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<mods:title id="D5CE6A1726C6E4370D3C28B1DF168A2B">New species and new records of mites of the family Laelapidae (Acari: Mesostigmata) associated with ants in Iran</mods:title>
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<subSubSection id="B3BC3BC8FFEBFF894EF4FE7CFE13FE05" box="[151,509,475,502]" pageId="1" pageNumber="23" type="nomenclature">
<paragraph id="FB196843FFEBFF894EF4FE7CFE13FE05" blockId="1.[151,509,475,502]" box="[151,509,475,502]" pageId="1" pageNumber="23">
<heading id="A051DF2FFFEBFF894EF4FE7CFE13FE05" bold="true" box="[151,509,475,502]" fontSize="11" level="1" pageId="1" pageNumber="23" reason="1">
<emphasis id="C9D2B451FFEBFF894EF4FE7CFE13FE05" bold="true" box="[151,509,475,502]" pageId="1" pageNumber="23">
Genus
<taxonomicName id="3CA613C0FFEBFF894E8DFE7BFE13FE05" authority="Berlese" authorityName="Berlese" box="[238,509,475,502]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894E8DFE7BFE75FE06" bold="true" box="[238,411,476,502]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
Berlese
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B3BC3BC8FFEBFF894EF4FD84FD99FDC8" box="[151,631,546,568]" pageId="1" pageNumber="23" type="reference_group">
<paragraph id="FB196843FFEBFF894EF4FD84FD99FDC8" blockId="1.[151,927,546,599]" box="[151,631,546,568]" pageId="1" pageNumber="23">
<treatmentCitationGroup id="DBB64F6DFFEBFF894EF4FD84FD99FDC8" box="[151,631,546,568]" pageId="1" pageNumber="23">
<taxonomicName id="3CA613C0FFEBFF894EF4FD84FD9DFDC8" authority="Berlese, 1916: 170" authorityName="Berlese" authorityPageNumber="170" authorityYear="1916" box="[151,627,546,568]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="subGenus" subGenus="Gymnolaelaps">
<emphasis id="C9D2B451FFEBFF894EF4FD84FEECFDC8" box="[151,258,547,568]" italics="true" pageId="1" pageNumber="23">Hypoaspis</emphasis>
(
<emphasis id="C9D2B451FFEBFF894F73FD85FE4CFDC8" box="[272,418,546,568]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
)
<treatmentCitation id="7A074E52FFEBFF894FD2FD85FD9DFDC8" author="Berlese" box="[433,627,546,568]" page="170" pageId="1" pageNumber="23" year="1916">
<bibRefCitation id="9F3715B2FFEBFF894FD2FD85FDAEFDC8" author="Berlese" box="[433,576,546,568]" pageId="1" pageNumber="23" refString="Berlese, A. (1916) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
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<subSubSection id="B3BC3BC8FFEBFF894EF4FDE5FC71FDA7" box="[151,927,577,599]" pageId="1" pageNumber="23" type="materials_examined">
<paragraph id="FB196843FFEBFF894EF4FDE5FC71FDA7" blockId="1.[151,927,546,599]" box="[151,927,577,599]" pageId="1" pageNumber="23">
<typeStatus id="241DD6E1FFEBFF894EF4FDE5FF26FDA7" box="[151,200,578,599]" pageId="1" pageNumber="23">Type</typeStatus>
species
<taxonomicName id="3CA613C0FFEBFF894F7CFDE6FD4AFDA7" authority="Berlese, 1892" authorityName="Berlese" authorityYear="1892" box="[287,676,577,599]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="C9D2B451FFEBFF894F7CFDE6FDFEFDA7" box="[287,528,577,599]" italics="true" pageId="1" pageNumber="23">Laelaps myrmecophilus</emphasis>
<bibRefCitation id="9F3715B2FFEBFF894C75FDE6FD4AFDA7" author="Berlese" box="[534,676,577,599]" pageId="1" pageNumber="23" refString="Berlese, A. (1892) Acari, Myriopoda et Scorpiones hucusque in Italia reperta, 69, 22 text pages + Plates 1 - 10. (Reprint by Junk, The Hague, 1979)." type="journal article" year="1892">Berlese, 1892</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="B3BC3BC8FFEBFF894EF4FD24FDC7FAF0" pageId="1" pageNumber="23" type="diagnosis">
<paragraph id="FB196843FFEBFF894EF4FD24FAD4FCBF" blockId="1.[151,1436,643,1280]" pageId="1" pageNumber="23">
<emphasis id="C9D2B451FFEBFF894EF4FD24FEFFFD6C" bold="true" box="[151,273,643,668]" pageId="1" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="3CA613C0FFEBFF894F7EFD23FE76FD6C" box="[285,408,644,668]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
usually with a three-tined palp tarsal claw, proximal tine occasionally reduced, rarely absent; genital shield large, abutting anal shield, with 13 pairs of setae in addition to the genital setae st5, all located on the edges of the shield; pre-sternal shield well-developed and sclerotised; a large subtriangular podal shield usually present posterior to coxa IV; epistome with irregularly serrated anterior margin; genu IV with nine setae including one ventral seta. Dorsal shield often with paired Zx setae between J and Z setae, unpaired Jx setae also often present; dorsal shield setae distally pointed, smooth or slightly serrated, not long and whip-like.
</paragraph>
<paragraph id="FB196843FFEBFF894EA4FCFBFDC7FAF0" blockId="1.[151,1436,643,1280]" pageId="1" pageNumber="23">
<emphasis id="C9D2B451FFEBFF894EA4FCFBFE5CFC84" bold="true" box="[199,434,860,884]" pageId="1" pageNumber="23">Notes on the genus.</emphasis>
The genus
<taxonomicName id="3CA613C0FFEBFF894C5BFCFBFD34FC84" box="[568,730,860,884]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894C5BFCFBFD34FC84" box="[568,730,860,884]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
has been collected in many parts of the world, almost always from ants' nests. The genus includes approximately 35 described species (
<bibRefCitation id="9F3715B2FFEBFF894DB5FCD8FB80FC67" author="Hunter" box="[982,1134,895,919]" pageId="1" pageNumber="23" refString="Hunter, P. E. (1967) Comments on Hypoaspis (Gymnolaelaps) Berlese, 1916, with description of a new species (Acarina: Dermanyssidae; Laelapinae). Journal of the Georgia Entomological Society, 2, 100 - 102 + Figure 1." type="journal article" year="1967">Hunter, 1967</bibRefCitation>
;
<bibRefCitation id="9F3715B2FFEBFF894A19FCD8FAD2FC67" author="Gu" box="[1146,1340,895,919]" pageId="1" pageNumber="23" refString="Gu, Y. - M. &amp; Guo, X. - G. (1997) A new species of the genus Gymnolaelaps from China (Acari: Laelapidae). Acta Zootaxonomica Sinica, 22, 246 - 248." type="journal article" year="1997">Gu &amp; Guo, 1997</bibRefCitation>
; Afifi &amp; Abdel-Halim, 1998).
<taxonomicName id="3CA613C0FFEBFF894FEBFC03FDC4FC4C" box="[392,554,932,956]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894FEBFC03FDC4FC4C" box="[392,554,932,956]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
has often been considered as a subgenus of a very broad genus
<taxonomicName id="3CA613C0FFEBFF894A97FC02FA84FC4C" box="[1268,1386,933,956]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894A97FC02FA84FC4C" box="[1268,1386,933,956]" italics="true" pageId="1" pageNumber="23">Hypoaspis</emphasis>
</taxonomicName>
, but we consider it to be a separate genus, following
<bibRefCitation id="9F3715B2FFEBFF894CD4FC6EFC72FC10" author="Evans" box="[695,924,968,992]" pageId="1" pageNumber="23" refString="Evans, G. O. &amp; Till, W. M. (1979) Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London, 35, 145 - 270." type="journal article" year="1979">Evans &amp; Till (1979)</bibRefCitation>
.
<bibRefCitation id="9F3715B2FFEBFF894DCBFC6FFB63FC10" box="[936,1165,968,992]" pageId="1" pageNumber="23" refString="Nemati, A. R., Mossadegh, M. S. &amp; Kamali, K. (2000) Some mites the Laelapidae (Acari: Mesostigmata) collected from ant and termite nests in Ahwaz Region, Khuzestan Province. In: Abstracts, 14 th Iranian Plant Protection Congress, Isfahan University of Technology, September 2000, Volume 1, p. 381." type="journal article">
Nemati
<emphasis id="C9D2B451FFEBFF894A61FC6EFBDDFC10" box="[1026,1075,968,992]" italics="true" pageId="1" pageNumber="23">et al</emphasis>
. (2000)
</bibRefCitation>
reported five species of
<taxonomicName id="3CA613C0FFEBFF894EF4FC4BFED7FBF4" box="[151,313,1004,1028]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894EF4FC4BFED7FBF4" box="[151,313,1004,1028]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
from Iran—
<taxonomicName id="3CA613C0FFEBFF894FA8FC4AFD1DFBF4" authority="Berlese, 1923" authorityName="Berlese" authorityYear="1923" box="[459,755,1004,1028]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="C9D2B451FFEBFF894FA8FC4AFE32FBF4" box="[459,476,1005,1028]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894F8FFC4BFDD6FBF4" box="[492,568,1004,1028]" italics="true" pageId="1" pageNumber="23">hospes</emphasis>
(
<bibRefCitation id="9F3715B2FFEBFF894C29FC4BFD05FBF4" author="Berlese" box="[586,747,1004,1028]" pageId="1" pageNumber="23" refString="Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia, 15, 237 - 262." type="journal article" year="1923">Berlese, 1923</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="3CA613C0FFEBFF894D61FC4AFB97FBF4" authority="Michael, 1891" authorityName="Michael" authorityYear="1891" box="[770,1145,1004,1028]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmophilus">
<emphasis id="C9D2B451FFEBFF894D61FC4AFCFDFBF4" box="[770,787,1005,1028]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894D40FC4BFC5BFBF4" box="[803,949,1004,1028]" italics="true" pageId="1" pageNumber="23">myrmophilus</emphasis>
(
<bibRefCitation id="9F3715B2FFEBFF894DA4FC4BFB81FBF4" author="Michael" box="[967,1135,1004,1028]" pageId="1" pageNumber="23" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 43, 638 - 653 + Plates XLIX - L." type="journal article" year="1891">Michael, 1891</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="3CA613C0FFEBFF894AE5FC4AFF3CFBD8" authority="Womersley, 1956" authorityName="Womersley" authorityYear="1956" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="vitzthumi">
<emphasis id="C9D2B451FFEBFF894AE5FC4AFB79FBF4" box="[1158,1175,1005,1028]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894ACBFC4BFAE1FBF4" box="[1192,1295,1004,1028]" italics="true" pageId="1" pageNumber="23">vitzthumi</emphasis>
<bibRefCitation id="9F3715B2FFEBFF894B7BFC4BFF3CFBD8" author="Womersley" pageId="1" pageNumber="23" refString="Womersley, H. (1956) On some new Acarina-Mesostigmata from Australia, New Zealand and New Guinea. Journal of the Linnean Society (Zoology), 42, 505 - 599." type="journal article" year="1956">Womersley, 1956</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="3CA613C0FFEBFF894EBCFBB7FDA0FBD8" authority="Womersley, 1956" authorityName="Womersley" authorityYear="1956" box="[223,590,1040,1064]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="australicus">
<emphasis id="C9D2B451FFEBFF894EBCFBB7FF1EFBD7" box="[223,240,1040,1063]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894F63FBB7FE93FBD8" box="[256,381,1040,1064]" italics="true" pageId="1" pageNumber="23">australicus</emphasis>
<bibRefCitation id="9F3715B2FFEBFF894FE5FBB7FDA0FBD8" author="Womersley" box="[390,590,1040,1064]" pageId="1" pageNumber="23" refString="Womersley, H. (1956) On some new Acarina-Mesostigmata from Australia, New Zealand and New Guinea. Journal of the Linnean Society (Zoology), 42, 505 - 599." type="journal article" year="1956">Womersley, 1956</bibRefCitation>
</taxonomicName>
, and an unidentified species. We follow
<bibRefCitation id="9F3715B2FFEBFF894A47FBB6FB28FBD8" author="Hunter" box="[1060,1222,1040,1064]" pageId="1" pageNumber="23" refString="Hunter, P. E. (1961) The genus Laelaspis, with descriptions of three new species (Acarina: Laelaptidae). Annals of the Entomological Society of America, 54, 672 - 683." type="journal article" year="1961">Hunter (1961)</bibRefCitation>
in placing
<taxonomicName id="3CA613C0FFEBFF894B29FBB7FF39FBBC" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="vitzthumi">
<emphasis id="C9D2B451FFEBFF894B29FBB7FAB5FBD7" box="[1354,1371,1040,1063]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894B0FFBB6FF39FBBC" italics="true" pageId="1" pageNumber="23">vitzthumi</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="3CA613C0FFEBFF894F17FB93FE31FBBC" box="[372,479,1076,1100]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894F17FB93FE31FBBC" box="[372,479,1076,1100]" italics="true" pageId="1" pageNumber="23">Laelaspis</emphasis>
</taxonomicName>
.
<bibRefCitation id="9F3715B2FFEBFF894F8CFB92FD7FFBBC" author="Hunter" box="[495,657,1076,1100]" pageId="1" pageNumber="23" refString="Hunter, P. E. (1967) Comments on Hypoaspis (Gymnolaelaps) Berlese, 1916, with description of a new species (Acarina: Dermanyssidae; Laelapinae). Journal of the Georgia Entomological Society, 2, 100 - 102 + Figure 1." type="journal article" year="1967">Hunter (1967)</bibRefCitation>
reported that the
<typeStatus id="241DD6E1FFEBFF894D3CFB92FC61FBBD" box="[863,911,1077,1101]" pageId="1" pageNumber="23">type</typeStatus>
species
<taxonomicName id="3CA613C0FFEBFF894D91FB92FB50FBBC" box="[1010,1214,1076,1100]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="C9D2B451FFEBFF894D91FB92FBEDFBBC" box="[1010,1027,1077,1100]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894A70FB93FB50FBBC" box="[1043,1214,1076,1100]" italics="true" pageId="1" pageNumber="23">myrmecophilus</emphasis>
</taxonomicName>
has three stronglydeveloped tines on the palp tarsal claw, as do most other species in the genus. Two species,
<taxonomicName id="3CA613C0FFEBFF894AF9FBFFFF36FB64" authority="Berlese, 1892" authorityName="Berlese" authorityYear="1892" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="canestrinii">
<emphasis id="C9D2B451FFEBFF894AF9FBFFFB45FB9F" box="[1178,1195,1112,1135]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894ADAFBFEFADFFB80" box="[1209,1329,1113,1136]" italics="true" pageId="1" pageNumber="23">canestrinii</emphasis>
(
<bibRefCitation id="9F3715B2FFEBFF894B5CFBFFFF3EFB64" author="Berlese" pageId="1" pageNumber="23" refString="Berlese, A. (1892) Acari, Myriopoda et Scorpiones hucusque in Italia reperta, 69, 22 text pages + Plates 1 - 10. (Reprint by Junk, The Hague, 1979)." type="journal article" year="1892">Berlese, 1892</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="3CA613C0FFEBFF894F6DFBDAFE50FB64" box="[270,446,1148,1172]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmophilus">
<emphasis id="C9D2B451FFEBFF894F6DFBDAFEF1FB64" box="[270,287,1149,1172]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894F4FFBDBFE50FB64" box="[300,446,1148,1172]" italics="true" pageId="1" pageNumber="23">myrmophilus</emphasis>
</taxonomicName>
, have a weakly-developed proximal tine, and in
<taxonomicName id="3CA613C0FFEBFF894DBEFBDAFB19FB64" authority="Hunter, 1967" authorityName="Hunter" authorityYear="1967" box="[989,1271,1148,1172]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="krantzi">
<emphasis id="C9D2B451FFEBFF894DBEFBDAFC00FB64" box="[989,1006,1149,1172]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894D9FFBDBFBA4FB64" box="[1020,1098,1148,1172]" italics="true" pageId="1" pageNumber="23">krantzi</emphasis>
(
<bibRefCitation id="9F3715B2FFEBFF894A3BFBDBFB1EFB64" author="Hunter" box="[1112,1264,1148,1172]" pageId="1" pageNumber="23" refString="Hunter, P. E. (1967) Comments on Hypoaspis (Gymnolaelaps) Berlese, 1916, with description of a new species (Acarina: Dermanyssidae; Laelapinae). Journal of the Georgia Entomological Society, 2, 100 - 102 + Figure 1." type="journal article" year="1967">Hunter, 1967</bibRefCitation>
)
</taxonomicName>
, there are only two tines.
<bibRefCitation id="9F3715B2FFEBFF894F6FFB07FDE0FB48" box="[268,526,1184,1208]" pageId="1" pageNumber="23" refString="Jalalizand, A. R., Ostovan, H., Kamali, K. &amp; Hatami, B. (2004) Investigation on the fauna of mites (Acari) on elm trees in Isfahan. In: Abstracts, 16 th Iranian Plant Protection Congress, University of Tabriz, August 2004, Volume 1, p. 255." type="journal article">
Jalalizand
<emphasis id="C9D2B451FFEBFF894FE0FB06FE5AFB48" box="[387,436,1184,1208]" italics="true" pageId="1" pageNumber="23">et al</emphasis>
. (2004)
</bibRefCitation>
reported
<taxonomicName id="3CA613C0FFEBFF894C1AFB07FCDAFB48" box="[633,820,1184,1208]" class="Arachnida" family="Laelapidae" genus="Pseudoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894C1AFB07FCDAFB48" box="[633,820,1184,1208]" italics="true" pageId="1" pageNumber="23">Pseudoparasitus</emphasis>
</taxonomicName>
sp. near
<taxonomicName id="3CA613C0FFEBFF894DF9FB07FB73FB48" authority="Womersley" authorityName="Womersley" box="[922,1181,1184,1208]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="australicus">
<emphasis id="C9D2B451FFEBFF894DF9FB07FBF8FB48" box="[922,1046,1184,1208]" italics="true" pageId="1" pageNumber="23">australicus</emphasis>
Womersley
</taxonomicName>
and
<taxonomicName id="3CA613C0FFEBFF894AB7FB06FF10FB2C" authority="Berlese" authorityName="Berlese" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="C9D2B451FFEBFF894AB7FB06FB0CFB48" box="[1236,1250,1185,1208]" italics="true" pageId="1" pageNumber="23">P</emphasis>
.
<emphasis id="C9D2B451FFEBFF894A92FB07FA72FB48" box="[1265,1436,1184,1208]" italics="true" pageId="1" pageNumber="23">myrmecophilus</emphasis>
(Berlese)
</taxonomicName>
from
<collectingCountry id="83B128D3FFEBFF894F22FB62FE9CFB2C" box="[321,370,1221,1244]" name="Iran" pageId="1" pageNumber="23">Iran</collectingCountry>
. We place both of these species in the genus
<taxonomicName id="3CA613C0FFEBFF894D01FB63FBEAFB2C" box="[866,1028,1220,1244]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C9D2B451FFEBFF894D01FB63FBEAFB2C" box="[866,1028,1220,1244]" italics="true" pageId="1" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
, but we did not find specimens of
<taxonomicName id="3CA613C0FFEBFF894BE7FB62FEACFAF0" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="C9D2B451FFEBFF894BE7FB62FA7BFB2C" box="[1412,1429,1221,1244]" italics="true" pageId="1" pageNumber="23">G</emphasis>
.
<emphasis id="C9D2B451FFEBFF894EF4FB4FFEACFAF0" box="[151,322,1256,1280]" italics="true" pageId="1" pageNumber="23">myrmecophilus</emphasis>
</taxonomicName>
in the present study.
</paragraph>
</subSubSection>
</treatment>
</document>