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<document ID-DOI="10.11865/zs.2022101" ID-ISSN="2095-6827" ID-Zenodo-Dep="7176146" ID-ZooBank="1C62670D-1CEB-4A0C-8A12-D8824F6B61DA" approvalRequired="14" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="6" checkinTime="1665296205732" checkinUser="felipe" docAuthor="Xin, Yafei, Jiang, Tongyao, Yao, Zhiyuan &amp; Li, Shuqiang" docDate="2022" docId="743FB854091EBC5CB1F1CE4AFDFFF837" docLanguage="en" docName="ZoolSyst.47.1.1–65.pdf" docOrigin="Zoological Systematics 47 (1)" docSource="http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Burmatheridon cetani Jiang &amp; Li 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="58" masterDocId="8806C02C0926BC65B179CF72FFCEFFBD" masterDocTitle="Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="57" updateTime="1665315257893" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xin, Yafei</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar Corresponding authors, E-mails: yaozy @ synu. edu. cn; lisq @ ioz. ac. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Tongyao</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yao, Zhiyuan</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>65</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11865/zs.2022101</mods:identifier>
<mods:identifier type="ISSN">2095-6827</mods:identifier>
<mods:identifier type="Zenodo-Dep">7176146</mods:identifier>
<mods:identifier type="ZooBank">1C62670D-1CEB-4A0C-8A12-D8824F6B61DA</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172618" ID-Zenodo-Dep="7172618" LSID="urn:lsid:plazi:treatment:743FB854091EBC5CB1F1CE4AFDFFF837" httpUri="http://treatment.plazi.org/id/743FB854091EBC5CB1F1CE4AFDFFF837" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<subSubSection box="[136,768,312,337]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph blockId="56.[136,768,312,337]" box="[136,768,312,337]" pageId="56" pageNumber="57">
<heading box="[136,768,312,337]" centered="true" fontSize="10" level="2" pageId="56" pageNumber="57" reason="2">
<emphasis bold="true" box="[136,608,312,337]" pageId="56" pageNumber="57">
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[136,511,312,337]" class="Arachnida" family="Tetrablemmidae" genus="Burmatheridon" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cetani" status="sp. nov.">
<emphasis bold="true" box="[136,380,312,337]" italics="true" pageId="56" pageNumber="57">Burmatheridon cetani</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel box="[523,608,313,337]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[638,761,313,337]" captionStart-0="Figure 52" captionStart-1="Figure 53" captionStart-2="Figure 54" captionStartId-0="54.[155,218,2055,2077]" captionStartId-1="55.[155,218,2025,2047]" captionStartId-2="56.[155,218,2030,2052]" captionTargetBox-0="[189,1399,241,2020]" captionTargetBox-1="[189,1398,378,1988]" captionTargetBox-2="[264,1323,432,1988]" captionTargetId-0="figure-54@54.[189,1399,241,2020]" captionTargetId-1="figure-42@55.[189,1398,378,1988]" captionTargetId-2="figure-54@56.[264,1323,432,1988]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionTargetPageId-2="56" captionText-0="Figure 52. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo). A–B. Right palp (A. Retrolateral view; B. Ventral view); C–D. Left palp (C. Ventral view; D. Retrolateral view). Scale bars=0.05mm." captionText-1="Figure 53. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." captionText-2="Figure 54. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo), left palp, ventral view. Scale bar = 0.05mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176264" figureDoi-1="http://doi.org/10.5281/zenodo.7176266" figureDoi-2="http://doi.org/10.5281/zenodo.7176268" httpUri-0="https://zenodo.org/record/7176264/files/figure.png" httpUri-1="https://zenodo.org/record/7176266/files/figure.png" httpUri-2="https://zenodo.org/record/7176268/files/figure.png" pageId="56" pageNumber="57">Figs 52–54</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176268" ID-Zenodo-Dep="7176268" httpUri="https://zenodo.org/record/7176268/files/figure.png" pageId="56" pageNumber="57" startId="56.[155,218,2030,2052]" targetBox="[264,1323,432,1988]" targetPageId="56">
<paragraph blockId="56.[155,1433,2030,2084]" pageId="56" pageNumber="57">
Figure 54.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[268,606,2030,2052]" class="Arachnida" family="Tetrablemmidae" genus="Burmatheridon" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cetani" status="sp. nov.">
<emphasis box="[268,483,2030,2052]" italics="true" pageId="56" pageNumber="57">Burmatheridon cetani</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[619,699,2030,2051]" pageId="56" pageNumber="57">
<taxonomicNameLabel box="[619,699,2030,2051]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42702Fo), left palp, ventral view. Scale bar = 0.05 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176270" ID-Zenodo-Dep="7176270" httpUri="https://zenodo.org/record/7176270/files/figure.png" pageId="57" pageNumber="58" startId="57.[155,218,1202,1224]" targetBox="[189,1399,270,1159]" targetPageId="57">
<paragraph blockId="57.[155,1432,1202,1256]" pageId="57" pageNumber="58">
Figure 55.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[260,600,1202,1224]" class="Arachnida" family="Theridiidae" genus="Cretotheridion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="champoi" status="sp. nov.">
<emphasis box="[260,490,1202,1223]" italics="true" pageId="57" pageNumber="58">Cretotheridion champoi</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[609,685,1203,1224]" pageId="57" pageNumber="58">
<taxonomicNameLabel box="[609,685,1203,1224]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42703Fo). A. Left palp, retrolateral view; B. Right palp, retrolateral view. Scale bars =0.05 mm.
</paragraph>
</caption>
<subSubSection pageId="57" pageNumber="58" type="materials_examined">
<paragraph blockId="57.[136,1452,1316,1930]" pageId="57" pageNumber="58">
<materialsCitation county="Hukawng Valley" location="Male" municipality="Late Cretaceous" pageId="57" pageNumber="58" specimenCount="1" typeStatus="holotype">
<typeStatus box="[189,292,1316,1340]" pageId="57" pageNumber="58">Holotype</typeStatus>
.
<location LSID="urn:lsid:plazi:treatment:743FB854091EBC5CB1F1CE4AFDFFF837:F9495F99091FBC5CB04DCA56FEA5FA81" box="[308,363,1316,1340]" county="Hukawng Valley" municipality="Late Cretaceous" name="Male" pageId="57" pageNumber="58">Male</location>
(IZCAS-Ar42702Fo).
<collectingMunicipality box="[622,801,1317,1341]" pageId="57" pageNumber="58">Late Cretaceous</collectingMunicipality>
amber from
<collectingCounty box="[957,1143,1316,1340]" pageId="57" pageNumber="58">Hukawng Valley</collectingCounty>
. Syninclusion includes one spiderling.
</materialsCitation>
</paragraph>
<paragraph blockId="57.[136,1452,1316,1930]" box="[189,1300,1386,1410]" pageId="57" pageNumber="58">Etymology. The species name is a popular Burmese boy’s name and means “hawk”; noun in apposition.</paragraph>
<paragraph blockId="57.[136,1452,1316,1930]" pageId="57" pageNumber="58">
Diagnosis. The new species can be distinguished from
<taxonomicName authorityName="Wunderlich" authorityYear="2018" box="[781,923,1421,1444]" class="Arachnida" family="Tetrablemmidae" genus="Burmatheridon" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="sinespinae">
<emphasis box="[781,923,1421,1444]" italics="true" pageId="57" pageNumber="58">B. sinespinae</emphasis>
</taxonomicName>
(see Wunderlich &amp; Müller, 2018: 117, figs 219– 222, photos 54–55) by the shape of the apophysis on the palp.
<taxonomicName authorityName="Wunderlich" authorityYear="2018" box="[789,929,1456,1479]" class="Arachnida" family="Tetrablemmidae" genus="Burmatheridon" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="sinespinae">
<emphasis box="[789,929,1456,1479]" italics="true" pageId="57" pageNumber="58">B. sinespinae</emphasis>
</taxonomicName>
has four apophyses (a serrated tegular apophysis, two thin tegular apophyses, and a long apophysis) that are different from those of
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[1014,1233,1490,1514]" class="Arachnida" family="Tetrablemmidae" genus="Burmatheridon" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="cetani" status="sp. nov.">
<emphasis box="[1014,1109,1490,1514]" italics="true" pageId="57" pageNumber="58">B. cetani</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[1245,1330,1490,1514]" pageId="57" pageNumber="58">
<taxonomicNameLabel box="[1245,1330,1490,1514]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has three apophyses (embolus, median apophysis, and conductor).
</paragraph>
<paragraph blockId="57.[136,1452,1316,1930]" pageId="57" pageNumber="58">
Description. Male. Total length 1.32; carapace 0.59 long, 0.44 wide; opisthosoma 0.73 long, 0.68 wide. Left palp: - (0.51 + 0.23 + -), left leg I: 4.04 (1.24 + 0.23 + 1.10 + 1.18 + 0.29), leg
<collectionCode box="[900,921,1594,1617]" pageId="57" pageNumber="58">II</collectionCode>
: 2.50 (0.75 + 0.18 + 0.59 + 0.64 + 0.34), leg
<collectionCode box="[1402,1433,1594,1617]" pageId="57" pageNumber="58">III</collectionCode>
: - (0.51 + 0.12 + - + - + -), leg IV: 2.18 (0.69 + 0.11 + 0.52 + 0.55 + 0.31); right palp: - (0.49 + 0.20 + -), right leg I: 3.57 (1.07 + 0.19 + 1.12 + 0.82 + 0.37), leg
<collectionCode box="[488,510,1664,1687]" pageId="57" pageNumber="58">II</collectionCode>
: 2.31 (0.78 + 0.22 + 0.60 + 0.73 + 0.35), leg
<collectionCode box="[986,1016,1664,1687]" pageId="57" pageNumber="58">III</collectionCode>
: - (- + 0.16 + 0.37 + 0.26 + 0.30), leg IV: - (0.63 + 0.28 + 0.56 + - + -). Eye sizes and interdistances: ALE 0.05, AME 0.04, PLE 0.05, PME 0.05; AME–AME 0.09, AME–ALE 0.06, PME–PLE 0.07, PME–PME 0.06. Habitus as in
<figureCitation box="[858,993,1732,1756]" captionStart="Figure 53" captionStartId="55.[155,218,2025,2047]" captionTargetBox="[189,1398,378,1988]" captionTargetId="figure-42@55.[189,1398,378,1988]" captionTargetPageId="55" captionText="Figure 53. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." figureDoi="http://doi.org/10.5281/zenodo.7176266" httpUri="https://zenodo.org/record/7176266/files/figure.png" pageId="57" pageNumber="58">Figs 53A–B</figureCitation>
. Carapace almost oval, yellow, short, with brown setae. Sternum yellowish, round, with dense setae. Labium not visible. Legs have two rows of strong setae on metatarsus and tarsus, 3 claws. Opisthosoma oval, yellowish.
</paragraph>
<paragraph blockId="57.[136,1452,1316,1930]" pageId="57" pageNumber="58">
Palp (
<figureCitation box="[252,388,1836,1860]" captionStart="Figure 52" captionStartId="54.[155,218,2055,2077]" captionTargetBox="[189,1399,241,2020]" captionTargetId="figure-54@54.[189,1399,241,2020]" captionTargetPageId="54" captionText="Figure 52. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo). A–B. Right palp (A. Retrolateral view; B. Ventral view); C–D. Left palp (C. Ventral view; D. Retrolateral view). Scale bars=0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176264" httpUri="https://zenodo.org/record/7176264/files/figure.png" pageId="57" pageNumber="58">Figs 52A–D</figureCitation>
,
<figureCitation box="[400,430,1836,1860]" captionStart="Figure 54" captionStartId="56.[155,218,2030,2052]" captionTargetBox="[264,1323,432,1988]" captionTargetId="figure-54@56.[264,1323,432,1988]" captionTargetPageId="56" captionText="Figure 54. Burmatheridon cetani Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42702Fo), left palp, ventral view. Scale bar = 0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176268" httpUri="https://zenodo.org/record/7176268/files/figure.png" pageId="57" pageNumber="58">54</figureCitation>
). Femur longer than patella, patella and tibia without apophyses. Embolus slender, sclerotized. Base of embolus irregular polygon shaped, serrated at edge, median apophysis originates from tegulum, sclerotized, slender, conductor deformed, subtegulum small.
</paragraph>
</subSubSection>
</treatment>
</document>
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