Notocerastes, Phaennis" box="[531,1183,1591,1616]" class="Insecta" family="Ulodidae" genus="Exohadrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AE59F9A1FD079066" box="[531,652,1591,1616]" italics="true" pageId="8" pageNumber="449">Exohadrus</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001AEEAF9AFFCAA9066" box="[672,801,1593,1616]" italics="true" pageId="8" pageNumber="449">Dipsaconia</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001AF7FF9AFFCF79066" box="[821,892,1593,1616]" italics="true" pageId="8" pageNumber="449">Meryx</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001AFDBF9AFFBA89066" box="[913,1059,1593,1616]" italics="true" pageId="8" pageNumber="449">Notocerastes</emphasis>
|
||
,
|
||
<emphasis id="CEB1EA93FFFED001A87DF9A1FB149066" box="[1079,1183,1591,1616]" italics="true" pageId="8" pageNumber="449">Phaennis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3BC54D02FFFED001A8A3F9A1FA1F9066" box="[1257,1428,1591,1616]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001A8A3F9A1FA1F9066" box="[1257,1428,1591,1616]" italics="true" pageId="8" pageNumber="449">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
; glandular pores present in
|
||
<taxonomicName id="3BC54D02FFFED001AD8EF9CAFDC69043" box="[452,589,1628,1653]" class="Insecta" family="Tenebrionidae" genus="Syrphetodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AD8EF9CAFDC69043" box="[452,589,1628,1653]" italics="true" pageId="8" pageNumber="449">Syrphetodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BC54D02FFFED001AECFF9CAFCBB9043" box="[645,816,1628,1653]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AECFF9CAFCBB9043" box="[645,816,1628,1653]" italics="true" pageId="8" pageNumber="449">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
; lateral flanges present on all or some of the ventrites, well-developed or reduced (some
|
||
<taxonomicName id="3BC54D02FFFED001AE6EF916FD0A90A1" box="[548,641,1664,1687]" class="Insecta" family="Tenebrionidae" genus="Ganyme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AE6EF916FD0A90A1" box="[548,641,1664,1687]" italics="true" pageId="8" pageNumber="449">Ganyme</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<taxonomicName id="3BC54D02FFFED001AE93F9E9FCE990AE" box="[729,866,1663,1688]" class="Insecta" family="Tenebrionidae" genus="Syrphetodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AE93F9E9FCE990AE" box="[729,866,1663,1688]" italics="true" pageId="8" pageNumber="449">Syrphetodes</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3BC54D02FFFED001AFEDF9E9FBD990AE" box="[935,1106,1663,1688]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AFEDF9E9FBD990AE" box="[935,1106,1663,1688]" italics="true" pageId="8" pageNumber="449">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
); ventrite 5 with or without apical grooves or impressions. Male with spiculum gastrale on sternite IX present; tergite IX completely fused to tergite X. Aedeagus tenebrionoid (
|
||
<figureCitation id="64FE2A04FFFED001AE55F951FD5890D6" box="[543,723,1735,1760]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="8" pageNumber="449">Figs 8F–G, J–L</figureCitation>
|
||
), symmetrical, not inverted; parameres partially fused together and articulated to phallobase, forming apicale, which lacks accessory lobes; anterior edge of penis without struts, apex acute to subacute (rounded in one dissected specimen of
|
||
<taxonomicName id="3BC54D02FFFED001AF17F899FC4F911E" box="[861,964,1807,1832]" class="Insecta" family="Ulodidae" genus="Arthopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFED001AF17F899FC4F911E" box="[861,964,1807,1832]" italics="true" pageId="8" pageNumber="449">Arthopus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="64FE2A04FFFED001AF9EF899FBC2911E" box="[980,1097,1807,1832]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="8" pageNumber="449">Figs 8H, I</figureCitation>
|
||
). Sternite VIII in female with long spiculum ventrale. Ovipositor (
|
||
<figureCitation id="64FE2A04FFFED001AE66F8A2FD0F917A" box="[556,644,1844,1869]" captionStart="FIGURES 7 A – N. A" captionStartId="20.[151,264,1802,1824]" captionTargetBox="[158,1422,199,1773]" captionTargetId="figure@20.[151,1436,193,1780]" captionTargetPageId="20" captionText="FIGURES 7 A – N. A, Pteroderes tuberosus Germain, head dorsal; B, Trachyderastes dipsaconiae Kaszab, maxilla; C, Brouniphylax sp., maxilla; D, Arthopus brouni Sharp, maxilla; E, Pteroderes tuberosus Germain, labium; F, Brouniphylax exiquus (Broun), antenna; G, Ganyme sp., prothorax frontal view. H – J, Metendosternite: H, Pteroderes tuberosus Germain; I, Brouniphylax exiquus (Broun); J, Trachyderas costulatum Fairmaire & Germain. K – L, Ulobostrichus monteithi sp. n., abdomen and detail; M, Pteroderes tuberosus Germain, abdomen; N, Brouniphylax sp., female ovipositor." httpUri="https://zenodo.org/record/258472/files/figure.png" pageId="8" pageNumber="449">Fig. 7N</figureCitation>
|
||
) long and slender, bursa typically saccular with a narrow spermathecal gland (
|
||
<figureCitation id="64FE2A04FFFED001ACA1F8C1FEC39146" box="[235,328,1879,1904]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="8" pageNumber="449">Fig. 8E</figureCitation>
|
||
); paraprocts distinctly longer than coxites, lightly sclerotized except for baculi, with coxites subdivided, apically slender, with well developed apical or subapical styli.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4DF650AFFFED000AC8DF836FDF6972A" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="449" type="discussion">
|
||
<paragraph id="FC7A3681FFFED000AC8DF836FDF6972A" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,285]" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="449">
|
||
<emphasis id="CEB1EA93FFFED001AC8DF836FEC4918F" bold="true" box="[199,335,1952,1977]" pageId="8" pageNumber="449">Comments.</emphasis>
|
||
The family description above is modified from that in
|
||
<bibRefCitation id="98544B70FFFED001AF85F837FB6F918F" box="[975,1252,1952,1977]" pageId="8" pageNumber="449" refString="Lawrence, J. F., Hastings, A. M., Dallwitz, M. J., Paine, T. A. & Zurcher, E. J. (1999 b) " Beetles of the World: A Key and Information System for Families and Subfamilies. " CD-ROM, Version 1.0 for MS-Windows. CSIRO Publishing, Melbourne." type="book">
|
||
Lawrence
|
||
<emphasis id="CEB1EA93FFFED001A80DF837FB09918F" box="[1095,1154,1952,1977]" italics="true" pageId="8" pageNumber="449">et al.</emphasis>
|
||
(1999b)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="98544B70FFFED001A955F836FEC591EA" author="Leschen" pageId="8" pageNumber="449" refString="Leschen, R. A. B. & Slipinski, S. A. (2010) Chapter 11.10. Ulodidae Pascoe, 1869. In: Leschen, R. A. B., Beutel, R. G. & Lawrence, J. F. (Eds.), Handbook of Zoology, Coleoptera Volume 2: Morphology and Systematics (Elateroidea, Bostrichformia, Cucujiformia partim). Walter de Gruyter, Berlin, pp. 559 - 563." type="journal article" year="2010">Leschen & Ślipiński (2010)</bibRefCitation>
|
||
based on the addition of genera described in this study and recent work by
|
||
<bibRefCitation id="98544B70FFFED000A8F0F852FF6F9686" author="Leschen" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="449" refString="Leschen, R. A. B. & Buckley, R. T. (2015) Revision and phylogeny of Syrphetodes (Coleoptera: Ulodidae): implications for biogeography, alpinization and conservation. Systematic Entomology, 40, 143 - 168. http: // dx. doi. org / 10.1111 / syen. 12094" type="journal article" year="2015">Leschen & Buckley (2015)</bibRefCitation>
|
||
. Larval characters and descriptions can be found in
|
||
<bibRefCitation id="98544B70FFFFD000AF0CFF01FBFE9686" author="Beutel" box="[838,1141,151,176]" pageId="9" pageNumber="450" refString="Beutel, R. G. & Friedrich, F. (2005) Comparative study of larvae of Tenebrionoidea (Coleoptera: Cucujiformia). European Journal of Entomology, 102 (2), 241 - 264. http: // dx. doi. org / 10.14411 / eje. 2005.037" type="journal article" year="2005">Beutel & Friedrich (2005)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="98544B70FFFFD000A8FBFF01FF6E96E2" author="Leschen" pageId="9" pageNumber="450" refString="Leschen, R. A. B. & Slipinski, S. A. (2010) Chapter 11.10. Ulodidae Pascoe, 1869. In: Leschen, R. A. B., Beutel, R. G. & Lawrence, J. F. (Eds.), Handbook of Zoology, Coleoptera Volume 2: Morphology and Systematics (Elateroidea, Bostrichformia, Cucujiformia partim). Walter de Gruyter, Berlin, pp. 559 - 563." type="journal article" year="2010">Leschen & Ślipiński (2010)</bibRefCitation>
|
||
. There are currently 16 genera and 41 species and further taxonomic work is needed to describe species in
|
||
<collectingCountry id="84D27611FFFFD000ACDDFF49FF7596CE" box="[151,254,223,248]" name="Australia" pageId="9" pageNumber="450">Australia</collectingCountry>
|
||
(
|
||
<taxonomicName id="3BC54D02FFFFD000AD44FF77FE0496CE" box="[270,399,225,248]" class="Insecta" family="Tenebrionidae" genus="Dipsaconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFFD000AD44FF77FE0496CE" box="[270,399,225,248]" italics="true" pageId="9" pageNumber="450">Dipsaconia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="EAC3FD08FFFFD000ADD4FF76FE4296C1" box="[414,457,224,248]" pageId="9" pageNumber="450" type="generic">2 n.</specimenCount>
|
||
spp.;
|
||
<taxonomicName id="3BC54D02FFFFD000AE44FF76FDE096C1" box="[526,619,224,247]" class="Insecta" family="Tenebrionidae" genus="Ganyme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFFD000AE44FF76FDE096C1" box="[526,619,224,247]" italics="true" pageId="9" pageNumber="450">Ganyme</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="EAC3FD08FFFFD000AE31FF76FD2E96C1" box="[635,677,224,248]" pageId="9" pageNumber="450" type="generic">2 n.</specimenCount>
|
||
spp.;
|
||
<taxonomicName id="3BC54D02FFFFD000AEA1FF49FCB196CE" box="[747,826,223,248]" class="Insecta" family="Tenebrionidae" genus="Ulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFFD000AEA1FF49FCB196CE" box="[747,826,223,248]" italics="true" pageId="9" pageNumber="450">Ulodes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="EAC3FD08FFFFD000AF03FF76FCFF96C1" box="[841,884,224,248]" pageId="9" pageNumber="450" type="generic">1 n.</specimenCount>
|
||
sp.),
|
||
<collectingCountry id="84D27611FFFFD000AFF9FF76FBC196CE" box="[947,1098,223,248]" name="New Zealand" pageId="9" pageNumber="450">New Zealand</collectingCountry>
|
||
(
|
||
<taxonomicName id="3BC54D02FFFFD000A816FF49FB7896CE" box="[1116,1267,223,248]" class="Insecta" family="Ulodidae" genus="Brouniphylax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFFD000A816FF49FB7896CE" box="[1116,1267,223,248]" italics="true" pageId="9" pageNumber="450">Brouniphylax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="EAC3FD08FFFFD000A948FF76FAA696C1" box="[1282,1325,224,248]" pageId="9" pageNumber="450" type="generic">3 n.</specimenCount>
|
||
spp.) and
|
||
<collectingCountry id="84D27611FFFFD000ACDDFE93FECE972B" box="[151,325,260,285]" name="New Caledonia" pageId="9" pageNumber="450">New Caledonia</collectingCountry>
|
||
(
|
||
<taxonomicName id="3BC54D02FFFFD000AD1EFE92FE74972B" box="[340,511,260,285]" class="Insecta" family="Ulodidae" genus="Trachyderastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEB1EA93FFFFD000AD1EFE92FE74972B" box="[340,511,260,285]" italics="true" pageId="9" pageNumber="450">Trachyderastes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="EAC3FD08FFFFD000AE47FE92FDBC972A" box="[525,567,260,284]" pageId="9" pageNumber="450" type="generic">4 n.</specimenCount>
|
||
spp.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |