174 lines
20 KiB
XML
174 lines
20 KiB
XML
<document id="C90CEAB7694524D04B081D585156989F" ID-DOI="10.1206/664.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="10108959" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630223030481" checkinUser="carolina" docAuthor="Platnick, Norman I. & Dupérré, Nadine" docDate="2009" docId="736787C1D944FFA4FF42FF6515C0F93C" docLanguage="en" docName="N3672.pdf" docOrigin="American Museum Novitates 3649" docSource="http://www.bioone.org/doi/abs/10.1206/664.1" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Heteroonops andros Platnick & Dupérré 2009, new species" docType="treatment" docVersion="4" lastPageNumber="38" masterDocId="8F5EFFB9D961FF81FFE7FFB61655FFE5" masterDocTitle="The Goblin Spider Genus Heteroonops (Araneae, Oonopidae), With Notes on Oonops" masterLastPageNumber="72" masterPageNumber="1" pageNumber="38" updateTime="1700018162531" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AD324573FEBA2A51D921EE6975E60859">The Goblin Spider Genus Heteroonops (Araneae, Oonopidae), With Notes on Oonops</mods:title>
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<mods:namePart id="7F9D42762844622C314D7FEFC27A9E9B">Platnick, Norman I.</mods:namePart>
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<mods:namePart id="A455BAB2EF2DC85684A6DB76E433F292">Dupérré, Nadine</mods:namePart>
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<mods:typeOfResource id="2D308142CF9B30F6A3EFDB06CAB1BD33">text</mods:typeOfResource>
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<mods:title id="A226A8D45B24D2E3537553FD58A94022">American Museum Novitates</mods:title>
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<mods:part id="4A76AA9602DD3CC4F789D78079DA3AE6">
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<mods:date id="84F7E2BE2DBD6D70E1BE0AB2CA3D1A52">2009</mods:date>
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<mods:number id="CA2DD5AE2CBC630F802161427ECCD95C">2009-11-30</mods:number>
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<mods:detail id="B1A1C5AE26317F4AD0CFFCCB41769EA8" type="volume">
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<mods:number id="F53A69B3B6E98CFA25A9CE8548B33448">3649</mods:number>
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<mods:url id="28472BD8B655098032C0F90A206C409F">http://www.bioone.org/doi/abs/10.1206/664.1</mods:url>
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<mods:classification id="E57E9D18851F680FB884B61B12DA5097">journal article</mods:classification>
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<mods:identifier id="03D7FBCE8F45EBB5EB2913E7397DA1BC" type="DOI">10.1206/664.1</mods:identifier>
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<mods:identifier id="3BDD14B79702E4EA76467E3028B9EA81" type="ISSN">0003-0082</mods:identifier>
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<treatment id="736787C1D944FFA4FF42FF6515C0F93C" LSID="urn:lsid:plazi:treatment:736787C1D944FFA4FF42FF6515C0F93C" httpUri="http://treatment.plazi.org/id/736787C1D944FFA4FF42FF6515C0F93C" lastPageNumber="38" pageId="37" pageNumber="38">
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<subSubSection id="B3D4655CD944FFA4FF42FF6517E9FEFE" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="FB7136D7D944FFA4FF42FF65144DFF0E" blockId="37.[165,536,211,235]" box="[165,536,211,235]" pageId="37" pageNumber="38">
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<heading id="A03981BBD944FFA4FF42FF65144DFF0E" box="[165,536,211,235]" centered="true" fontSize="36" level="2" pageId="37" pageNumber="38" reason="8">
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<taxonomicName id="3CCE4D54D944FFA4FF42FF6517D0FF0E" ID-CoL="3LBLG" authority="Platnick & Dupérré, 2009" authorityName="Platnick & Dupérré" authorityYear="2009" box="[165,389,211,235]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="andros" status="sp. nov.">
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<emphasis id="C9BAEAC5D944FFA4FF42FF6517D0FF0E" bold="true" box="[165,389,211,235]" italics="true" pageId="37" pageNumber="38">Heteroonops andros</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="D28957BED944FFA4FE72FF65144DFF0E" box="[405,536,211,235]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="FB7136D7D944FFA4FEE6FEB517E9FEFE" blockId="37.[257,444,259,283]" box="[257,444,259,283]" pageId="37" pageNumber="38">
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<heading id="A03981BBD944FFA4FEE6FEB517E9FEFE" box="[257,444,259,283]" centered="true" fontSize="36" level="3" pageId="37" pageNumber="38" reason="7">
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<figureCitation id="63F52A52D944FFA4FEE6FEB517E9FEFE" box="[257,444,259,283]" captionStart="Figs" captionStartId="38.[151,198,1673,1695]" captionTargetBox="[216,1138,210,1639]" captionTargetId="figure-1@38.[216,1139,210,1639]" captionTargetPageId="38" captionText="Figs. 169–175. Heteroonops andros, new species, female. 169. Carapace, dorsal view. 170. Habitus, ventral view. 171. Same, lateral view. 172. Carapace and palp, anterior view. 173. Epigastric region, ventral view. 174. Genitalia, ventral view. 175. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10128204" httpUri="https://zenodo.org/record/10128204/files/figure.png" pageId="37" pageNumber="38">Figures 169–175</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3D4655CD944FFA4FF95FE881760FE2E" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="FB7136D7D944FFA4FF95FE881760FE2E" blockId="37.[88,614,318,1163]" pageId="37" pageNumber="38">
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<materialsCitation id="4BA63C8AD944FFA4FF95FE881764FE2E" accessionNumber="OON 21177" collectingDate="1994-05-01" collectingDateMax="1994-05-06" collectingDateMin="1994-05-01" collectionCode="AMNH" collectorName="R. Anderson" county="Bahama Islands" location="Female" municipality="Andros Island" pageId="37" pageNumber="38" specimenCount="1" typeStatus="holotype">
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TYPE:
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<location id="FE11600CD944FFA4FF2BFE88174AFEB3" LSID="urn:lsid:plazi:treatment:736787C1D944FFA4FF42FF6515C0F93C:FE11600CD944FFA4FF2BFE88174AFEB3" box="[204,287,318,342]" county="Bahama Islands" municipality="Andros Island" name="Female" pageId="37" pageNumber="38">Female</location>
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<typeStatus id="24758875D944FFA4FECEFE8817D8FEB3" box="[297,397,318,342]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
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taken in a
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<location id="FE11600CD944FFA4FDF2FE881430FEB3" LSID="urn:lsid:plazi:treatment:736787C1D944FFA4FF42FF6515C0F93C:FE11600CD944FFA4FDF2FE881430FEB3" box="[533,613,318,342]" county="Bahama Islands" municipality="Andros Island" name="Berlese" pageId="37" pageNumber="38">Berlese</location>
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sample of coastal coppice litter at
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<location id="FE11600CD944FFA4FE3BFEED1434FE96" LSID="urn:lsid:plazi:treatment:736787C1D944FFA4FF42FF6515C0F93C:FE11600CD944FFA4FE3BFEED1434FE96" box="[476,609,347,371]" county="Bahama Islands" municipality="Andros Island" name="Pigeon Cay" pageId="37" pageNumber="38">Pigeon Cay</location>
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,
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<collectingMunicipality id="1B15ACADD944FFA4FFBFFECE1753FE75" box="[88,262,376,400]" pageId="37" pageNumber="38">Andros Island</collectingMunicipality>
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,
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<collectingCounty id="12104E5BD944FFA4FEFDFECE178CFE75" box="[282,473,376,400]" pageId="37" pageNumber="38">Bahama Islands</collectingCounty>
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(
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<date id="8F701017D944FFA4FE16FECE16C5FE48" pageId="37" pageNumber="38" value="1994-05-01" valueMax="1994-05-06" valueMin="1994-05-01">
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<collectingDate id="9F34E9FFD944FFA4FE16FECE16C5FE48" pageId="37" pageNumber="38" value="1994-05-01" valueMax="1994-05-06" valueMin="1994-05-01">May 1–6, 1994</collectingDate>
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</date>
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;
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<collectorName id="563B5301D944FFA4FF41FE231713FE48" box="[166,326,405,429]" pageId="37" pageNumber="38">R. Anderson</collectorName>
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), deposited in
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<collectionCode id="9DDFAE12D944FFA4FDECFE231430FE48" box="[523,613,405,429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="37" pageNumber="38">AMNH</collectionCode>
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(PBI_OON 21177)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3D4655CD944FFA4FF95FE661414FDE3" pageId="37" pageNumber="38" type="etymology">
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<paragraph id="FB7136D7D944FFA4FF95FE661414FDE3" blockId="37.[88,614,318,1163]" pageId="37" pageNumber="38">
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ETYMOLOGY: The specific name is a noun in apposition taken from the
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<typeStatus id="24758875D944FFA4FE4BFE581789FDE3" box="[428,476,494,518]" pageId="37" pageNumber="38">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3D4655CD944FFA4FF95FDBD17ABFDBB" pageId="37" pageNumber="38" type="diagnosis">
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<paragraph id="FB7136D7D944FFA4FF95FDBD17ABFDBB" blockId="37.[88,614,318,1163]" pageId="37" pageNumber="38">DIAGNOSIS: Females can be recognized by the triangular anterior receptaculum with a wide, translucent sac (figs. 173–175).</paragraph>
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</subSubSection>
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<subSubSection id="B3D4655CD944FFA4FF95FDD21449FB88" pageId="37" pageNumber="38" type="description">
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<paragraph id="FB7136D7D944FFA4FF95FDD21705FD99" blockId="37.[88,614,318,1163]" box="[114,336,612,636]" pageId="37" pageNumber="38">MALE: Unknown.</paragraph>
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<paragraph id="FB7136D7D944FFA4FF95FD371753FBB5" blockId="37.[88,614,318,1163]" pageId="37" pageNumber="38">FEMALE (PBI_OON 21177, figs. 169– 172): Total length 1.80. Palpal patella prolaterally dilated at half its length. Abdominal venter without pair of subcuticular dark spots just anterior of spinnerets. Leg spination (leg II missing): femora: I d1-0-0; III, IV d1-0-1; tibiae: I v0-2-2; III p1-1-0, v0-2-2, r0-1-0; IV d0-1-0, p1-0-1, v0-1-2, r1-0-1; metatarsi: I v2- 0-2; III d1-0-0, v0-2-2, r1-0-1; IV d1-0-0, p1-0- 1, v2-1-2, r1-0-1. Palpal spination: femur v1-1- 1, patella p0-1-1, tibia p1-1-2, v1-1-2, tarsus p1-3-1, v2-1-2. Anterior receptaculum consisting of sclerotized triangle leading to membranous sac; posterior receptaculum with presumed secretory glands arrayed in rows (figs. 173–175).</paragraph>
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<paragraph id="FB7136D7D944FFA4FF95FBE31449FB88" blockId="37.[88,614,318,1163]" box="[114,540,1109,1133]" pageId="37" pageNumber="38">OTHER MATERIAL EXAMINED: None.</paragraph>
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</subSubSection>
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<subSubSection id="B3D4655CD944FFA4FF95FBC515C0F93C" pageId="37" pageNumber="38" type="distribution">
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<paragraph id="FB7136D7D944FFA4FF95FBC517BFFB6E" blockId="37.[88,614,318,1163]" box="[114,490,1139,1163]" pageId="37" pageNumber="38">DISTRIBUTION: Bahama Islands.</paragraph>
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<paragraph id="FB7136D7D944FFA4FF16FB051798FB2E" blockId="37.[241,461,1203,1227]" box="[241,461,1203,1227]" pageId="37" pageNumber="38">The Cuban Species</paragraph>
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<paragraph id="FB7136D7D944FFA4FF95FB581761F93F" blockId="37.[88,614,1262,1754]" pageId="37" pageNumber="38">
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The paper on Cuban oonopids by
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<bibRefCitation id="9F5F4B26D944FFA4FFBFFABD17ACFAC6" author="Dumitresco, M. & M. Georgesco" box="[88,505,1291,1315]" pageId="37" pageNumber="38" pagination="65 - 114" refId="ref29872" refString="Dumitresco, M., and M. Georgesco. 1983. Sur les Oonopidae (Araneae) de Cuba. Resultats des Expeditions Biospeologiques Cubano- Roumaines a Cuba 4: 65 - 114." type="journal article" year="1983">Dumitresco and Georgesco (1983)</bibRefCitation>
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includes descriptions of at least three species that may belong to
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<taxonomicName id="3CCE4D54D944FFA4FF36FAF0170BFABB" authorityName="Dalmas" authorityYear="1916" box="[209,350,1350,1374]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="C9BAEAC5D944FFA4FF36FAF0170BFABB" box="[209,350,1350,1374]" italics="true" pageId="37" pageNumber="38">Heteroonops</emphasis>
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</taxonomicName>
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, but all three descriptions are problematic. As indicated above, the specimens studied by those investigators are not available on loan, but probably wouldn’t prove to be very helpful even if they could be re-examined. Judging from the detailed figures in their paper, Dumitresco and Georgesco treated the specimens like mites, and probably mounted them in Hoyer’s medium on ‘‘permanent’’ slides. During that process, the male and female genitalia would have been smashed flat by the weight of the cover slips, making comparisons with normal, three-dimensional specimens difficult.
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</paragraph>
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<paragraph id="FB7136D7D944FFA4FD57FF651528FCDB" blockId="37.[662,1188,211,1753]" pageId="37" pageNumber="38">
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<materialsCitation id="4BA63C8AD944FFA4FD57FF65152FFCDB" collectorName="de Pinos" country="Iceland" location="Isla" pageId="37" pageNumber="38" specimenCount="2" specimenCount-female="2">
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The only Cuban specimens of the genus we have been able to examine are the
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<specimenCount id="EDC8FD5ED944FFA4FBFAFF4712F1FEEC" box="[1053,1188,241,265]" pageId="37" pageNumber="38" type="female">two females</specimenCount>
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of
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<taxonomicName id="3CCE4D54D944FFA4FD51FEB91503FEC2" authorityName="Simon" authorityYear="1891" box="[694,854,271,295]" class="Arachnida" family="Oonopidae" genus="Oonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="spinimanus">
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<emphasis id="C9BAEAC5D944FFA4FD51FEB91503FEC2" box="[694,854,271,295]" italics="true" pageId="37" pageNumber="38">H. spinimanus</emphasis>
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</taxonomicName>
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recorded above.
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<bibRefCitation id="9F5F4B26D944FFA4FBF9FEB81593FEA1" author="Dumitresco, M. & M. Georgesco" pageId="37" pageNumber="38" pagination="65 - 114" refId="ref29872" refString="Dumitresco, M., and M. Georgesco. 1983. Sur les Oonopidae (Araneae) de Cuba. Resultats des Expeditions Biospeologiques Cubano- Roumaines a Cuba 4: 65 - 114." type="journal article" year="1983">Dumitresco and Georgesco (1983: 88</bibRefCitation>
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, pl. 13, figs. 1–5) recorded
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<specimenCount id="EDC8FD5ED944FFA4FCE4FEFC15D7FE87" box="[771,898,330,354]" pageId="37" pageNumber="38" type="female">one female</specimenCount>
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of that species from the
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<collectingCountry id="83D97647D944FFA4FD71FEDE14EAFE65" box="[662,703,360,384]" name="Iceland" pageId="37" pageNumber="38">Isla</collectingCountry>
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<collectorName id="563B5301D944FFA4FD2EFEDE1565FE65" box="[713,816,360,384]" pageId="37" pageNumber="38">de Pinos</collectorName>
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, but we are not convinced that this female was correctly identified. First, as noted above, Dumitresco and Georgesco indicated that this specimen lacks a dilation on the palpal patella; in addition, their illustrations show that the anterior receptaculum is scarcely expanded at its anterior end. One possible complication is that the overall appearance of the female genitalia, in this species (and perhaps the others), depends on whether or not the epigastric furrow is open; a conical projection at the rear of the anterior portion of the female genitalia apparently inserts into a conical invagination at the front of the posterior portion, when the furrow is closed (see fig. 131)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="FB7136D7D944FFA4FD57FCF21553FB3A" blockId="37.[662,1188,211,1753]" pageId="37" pageNumber="38">
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They described a second species as ‘‘
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<emphasis id="C9BAEAC5D944FFA4FD43FCD41586FC9F" box="[676,979,866,890]" italics="true" pageId="37" pageNumber="38">
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<taxonomicName id="3CCE4D54D944FFA4FD43FCD41564FC9F" authorityName="Dalmas" authorityYear="1916" box="[676,817,866,890]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Heteroonops</taxonomicName>
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(?)
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<taxonomicName id="3CCE4D54D944FFA4FC9BFCD41586FC9F" authority="Platnick & Dupérré, 2009" authorityName="Platnick & Dupérré" authorityYear="2009" box="[892,979,866,890]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="colombi" status="sp. nov.">colombi</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="D28957BED944FFA4FC02FCD41265FC9F" box="[997,1072,866,890]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
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’’ (
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<bibRefCitation id="9F5F4B26D944FFA4FBBFFCD41258FC7D" author="Dumitresco, M. & M. Georgesco" pageId="37" pageNumber="38" pagination="65 - 114" refId="ref29872" refString="Dumitresco, M., and M. Georgesco. 1983. Sur les Oonopidae (Araneae) de Cuba. Resultats des Expeditions Biospeologiques Cubano- Roumaines a Cuba 4: 65 - 114." type="journal article" year="1983">Dumitresco and Georgesco, 1983: 90</bibRefCitation>
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, pl. 14, figs. 1, 2). Again, they had a single female, from the Cueva Cinco Cuevas in
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<collectingRegion id="390AF835D944FFA4FBF3FC0D14A9FC14" country="Cuba" name="La Habana" pageId="37" pageNumber="38">La Habana province</collectingRegion>
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. That female is missing both pedipalps, and hence its generic placement remains debatable. They presented only a ventral view of the female genitalia, which do not closely resemble those of any of the species of
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<taxonomicName id="3CCE4D54D944FFA4FD71FBD81576FB63" authorityName="Dalmas" authorityYear="1916" box="[662,803,1134,1158]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9BAEAC5D944FFA4FD71FBD81576FB63" box="[662,803,1134,1158]" italics="true" pageId="37" pageNumber="38">Heteroonops</emphasis>
|
||
</taxonomicName>
|
||
known to us. The identity and placement of this species will remain enigmatic until topotypical specimens become available for study.
|
||
</paragraph>
|
||
<paragraph id="FB7136D7D944FFA4FD57FB5315C0F93C" blockId="37.[662,1188,211,1753]" pageId="37" pageNumber="38">
|
||
Finally, those authors (Dumitesco and Gerogesco, 1983: 81, pl. 10, figs. 1–7) assigned to
|
||
<taxonomicName id="3CCE4D54D944FFA4FD5AFA97125FFADC" authority="Chickering" authorityName="Chickering" authorityYear="1971" box="[701,1034,1313,1337]" class="Arachnida" family="Oonopidae" genus="Oonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="castellus">
|
||
<emphasis id="C9BAEAC5D944FFA4FD5AFA97152AFADC" box="[701,895,1313,1337]" italics="true" pageId="37" pageNumber="38">Oonops castellus</emphasis>
|
||
Chickering
|
||
</taxonomicName>
|
||
<specimenCount id="EDC8FD5ED944FFA4FBFDFA971485FAB2" pageId="37" pageNumber="38" type="generic">three specimens</specimenCount>
|
||
from two widely separated Cuban localities. They noted some discrepancies between their specimens and those described from the
|
||
<collectingCountry id="83D97647D944FFA4FCEAFA2E15E1FA55" box="[781,948,1432,1456]" name="Virgin Islands" pageId="37" pageNumber="38">Virgin Islands</collectingCountry>
|
||
by Chickering, but concluded that the differences did not justify the establishment of a new species for any of the Cuban specimens. Their illustrations suggest that these Cuban specimens may belong to
|
||
<taxonomicName id="3CCE4D54D944FFA4FCF0F99B15F1F9A0" authorityName="Dalmas" authorityYear="1916" box="[791,932,1581,1605]" class="Arachnida" family="Oonopidae" genus="Heteroonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9BAEAC5D944FFA4FCF0F99B15F1F9A0" box="[791,932,1581,1605]" italics="true" pageId="37" pageNumber="38">Heteroonops</emphasis>
|
||
</taxonomicName>
|
||
, but they also leave little doubt that they do not belong to
|
||
<taxonomicName id="3CCE4D54D944FFA4FB61F9FD14A1F965" authorityName="Chickering" authorityYear="1971" class="Arachnida" family="Oonopidae" genus="Oonops" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="castellus">
|
||
<emphasis id="C9BAEAC5D944FFA4FB61F9FD14A1F965" italics="true" pageId="37" pageNumber="38">H. castellus</emphasis>
|
||
</taxonomicName>
|
||
. Here again, the generic and specific placement of these specimens will remain indeterminate until topotypical material can be obtained for study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |