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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.236.108651" ID-Pensoft-Pub="1314-2003-236-97" ID-Pensoft-UUID="B3D2DC6712FC5FA697A2387ADAAB8A62" ModsDocID="1314-2003-236-97" checkinTime="1701930837961" checkinUser="pensoft" docAuthor="de Mestier, Astrid, Campos Pineda, Ernesto, Cedeno Fonseca, Marco &amp; Ortiz, Orlando O." docDate="2023" docId="724A0DAA225B5D1F9EF2BD8ECFA63B9B" docLanguage="en" docName="PhytoKeys 236: 97-112" docOrigin="PhytoKeys 236" docPubDate="2023-12-06" docSource="http://dx.doi.org/10.3897/phytokeys.236.108651" docTitle="Casearia isthmica de Mestier &amp; O. Ortiz 2023, sp. nov." docType="treatment" docVersion="2" id="B3D2DC6712FC5FA697A2387ADAAB8A62" lastPageNumber="97" masterDocId="B3D2DC6712FC5FA697A2387ADAAB8A62" masterDocTitle="A new species of Casearia Jacq. (Salicaceae) from Central Panama and insights into its phylogenetic position within the genus" masterLastPageNumber="112" masterPageNumber="97" pageNumber="97" updateTime="1701931139480" updateUser="ExternalLinkService">
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<mods:title id="E5535E52494839AE99DB1120E7E35A02">A new species of Casearia Jacq. (Salicaceae) from Central Panama and insights into its phylogenetic position within the genus</mods:title>
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<mods:namePart id="02B5E1396CD1E58B93C5A4DB9D511281">de Mestier, Astrid</mods:namePart>
<mods:nameIdentifier id="370945D8CD5FECCF46125B34816AC084" type="ORCID">https://orcid.org/0000-0002-9976-8196</mods:nameIdentifier>
<mods:affiliation id="56E797B3F84244B35D274A53B54EB7D8">Institut fuer Biologie - Systematische Botanik und Pflanzengeographie, Freie Universitaet Berlin, Altensteinstrasse 6, 14195 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier id="362FC2BEA4A93D9363F42A043302628E" type="email">astriddemestier@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D13AC71FE1196134A18A6555981959BC">Campos Pineda, Ernesto</mods:namePart>
<mods:nameIdentifier id="C2E0235FF909EFB95614931BFE979514" type="ORCID">https://orcid.org/0000-0001-8606-5785</mods:nameIdentifier>
<mods:affiliation id="EDD5EC8ABC72532E2FD09B3D721B0491">Smithsonian Tropical Research Institute, Box 0843 - 03092, Balboa, Ancon, Panama</mods:affiliation>
</mods:name>
<mods:name id="EE7E834C4DF3BCBDC1B502FB64A67F35" type="personal">
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<mods:namePart id="12822E6DDA5980ECCA637078BCDB11CC">Cedeno Fonseca, Marco</mods:namePart>
<mods:nameIdentifier id="DD51D99C27FBBDBD0C137BC9F2C18FF5" type="ORCID">https://orcid.org/0000-0003-0119-0427</mods:nameIdentifier>
<mods:affiliation id="DE98ABF943CE2A3C6810A846B91A33C9">Institut fuer Biologie - Systematische Botanik und Pflanzengeographie, Freie Universitaet Berlin, Altensteinstrasse 6, 14195 Berlin, Germany &amp; Botanischer Garten und Botanisches Museum Berlin, Freie Universitaet Berlin, Koenigin-Luise-Str. 6 - 8, 14195 Berlin, Germany &amp; Herbario Luis Fournier Origgi (USJ), Centro de Investigacion en Biodiversidad y Ecologia Tropical, Universidad de Costa Rica, Apdo. 11501 - 2060, San Jose ́, Costa Rica</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="8B07207BFFE38BE884CEF3FD1942697B">Ortiz, Orlando O.</mods:namePart>
<mods:nameIdentifier id="724B3539EB7A22C786D957BE30656FF1" type="ORCID">https://orcid.org/0000-0002-7805-0046</mods:nameIdentifier>
<mods:affiliation id="52A851C798EA450F0E35C4DA4AF6C4D8">Institut fuer Biologie - Systematische Botanik und Pflanzengeographie, Freie Universitaet Berlin, Altensteinstrasse 6, 14195 Berlin, Germany &amp; Jardin Botanico Lankester, Universidad de Costa Rica, Apdo. 302 - 7050, Cartago, Costa Rica &amp; Herbario PMA, Universidad de Panama, Estafeta Universitaria, Panama City, Panama</mods:affiliation>
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<mods:part id="17E63C49BF3A5FF146073E923ED00A0B">
<mods:date id="479171CAF6FBBA12CDE7CEBE1771EF50">2023</mods:date>
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<mods:number id="3C790533B7FA2998D37EE821289CF706">236</mods:number>
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<mods:identifier id="54FEDACA490EB1CFF4D6D2CEADDB3FE1" type="DOI">http://dx.doi.org/10.3897/phytokeys.236.108651</mods:identifier>
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<treatment id="724A0DAA225B5D1F9EF2BD8ECFA63B9B" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B" httpUri="http://treatment.plazi.org/id/724A0DAA225B5D1F9EF2BD8ECFA63B9B" lastPageNumber="97" pageId="0" pageNumber="97">
<subSubSection id="D85D0EE44D56E6707802479396116DE3" pageId="0" pageNumber="97" type="nomenclature">
<paragraph id="3F56FD75A8A8713DE90E63AE70BB6698" pageId="0" pageNumber="97">
<taxonomicName id="4049D497BAE8895C90F3FEE4DAF2680B" LSID="724A0DAA-225B-5D1F-9EF2-BD8ECFA63B9B" authority="de Mestier &amp; O. Ortiz" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Salicaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica" status="sp. nov.">Casearia isthmica de Mestier &amp; O.Ortiz</taxonomicName>
<taxonomicNameLabel id="F86E64C28633F309343B75DBB1E27D59" pageId="0" pageNumber="97">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2BB616E297CA0715CAFF3AC39B39086C" pageId="0" pageNumber="97" type="description">
<paragraph id="0B46C3C13F340F85D6F8267BDF889FD8" pageId="0" pageNumber="97">
<figureCitation id="407300C25A067E4DED295764B6C25A8C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Casearia isthmica de Mestier &amp; O. Ortiz A flowering branch B inflorescence C flowers at anthesis D fruiting branch E leaves F ripe fruit G bark. Photos A, C, E (back side of the leaf) by Carmen Galdames (Galdames 6153). Photos D, E (front side of the leaf) by Carmen Galdames (Galdames 6642). Photo B by Ernesto Campos (Sumich 151). Photos G, E by Ernesto Campos Plate by Marco Cedeno-Fonseca." figureDoi="10.3897/phytokeys.236.108651.figure3" httpUri="https://binary.pensoft.net/fig/948452" pageId="0" pageNumber="97">Figs 3</figureCitation>
<figureCitation id="F661A493494413912AD1DE520853FAC6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Casearia isthmica de Mestier &amp; O. Ortiz A branches B twig and stipules C trunk base and roots D lenticellate bark. Photos A, B by Carmen Galdames (Galdames 6642). Photo C by Ernesto Campos (Campos 1087). Photo D by Ernesto Campos. Plate by Marco Cedeno-Fonseca." figureDoi="10.3897/phytokeys.236.108651.figure4" httpUri="https://binary.pensoft.net/fig/948453" pageId="0" pageNumber="97">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8F5FF7BF51460BF5CE5102F07F492902" pageId="0" pageNumber="97" type="materials_examined">
<paragraph id="46FBB6D182B973F5044777D3051F1E99" pageId="0" pageNumber="97">
<typeStatus id="800887563BBBAE21B92911032AAA6C97">Type</typeStatus>
.
</paragraph>
<paragraph id="2AF8A815BE8B61F7CE96F729896CD261" pageId="0" pageNumber="97">
<materialsCitation id="DFEAF5DFEA4E045FA97C61C2C70A47EB" collectingDate="2022-04-27" collectorName="La Rana Dorada, E. Campos, J. Sumich" country="Panama" elevation="100" latitude="8.692223" location="La Rana Dorada" longLatPrecision="21" longitude="-79.92611" municipality="Parque Nacional Altos de Campana" specimenCount="1" stateProvince="Distrito de Capira" typeStatus="holotype">
<emphasis id="B31A07A240FD451C8A36E8FF87A27570" bold="true" pageId="0" pageNumber="97">
<collectingCountry id="56F77FA8DA7D93290B195F3F49CF5B12" name="Panama">Panama</collectingCountry>
.
<collectingRegion id="295730EBB17BF780D13D8FEC2DC464CD">Distrito de Capira</collectingRegion>
</emphasis>
:
<collectingMunicipality id="AB9A9C27BC52D2B3BEEF67D75BD7A92C">Parque Nacional Altos de Campana</collectingMunicipality>
, sendero
<collectorName id="74257076EF4EC0315501CE95AA831E5D">
<location id="3B7BEF8592B66C7BE547F8118B86E3A6" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:3B7BEF8592B66C7BE547F8118B86E3A6" country="Panama" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Parque Nacional Altos de Campana" name="La Rana Dorada" stateProvince="Distrito de Capira">La Rana Dorada</location>
</collectorName>
, ca.
<quantity id="3EDCBA0A4CF37F57DCC5FC8AE2B1964E" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="8055634502BADC1791D4BE868D792C59" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
desde la entrada,
<geoCoordinate id="02C3CE25815D50C98F70C431D3378ED1" degrees="8" direction="north" minutes="41" orientation="latitude" precision="15" seconds="32" value="8.692223">8°41'32&quot;N</geoCoordinate>
,
<geoCoordinate id="F722635B0D1BCBA7D4C984A0356E1F62" degrees="79" direction="west" minutes="55" orientation="longitude" precision="15" seconds="34" value="-79.92611">79°55'34&quot;W</geoCoordinate>
,
<quantity id="65A446C6186F11BCB0B51A671FC7F961" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">
<elevation id="56B756BB40A0E6308322DDED7CE006A9" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">839 m</elevation>
</quantity>
,
<collectingDate id="F758BA23EA9FFA5637B63F6ED3AB31B0" value="2022-04-27">27 April 2022</collectingDate>
,
<emphasis id="FB5E1CB553126243C97471059AAAEF41" italics="true" pageId="0" pageNumber="97">
<collectorName id="F9A70EE504395262D45331A80C25FEB8">E. Campos</collectorName>
&amp;
<collectorName id="726495CA60A0627FDB753B93FF1143DF">J. Sumich</collectorName>
1329
</emphasis>
(SAL302) (
<typeStatus id="C43164215B59AB1B0892C12A26234D89">holotype</typeStatus>
: PMA-127644!; isotypes: B-101233230!, SCZ! two sheets, barcodes: 20033 and 20034)
</materialsCitation>
.
</paragraph>
<paragraph id="CAC98FF9426BA4AEA92582022ED4AA93" pageId="0" pageNumber="97">
<taxonomicName id="A170CC42AFA9CF4E4E3EAD712C5A2284" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="D856F187B1D77C7244D92D2F23D4D9FC" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
shares morphological similarities with
<taxonomicName id="9D91E6E343EC8AF8E3BF1E04A4F6606D" lsidName="C. laetioides" pageId="0" pageNumber="97" rank="species" species="laetioides">
<emphasis id="26824BAAC79FDE401CA4C914D3247A21" italics="true" pageId="0" pageNumber="97">C. laetioides</emphasis>
</taxonomicName>
(=
<taxonomicName id="1C5AE449634EF6B172C2F36C12A4F7B7" authorityName="Britton &amp; Millspaugh" authorityYear="1920" baseAuthorityName="Swartz" class="Magnoliopsida" family="Salicaceae" genus="Zuelania" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Zuelania guidonia" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="guidonia">
<emphasis id="675A8535F4857820053424EF81D837EF" italics="true" pageId="0" pageNumber="97">Zuelania guidonia</emphasis>
</taxonomicName>
(Sw.) Britton &amp; Millsp),
<taxonomicName id="5D473279AF5B1F346DB17881C4D26DB8" lsidName="C. sanchezii" pageId="0" pageNumber="97" rank="species" species="sanchezii">
<emphasis id="445F8104AD254CDAA7F68E2644F1D42F" italics="true" pageId="0" pageNumber="97">C. sanchezii</emphasis>
</taxonomicName>
and
<taxonomicName id="CBBA50004A63BB681A6FE0FEF2BF209E" lsidName="C. tremula" pageId="0" pageNumber="97" rank="species" species="tremula">
<emphasis id="E2516499F192A9A8FD3FC35429CE15C5" italics="true" pageId="0" pageNumber="97">C. tremula</emphasis>
</taxonomicName>
. From both the plastid and nuclear trees,
<taxonomicName id="B4B5482BA4BF10DEA9B016CDD36BB219" lsidName="C. laetioides" pageId="0" pageNumber="97" rank="species" species="laetioides">
<emphasis id="C1ABC80B235C44767A6AF15C22DA4A12" italics="true" pageId="0" pageNumber="97">C. laetioides</emphasis>
</taxonomicName>
can be hypothesized as sister species to the newly discovered
<taxonomicName id="61933031B71D415603529864F27C6AF4" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="641BC6FFA89FA32554ECA17991369E56" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
. In terms of morphology, this new taxon differs from
<taxonomicName id="17C6256A29C9813DC7BCE80D422A17C0" lsidName="C. laetioides" pageId="0" pageNumber="97" rank="species" species="laetioides">
<emphasis id="36FE7290DC0B4BCF5A721853EB61D213" italics="true" pageId="0" pageNumber="97">C. laetioides</emphasis>
</taxonomicName>
(A. Rich.) Northr. by having essentially glabrous leaf blades (vs. pubescent), a prominent style (vs. absent or sometimes poorly developed), greenish-white sepals (vs. yellowish), 12 staminodia and stamens (vs. 15-20 staminodia and 20-40 stamens), and glabrous fruits (vs. pubescent). Furthermore,
<taxonomicName id="7D8D4C80E8F7B73306300C3E6A28DE6F" lsidName="C. sanchezii" pageId="0" pageNumber="97" rank="species" species="sanchezii">
<emphasis id="42D62449C9EE3E02DDFAA60F00E4C26E" italics="true" pageId="0" pageNumber="97">C. sanchezii</emphasis>
</taxonomicName>
differs morphologically from
<taxonomicName id="E3AE51A1A610048B282FA0E7D3908CF8" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="1FCA32153A3C350D39ED67625AFB4E8F" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
in presenting sessile inflorescence, crenate to subentire leaf blade and a longer petiole of
<quantity id="87207587AF0722C96EDDCA0990D78D6E" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1 to 2.5 cm</quantity>
.
<taxonomicName id="F433E9AFC5F6B6CD91C9F9C9F1CAE0FD" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="AF1B832F879F3337673BF7A13A3F1509" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4623D4773C53223976FDD5C2487B4867" lsidName="C. tremula" pageId="0" pageNumber="97" rank="species" species="tremula">
<emphasis id="19A18614C379D608297193D8B1518A34" italics="true" pageId="0" pageNumber="97">C. tremula</emphasis>
</taxonomicName>
in having congested inflorescences with 12-20 flowers (vs. loose inflorescences with 3 to 10 flowers), 12 stamens arranged in one row (vs. 15 to 24 stamens in two rows), and pubescent styles (vs. glabrous). The discriminating morphological characteristics between
<taxonomicName id="5639DFDCD3E3E54966132324AE689B0B" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="9344077AA6F3F3894FD42ED45C82E797" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
and its congeners were listed in Table
<tableCitation id="63736A3AD10800858B7FF63C36807442" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of the morphological characteristic of the newly described species and its closest relatives." httpUri="http://table.plazi.org/id/B331B1DCF5A492C402F52CF91C62AEF2" pageId="0" pageNumber="97" tableUuid="B331B1DCF5A492C402F52CF91C62AEF2">1</tableCitation>
.
<taxonomicName id="4B215D865C84B54CBAC4E8C14EDE1932" lsidName="C. sanchezii" pageId="0" pageNumber="97" rank="species" species="sanchezii">
<emphasis id="64A26C1063B9153006F355044D654D4B" italics="true" pageId="0" pageNumber="97">C. sanchezii</emphasis>
</taxonomicName>
is found in the cloud forest of
<collectingCountry id="5B4FBDA69ECBA912E9A497ABB63AB438" name="Mexico">Mexico</collectingCountry>
and
<collectingCountry id="C8A422859B1852D6963ED915FD489C80" name="El Salvador">El Salvador</collectingCountry>
at elevation of
<quantity id="AA8A7DDD04A6399EA3766C69526E7B78" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">1600 to 2000 m</quantity>
, whereas
<taxonomicName id="0E5B00C25C47A7EDEA8581DB6B35EF21" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="B7B5F0C1718FD3618F81C98EDF3E9BBB" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
is endemic to Central
<collectingCountry id="CBDC208E2E1007A04EC0D001D413E985" name="Panama">Panama</collectingCountry>
and occurs at
<specimenCount id="77B6829E5C9011C316840DCDBB092A62" type="generic">
<quantity id="3B928911B0E6BC14E2BB41DD8AC28AFD" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
</paragraph>
<caption id="FBF67152E0F184E1EB52FF5E0CA06AEA" doi="10.3897/phytokeys.236.108651.figure3" httpUri="https://binary.pensoft.net/fig/948452" pageId="0" pageNumber="97" start="Figure 3" startId="F3">
<paragraph id="C3AD034CC10ECC0D6CF7C154A7A22FBE" pageId="0" pageNumber="97">
<emphasis id="BB1D650055CA8893F4BE3B283A54B4A7" bold="true" pageId="0" pageNumber="97">Figure 3.</emphasis>
<taxonomicName id="B7645DF350151DA7A23D67A5AB08BD2C" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="08CFB65126A00126C0AED89DCD4B7404" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
de Mestier &amp; O. Ortiz
<emphasis id="C5101432E877B643BE839C86F797F0F8" bold="true" pageId="0" pageNumber="97">A</emphasis>
flowering branch
<emphasis id="EE0B6808400729655B476BDAA2A118F3" bold="true" pageId="0" pageNumber="97">B</emphasis>
inflorescence
<emphasis id="94EFF64F811742C8501CCF7EAE493E7E" bold="true" pageId="0" pageNumber="97">C</emphasis>
flowers at anthesis
<emphasis id="B04E2EA858AD8723FB71E8524EC23FBB" bold="true" pageId="0" pageNumber="97">D</emphasis>
fruiting branch
<emphasis id="68D43A180C54E146860CFB33BEA6073D" bold="true" pageId="0" pageNumber="97">E</emphasis>
leaves
<emphasis id="C5E8BF262C14CCC2356AABAA2D3A48B8" bold="true" pageId="0" pageNumber="97">F</emphasis>
ripe fruit
<emphasis id="DE7DAFD420D477BAF89D40B5A4063306" bold="true" pageId="0" pageNumber="97">G</emphasis>
bark. Photos
<emphasis id="EB0653FB7DD300A522459B3368EE8644" bold="true" pageId="0" pageNumber="97">A, C, E</emphasis>
(back side of the leaf) by Carmen Galdames (Galdames 6153). Photos
<emphasis id="7ACADEA482E30AF63B7725D31D39B73C" bold="true" pageId="0" pageNumber="97">D, E</emphasis>
(front side of the leaf) by Carmen Galdames (Galdames 6642). Photo B by Ernesto Campos (Sumich 151). Photos
<emphasis id="5C09415C6C808401BCBC6E7DBD2D0B79" bold="true" pageId="0" pageNumber="97">G, E</emphasis>
by Ernesto Campos Plate by Marco
<normalizedToken id="899CE31D4E13FE9A578D1C3B8FB79305" originalValue="Cedeño-Fonseca">Cedeno-Fonseca</normalizedToken>
.
</paragraph>
</caption>
<caption id="1EE1A19371BA81B38B02581D447BBC12" doi="10.3897/phytokeys.236.108651.figure4" httpUri="https://binary.pensoft.net/fig/948453" pageId="0" pageNumber="97" start="Figure 4" startId="F4">
<paragraph id="7BBD9CC0060D9785436849A9AF2DE3A4" pageId="0" pageNumber="97">
<emphasis id="5C80723A40E422AC0B7C87E86914C59C" bold="true" pageId="0" pageNumber="97">Figure 4.</emphasis>
<taxonomicName id="B88C6F4508025492F670C3E12EF102B2" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="AD071867C6F2F5B09E66B1BCC3465B31" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
de Mestier &amp; O. Ortiz
<emphasis id="9AB945DD0BBC67241007062496AD6778" bold="true" pageId="0" pageNumber="97">A</emphasis>
branches
<emphasis id="863D7FA7FFF8A09D5D3C49316B65C8B5" bold="true" pageId="0" pageNumber="97">B</emphasis>
twig and stipules
<emphasis id="34B9882F9E00A22BFAD865C9D19F0163" bold="true" pageId="0" pageNumber="97">C</emphasis>
trunk base and roots
<emphasis id="439E29A72BEB38E4E2760397EFAF208D" bold="true" pageId="0" pageNumber="97">D</emphasis>
lenticellate bark. Photos
<emphasis id="56F0306320209A04EC087E1A99D68925" bold="true" pageId="0" pageNumber="97">A, B</emphasis>
by Carmen Galdames (Galdames 6642). Photo
<emphasis id="C3910DAD12C3F87AD67CDA51ADA8FA45" bold="true" pageId="0" pageNumber="97">C</emphasis>
by Ernesto Campos (Campos 1087). Photo
<emphasis id="F4492009F4BC9480511A4928F6931BBD" bold="true" pageId="0" pageNumber="97">D</emphasis>
by Ernesto Campos. Plate by Marco
<normalizedToken id="9D0241283529F1439DEB414D0C58ED0F" originalValue="Cedeño-Fonseca">Cedeno-Fonseca</normalizedToken>
.
</paragraph>
</caption>
<caption id="B331B1DCF5A492C402F52CF91C62AEF2" ID-Table-UUID="B331B1DCF5A492C402F52CF91C62AEF2" httpUri="http://table.plazi.org/id/B331B1DCF5A492C402F52CF91C62AEF2" pageId="0" pageNumber="97" start="Table 1" startId="T1">
<paragraph id="5FCAF3A5AE3B243DC571C286CFF61C04" pageId="0" pageNumber="97">
<emphasis id="09B8230F874FB8D336A3366DB50B86EB" bold="true" pageId="0" pageNumber="97">Table 1.</emphasis>
Comparison of the morphological characteristic of the newly described species and its closest relatives.
</paragraph>
</caption>
<paragraph id="A7921A37E586E13ACA497A6A27783451" pageId="0" pageNumber="97">
<table id="B334DDE26FA5A0CBE045D2B672E5316C" pageId="0" pageNumber="97">
<tr id="7E4396EFAA2CFBEC7F8363B809404103" pageId="0" pageNumber="97">
<th id="C9162549C159CDEA53A40D49A3C04A0A" colspan="1" isEmpty="true" pageId="0" pageNumber="97" rowspan="1">-</th>
<th id="A0EECCCDB7EE089911DDDB65C73D6452" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<taxonomicName id="2A26B2453E1680666CE5E8FEA97F2EA2" lsidName="C. isthmica" pageId="0" pageNumber="97" rank="species" species="isthmica">
<emphasis id="D1CEFDA62E0B3BCD4DA874A2F6A88A04" italics="true" pageId="0" pageNumber="97">C. isthmica</emphasis>
</taxonomicName>
</th>
<th id="A94449C058C8FB9FB370A0DDD6DB79D6" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<taxonomicName id="9E8789A77B8993B1BDB5F9CB3BFD66ED" lsidName="C. laetioides" pageId="0" pageNumber="97" rank="species" species="laetioides">
<emphasis id="55A9F554F98118FF5626383498B3F372" italics="true" pageId="0" pageNumber="97">C. laetioides</emphasis>
</taxonomicName>
</th>
<th id="696B7E1B91222556E4CDCA0DC6A2D398" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<taxonomicName id="859AC53CD6BB591E2C27488D02FE8BA3" lsidName="C. sanchezii" pageId="0" pageNumber="97" rank="species" species="sanchezii">
<emphasis id="2DD9276DD4BBC9359053DF6209C92FA7" italics="true" pageId="0" pageNumber="97">C. sanchezii</emphasis>
</taxonomicName>
</th>
<th id="13A3A572DF351E8BABA007214C792BEA" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<taxonomicName id="2EC0473FCCA06422E22E801441633367" lsidName="C. tremula" pageId="0" pageNumber="97" rank="species" species="tremula">
<emphasis id="2126FB33052DC66D1460FAC3E62DFC43" italics="true" pageId="0" pageNumber="97">C. tremula</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="F5EB7088FEEB6C08A525DA67529D4DB0" pageId="0" pageNumber="97">
<td id="E9B1024BE991836E8B4F3DF731C83B43" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="11AA2818BF71E1FF27B3CA6653F52141" bold="true" pageId="0" pageNumber="97">Elevation (m</emphasis>
)
</td>
<td id="A8A78C291B9D26D2F101076F6BEF7E4F" colspan="1" pageId="0" pageNumber="97" rowspan="1">(600-)800-900</td>
<td id="1889E981603CCA8377C5CF4C96F4E85E" colspan="1" pageId="0" pageNumber="97" rowspan="1">0-500</td>
<td id="D76849D579345858054888F00159B6B5" colspan="1" pageId="0" pageNumber="97" rowspan="1">1600-2200</td>
<td id="3D328000FCE4F57CE5C42173948C7EAD" colspan="1" pageId="0" pageNumber="97" rowspan="1">0-600</td>
</tr>
<tr id="EFB0BE7B3A5C5014A171E9596CA016F7" pageId="0" pageNumber="97">
<td id="8837E57E6BF4C61E8150BA75AED0AF95" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="837328C99443D278BDCBD37B5D996EDD" bold="true" pageId="0" pageNumber="97">Habitat</emphasis>
</td>
<td id="6113F93A82C094729E88A6A0F266EE39" colspan="1" pageId="0" pageNumber="97" rowspan="1">cloud forests</td>
<td id="1645B2B41C9894CCA3FD70C7D7E3E377" colspan="1" pageId="0" pageNumber="97" rowspan="1">wet or dry lowland forests</td>
<td id="9B3F6AF04D80C1028CB2CF225B713DD2" colspan="1" pageId="0" pageNumber="97" rowspan="1">cloud forests</td>
<td id="695A6246F396F44BA935B1444F640014" colspan="1" pageId="0" pageNumber="97" rowspan="1">wet or dry lowland forests</td>
</tr>
<tr id="345E0388B0904BA97B2FF60562E72B73" pageId="0" pageNumber="97">
<td id="071D146FC9C7F0E80D27B9C627C3210E" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="3077B44827EEFE85C9CADCE5D965FA2A" bold="true" pageId="0" pageNumber="97">Petiole</emphasis>
</td>
<td id="1B560AD416749CB3DAE491703BD27989" colspan="1" pageId="0" pageNumber="97" rowspan="1">caniculate</td>
<td id="414C158799E77E144150CF745472628E" colspan="1" pageId="0" pageNumber="97" rowspan="1">non-caniculate</td>
<td id="E4E3D5A913D270DFE882676D51E43505" colspan="1" pageId="0" pageNumber="97" rowspan="1">caniculate</td>
<td id="25BA3DEB71F8CF6CF1880AB5B33386AA" colspan="1" pageId="0" pageNumber="97" rowspan="1">non-caniculate</td>
</tr>
<tr id="4E0CA28E55DB3978529709F9A22775E5" pageId="0" pageNumber="97">
<td id="EE391B2178E9259BB17ABA0B7F98FB4B" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="7867396C10DE88AE108C67A9D3E531CD" bold="true" pageId="0" pageNumber="97">Petiole size (cm)</emphasis>
</td>
<td id="9363B5231F10072D7F0582FD30FF07BA" colspan="1" pageId="0" pageNumber="97" rowspan="1">0.5-0.8</td>
<td id="494A305C64C6E2CE7EC133F716BC2CEC" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<quantity id="16BAFCE3E12F3A021F0DC31046E6714C" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 cm</quantity>
</td>
<td id="8B9D04FC2DD51CD33AD4D8C895417CD2" colspan="1" pageId="0" pageNumber="97" rowspan="1">
(0.4-)
<quantity id="D821224C497CE7E77C40FF306F764178" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1-2.5 cm</quantity>
</td>
<td id="43DE1F89421213DCFB8708143CCF9526" colspan="1" pageId="0" pageNumber="97" rowspan="1">(0.5-)1-1.8(-2.4)</td>
</tr>
<tr id="4CC7481115643A317D9B90B0E53DC47F" pageId="0" pageNumber="97">
<td id="3DBBDE21BD7F5FD2F3E998FAFAE4FE21" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="E709F8F14B9071C983B4701849D30A9E" bold="true" pageId="0" pageNumber="97">Leaf blade</emphasis>
</td>
<td id="BB1B1142E7697E7FFD54DBC5B37DBF3D" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
<td id="25AFD6A170F045156C04AD5BD652A040" colspan="1" pageId="0" pageNumber="97" rowspan="1">pubescent</td>
<td id="FFA695EF7BE1940FD902315371DDF466" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
<td id="E31CA379312C41BD473D9C6C3D5FFA57" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
</tr>
<tr id="20C88F03007597387E4CF221FA09A2D2" pageId="0" pageNumber="97">
<td id="7CD1847813819CEFB46F0036C546ECC1" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="A6A6203424A8D60188C2F37012B5C29E" bold="true" pageId="0" pageNumber="97">Leaf base</emphasis>
</td>
<td id="94832A23B0769DE807F2B9C740CB02B9" colspan="1" pageId="0" pageNumber="97" rowspan="1">asymmetrical, rounded to acute</td>
<td id="889F1F0E7A2A8A3BA71CDEAE6E302E55" colspan="1" pageId="0" pageNumber="97" rowspan="1">asymmetrical, obtuse to rotundate</td>
<td id="8B4C2BE801C4E88D399DCE36116615AD" colspan="1" pageId="0" pageNumber="97" rowspan="1">symmetrical, obtuse, cuneate or subcordate</td>
<td id="926542F5D9B6AE93E8017425D737B951" colspan="1" pageId="0" pageNumber="97" rowspan="1">asymmetrical, cuneate to rounded</td>
</tr>
<tr id="63EC3F03C17BF93C1757FC5577FA8374" pageId="0" pageNumber="97">
<td id="7EA89D3109AC854E019C11E786C4EE0C" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="F91F31E60D4D62B578FA5B21B8F7A4F0" bold="true" pageId="0" pageNumber="97">Leaf margins</emphasis>
</td>
<td id="545561F67B6081A6F10F1805C7D2354A" colspan="1" pageId="0" pageNumber="97" rowspan="1">serrate to crenate</td>
<td id="B507D527C3189F297287B9FAD8F9660B" colspan="1" pageId="0" pageNumber="97" rowspan="1">serrate</td>
<td id="D36D4B77341784CB8EAC03094E0E6733" colspan="1" pageId="0" pageNumber="97" rowspan="1">crenate to subentire</td>
<td id="36A83AF34BA1856EC1B6BB9CF6D72020" colspan="1" pageId="0" pageNumber="97" rowspan="1">serrate-crenulate</td>
</tr>
<tr id="FE691FF436BA9F8CF41055B0E4860740" pageId="0" pageNumber="97">
<td id="927CF604C87FB35BF3CFE4E3882517BE" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="0D3923D90345C9A1D10AAAEB593EAC1F" bold="true" pageId="0" pageNumber="97">Leaf apex</emphasis>
</td>
<td id="1B174118DA427E3D00E21843FD2D3EDC" colspan="1" pageId="0" pageNumber="97" rowspan="1">subcaudate to acuminate</td>
<td id="56D280D300DB3F940C578925CACB546F" colspan="1" pageId="0" pageNumber="97" rowspan="1">acuminate</td>
<td id="DEE537F073C53A7E5948371B91591C64" colspan="1" pageId="0" pageNumber="97" rowspan="1">acute to acuminate</td>
<td id="FE52EAF2E79E785A4D32CFCABCD2A960" colspan="1" pageId="0" pageNumber="97" rowspan="1">attenuate to subacuminate</td>
</tr>
<tr id="47F08AD16FBAEB65A286E1B54D12FA2E" pageId="0" pageNumber="97">
<td id="70B70CCF7F2DE3834ABCD81698807041" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="8AC9558ADD68AC01A0728F1C7E15BA52" bold="true" pageId="0" pageNumber="97">
Inflorescence
<typeStatus id="94A36814B69AC222EF08D54AE19C82CF">type</typeStatus>
</emphasis>
</td>
<td id="002E83AF46E0E118FE32A2217DFFABE3" colspan="1" pageId="0" pageNumber="97" rowspan="1">pedunculate, umbelliform</td>
<td id="D17C0300399E623007BE75A92B39F4EB" colspan="1" pageId="0" pageNumber="97" rowspan="1">sessile, fascicle</td>
<td id="29EF74B1DCB3A41914FBA7398BCF2EEE" colspan="1" pageId="0" pageNumber="97" rowspan="1">sessile, umbelliform</td>
<td id="4F98663608DB8CD536CBFD5F48AB3F38" colspan="1" pageId="0" pageNumber="97" rowspan="1">pedunculate, fascicle or corymb</td>
</tr>
<tr id="53B3040137E6F09C3F882A770120A012" pageId="0" pageNumber="97">
<td id="AC4C71E387A23E2A11D5B500B19BBD1B" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="0DBB37020C54E6963A0A209AE01A311D" bold="true" pageId="0" pageNumber="97">Flower number per inflorescence</emphasis>
</td>
<td id="6F9F3787661193347F6F02F02C645E0A" colspan="1" pageId="0" pageNumber="97" rowspan="1">12-20</td>
<td id="266C0A90EB784A43ABF9082FF51A89F9" colspan="1" pageId="0" pageNumber="97" rowspan="1">15</td>
<td id="5FAD1702C39B396642590AD6FE1E6654" colspan="1" pageId="0" pageNumber="97" rowspan="1">15-20</td>
<td id="1C81C637F828FB363B754AED67E5A887" colspan="1" pageId="0" pageNumber="97" rowspan="1">3-10</td>
</tr>
<tr id="FB0947B6E006D35D2DA98101D2AF25DB" pageId="0" pageNumber="97">
<td id="BCC9D56F420C8B0B716F01633BCBB666" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="7D242422368D4DC3F03D6494FE45FD21" bold="true" pageId="0" pageNumber="97">Stamens</emphasis>
</td>
<td id="F885AFE81DBD24BA4BB831E063C01597" colspan="1" pageId="0" pageNumber="97" rowspan="1">12</td>
<td id="19D4E111F848755AF495D9775D4B11C5" colspan="1" pageId="0" pageNumber="97" rowspan="1">20-40</td>
<td id="415099C90E9C359599CFE3AA4F6E250B" colspan="1" pageId="0" pageNumber="97" rowspan="1">12-15</td>
<td id="5CE899381AB45F2611ED9B0557D35360" colspan="1" pageId="0" pageNumber="97" rowspan="1">15-24</td>
</tr>
<tr id="F7A34502176488E481989F6C16317E10" pageId="0" pageNumber="97">
<td id="DACFD66F6F8273C00BB8C084CD6DB1C5" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="B8E29C978665C726C8D435F4A50EABA3" bold="true" pageId="0" pageNumber="97">Style</emphasis>
</td>
<td id="9F5B01597A8C2EC51C6BCC62EE7AA5A8" colspan="1" pageId="0" pageNumber="97" rowspan="1">present, pubescent</td>
<td id="9A0B42F7927B89FBC07EE62A949D9CAB" colspan="1" pageId="0" pageNumber="97" rowspan="1">absent</td>
<td id="078F60A1FB07E37308983F3147EC6CA3" colspan="1" pageId="0" pageNumber="97" rowspan="1">present, glabrous</td>
<td id="9BEC8C515DB675AAEEEB3B578BE65210" colspan="1" pageId="0" pageNumber="97" rowspan="1">present, glabrous</td>
</tr>
<tr id="F38690AF5CE017637766276A868199A9" pageId="0" pageNumber="97">
<td id="E5F6FDF132B50950D5D03CA2553289BC" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="7D6D9CE6875A44688D5566F5B66CDAD0" bold="true" pageId="0" pageNumber="97">Stamens in the same row as staminodia</emphasis>
</td>
<td id="94A39B9F11BA16B4710BC6E009D2677E" colspan="1" pageId="0" pageNumber="97" rowspan="1">yes</td>
<td id="A2912368BDD012940659A7710474158D" colspan="1" pageId="0" pageNumber="97" rowspan="1">yes</td>
<td id="28688C03E26912BB4B4D299DF9D45CD1" colspan="1" pageId="0" pageNumber="97" rowspan="1">yes</td>
<td id="8B802C5887BA5B634C48364545883A49" colspan="1" pageId="0" pageNumber="97" rowspan="1">no</td>
</tr>
<tr id="E244DD0CDF43AF834590606B34B6D086" pageId="0" pageNumber="97">
<td id="603F9DA9D4DA870046A04171C9C6705D" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="9AE1715C0FD471206E648179D8024756" bold="true" pageId="0" pageNumber="97">Sepals</emphasis>
</td>
<td id="D5957465D2A914BDE09A2E273FF97B78" colspan="1" pageId="0" pageNumber="97" rowspan="1">5</td>
<td id="E1455FB8E6902E0186123A6D88E90E52" colspan="1" pageId="0" pageNumber="97" rowspan="1">4-5</td>
<td id="E0662C8227E95FA282960221AE837B60" colspan="1" pageId="0" pageNumber="97" rowspan="1">5</td>
<td id="6E1C4D7A576BABF10C90747BBDC5A839" colspan="1" pageId="0" pageNumber="97" rowspan="1">(5-)6-9</td>
</tr>
<tr id="3C2800398B7AF027FBDB18C4C6C5671A" pageId="0" pageNumber="97">
<td id="38958D09FC1BCAC7F64678CCBC9E7EB8" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="B169E7DCEB59EC0D99AA9D0017F3E47D" bold="true" pageId="0" pageNumber="97">Sepal color</emphasis>
</td>
<td id="B9CC09581B1EEDAF64B22E1F73979886" colspan="1" pageId="0" pageNumber="97" rowspan="1">greenish-white</td>
<td id="4964833798D8756668A380F8E6635D1C" colspan="1" pageId="0" pageNumber="97" rowspan="1">yellow</td>
<td id="2BD54760A661941EFFD148B090515566" colspan="1" pageId="0" pageNumber="97" rowspan="1">white</td>
<td id="DBB4556F2F0609AEDD80EC6CAC49719F" colspan="1" pageId="0" pageNumber="97" rowspan="1">greenish-white</td>
</tr>
<tr id="D14E06A686AA536BCD6A856A7B4C7110" pageId="0" pageNumber="97">
<td id="A314FC692851321C62255AE688EB2C7E" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="929049C6C8ACA203A32349521C2CCDFB" bold="true" pageId="0" pageNumber="97">Fruit pubescence</emphasis>
</td>
<td id="C6344CB25854160A8404E2B2234A9519" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
<td id="7D6ACFF780D9254D90723E3F8C00C90C" colspan="1" pageId="0" pageNumber="97" rowspan="1">pubescent</td>
<td id="01A57AEFBEE0C087C0CF5D8CC53C843B" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
<td id="F47748A27ADE7326DD57C5D01A74172C" colspan="1" pageId="0" pageNumber="97" rowspan="1">glabrous</td>
</tr>
<tr id="DFD4BBCD69372A062E886C21B3FADB00" pageId="0" pageNumber="97">
<td id="9B08C9F109D7780E7AF75BAD4AFE6DBB" colspan="1" pageId="0" pageNumber="97" rowspan="1">
<emphasis id="2C1C0375567857407BDD8D4E0B277D20" bold="true" pageId="0" pageNumber="97">Fruit color</emphasis>
</td>
<td id="4A9B53698E3AF2ED03044001AEF576EB" colspan="1" pageId="0" pageNumber="97" rowspan="1">red pinkish</td>
<td id="8DF1CD8A94421E2405DDC21C9F43AF96" colspan="1" pageId="0" pageNumber="97" rowspan="1">yellowish-green</td>
<td id="9EEE65858672B782A2F50345C1AAD304" colspan="1" pageId="0" pageNumber="97" rowspan="1">red to dark red</td>
<td id="791216A3088A6202FFFC778FAC9ABCB8" colspan="1" pageId="0" pageNumber="97" rowspan="1">purple red</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="E5A4F7543F2FA877901ECC442B672780" pageId="0" pageNumber="97" type="description">
<paragraph id="DFDBF04B9DAAD7C7EB6FAF2B57A1FEA7" pageId="0" pageNumber="97">Description.</paragraph>
<paragraph id="7E79531D786B9A15E18E7169106D7D1A" pageId="0" pageNumber="97">
Tree, up to 1520 m tall; trunk straight, with bark cream, highly lenticellate; linear lenticels, arranged horizontally and vertically; branches zig-zag in shape, brownish, glabrous, with white lenticels. Stipules 4.0-6.8
<normalizedToken id="738F074B27B5D1B5656DFE1299C67DEC" originalValue="×">x</normalizedToken>
2.5-4 mm, glabrous, ovate or triangular, persistent on the upper part (distally) and then deciduous. Leaves alternate, simple, deciduous when flowering; petiole 0.5-0.8 cm, canaliculate, glabrous; leaf blade 3.5-8.8
<normalizedToken id="C00CC39372613CF6FAA70881D38A801A" originalValue="×">x</normalizedToken>
0.7-2.6 cm, subcoriaceous or coriaceous, drying brownish or blackish, lanceolate to obovate, asymmetrical, rounded or sometimes acute at the base, subcaudate-acuminate at the apex, densely pellucid punctate, glabrous on both sides, although slightly puberulous on the major abaxial veins; margins slightly serrate to crenate, marginal teeth more frequent in the upper half of the leaf blade; venation pinnate, lateral secondary veins in 5-10 ascending pairs, higher-order veins forming a dense reticulation, prominent on the abaxial surface. Inflorescences pedunculate, congested, umbeliform, each unit 12-20 flowered; peduncle 3 mm; bracts ca. 3.2 mm, coriaceous, ovate-lanceolate, greenish; pedicels ca. 8 mm, terete, basally articulate, green, glabrous. Floral buds oblong-obovate; flowers bisexual; sepals 5, ca. 4.0
<normalizedToken id="164230B1FFAB6F3F6A6582E4034BC445" originalValue="×">x</normalizedToken>
1.6 mm, seemly free, oblong-obovate, glabrescent, greenish-white, white internally and greenish externally; staminodia 12, white, ligulate, white-hirsute; stamens 12 on the same row as the smaller staminodia and alternating with them, densely pubescent; filaments equal, free, white-hirsute; anthers elliptic, creamy, white-hirsute externally; ovary greenish, ovate, white-hirsute; style undivided, white-hirsute; stigma capitate, creamy. Fruits fleshy, orbicular, up to 5 cm diameter, very glossy, green during development, turning black-purple externally and pinkish or red internally when ripe, covered on the lower part by the sepals, dehiscent by 3 valves; seeds many, creamy-white, aril orange.
</paragraph>
</subSubSection>
<subSubSection id="54AA588DC596A36ADADB9F4243C9B12A" pageId="0" pageNumber="97" type="distribution">
<paragraph id="F911A30994218E9EECCEBE5150DC6DB3" pageId="0" pageNumber="97">Distribution and habitat.</paragraph>
<paragraph id="874C3DF789575F763F59440CC5C92BEB" pageId="0" pageNumber="97">
Endemic to Central Panama (
<normalizedToken id="83C0855628D7EFE7579DF64B08ADABD4" originalValue="Panamá">Panama</normalizedToken>
and
<normalizedToken id="3A4FD9C381A1C965AF8F8F11A9F7D23F" originalValue="Panamá">Panama</normalizedToken>
Oeste provinces) (Fig.
<figureCitation id="7D5536CD23BCFD4D82677E9FDE9CE321" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Geographic distribution of Casearia isthmica." figureDoi="10.3897/phytokeys.236.108651.figure5" httpUri="https://binary.pensoft.net/fig/948454" pageId="0" pageNumber="97">5</figureCitation>
), documented only from cloud forests at elevations between (600-)800-900 m, in the premontane rain forest life zone.
</paragraph>
<caption id="4B6BCB59E93514F7A74A27118E2967E8" doi="10.3897/phytokeys.236.108651.figure5" httpUri="https://binary.pensoft.net/fig/948454" pageId="0" pageNumber="97" start="Figure 5" startId="F5">
<paragraph id="4B8FD92B99AD5FB93BCB9A7876566619" pageId="0" pageNumber="97">
<emphasis id="95DEC4EF15EB401BBB877DCA2A57E46B" bold="true" pageId="0" pageNumber="97">Figure 5.</emphasis>
Geographic distribution of
<taxonomicName id="1AF07DC4FE021095C95760F69582728B" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="60385D8AA0E52B50A7FF35E307654AC9" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="849EF356DC340961E50BFBF903B81CF5" pageId="0" pageNumber="97" type="phenology">
<paragraph id="45710B66B77021CB4BDD4BC49B4D7FC5" pageId="0" pageNumber="97">Phenology.</paragraph>
<paragraph id="CAADA6E032C52D50BAE6FC10D1E61BD2" pageId="0" pageNumber="97">Flowering in April, May. Fruiting in June, July.</paragraph>
</subSubSection>
<subSubSection id="E22D5B522F27FE7C8AF5ECD7258DF117" pageId="0" pageNumber="97" type="etymology">
<paragraph id="5D7F3A75020DAF668AD1C7C0912F335B" pageId="0" pageNumber="97">Etymology.</paragraph>
<paragraph id="AD315253C2033715AD2B45C204A9E933" pageId="0" pageNumber="97">The specific epithet refers to the geographic distribution of the new species, which is restricted to the Isthmus of Panama.</paragraph>
</subSubSection>
<subSubSection id="9BF73AAEEC622D3C5D6716602286B89D" pageId="0" pageNumber="97" type="preliminary conservation status">
<paragraph id="91DF2C5F680DD557ECF5BE4B79E04851" pageId="0" pageNumber="97">Preliminary conservation status.</paragraph>
<paragraph id="D50D43A0D83BE8682C7EEB8115FC9F28" pageId="0" pageNumber="97">
<taxonomicName id="58649CA682F0CA369728B62787347CDA" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="B2B8FA77C9887AD6CD16AD38089EF545" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
is known from ten collections made in two locations (Altos de Campana and Altos de Pacora). These collections were made within or very close to the external limits of protected areas (Altos de Campana National Park and Chagres National Park). Although they are protected areas, both locations face moderate anthropic disturbances, mainly in the borders such as Altos de Pacora (Chagres National Park). The highest threat facing this taxon is the loss of habitat caused by new road constructions and building of housing, as well as destructive tourist activities that are not environmentally sustainable, such as clandestine motorcycle races through the forest. Because this species has a limited distribution (EOO: 220 km2; AOO: 16 km2), the effect on its natural habitat may be extremely critical and may compromise its conservation.
<taxonomicName id="0828E9BA5727659E1770907D12C63505" authorityName="de Mestier &amp; O. Ortiz" authorityYear="2023" class="Magnoliopsida" family="Samydaceae" genus="Casearia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casearia isthmica" order="Malpighiales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis id="58FB70D7AF0E75E0AB146668AC772755" italics="true" pageId="0" pageNumber="97">Casearia isthmica</emphasis>
</taxonomicName>
must be considered as Endangered [EN B1ab(iii)+2ab(iii)].
</paragraph>
</subSubSection>
<subSubSection id="EE5176049B7AF03564D99F2334EB2A86" pageId="0" pageNumber="97" type="materials_examined">
<paragraph id="70DEF630173A389FA7AF2F8FBD548F05" pageId="0" pageNumber="97">Selected specimens examined.</paragraph>
<paragraph id="79A1F71D42DFE44B63F76C1F9A544C88" pageId="0" pageNumber="97">
<materialsCitation id="B3D4217342389FA775CA29DA4DBB8650" collectingDate="1992-05-21" collectingDateMax="2022-04-27" collectingDateMin="1992-05-21" collectorName="de Interpretacion, M. D. Correa, Sendero El Tigre, C. Galdames, H. Valdez, de Capira, de la Rana Dorada, E. Campos, La Rana Dorada, J. Sumich" country="Panama Panama" county="Parque Nacional Altos de Campana" elevation="850" latitude="8.692223" location="Camino Zamora" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" specimenCount="1" stateProvince="Oeste Province">
<emphasis id="2648E3817265797027D958982D4155DF" bold="true" pageId="0" pageNumber="97">
<collectingCountry id="26D1283FE24E82D0C5023E54CABD381B" name="Panama Panama">
Panama
<normalizedToken id="174447563D93656A40D68842CEEFBB5E" originalValue="Panamá">Panama</normalizedToken>
</collectingCountry>
<collectingRegion id="0C754ADC1932E8EFFE2410671F1C5D7B">Oeste Province</collectingRegion>
</emphasis>
:
<collectingCounty id="F9EBF8631C692ECB2C8A280161055F52">Parque Nacional Altos de Campana</collectingCounty>
;
<collectingMunicipality id="38350275044C102ED986E788B866E08E">
Sendero
<collectorName id="DEF9B7C4CB629D9D8B3DCEE08BF17E04">
de
<normalizedToken id="404169EE1CF063513B94BAF12A90BEFB" originalValue="Interpretación">Interpretacion</normalizedToken>
</collectorName>
</collectingMunicipality>
;
<quantity id="F5861D396F3DE3E296971189226EB0F2" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
del campamento de los guardaparques de INRENARE; bosque muy
<normalizedToken id="E3013768C692856A8A0C16D803895466" originalValue="húmedo">humedo</normalizedToken>
tropical premontano;
<location id="9AFB6B5F979FA4FB0D8CA9109BB3583F" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:9AFB6B5F979FA4FB0D8CA9109BB3583F" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Camino Zamora" stateProvince="Oeste Province">Camino Zamora</location>
;
<geoCoordinate id="837A55A6496CC1A19FF16132659C8686" degrees="8" direction="north" minutes="40" orientation="latitude" precision="925" value="8.666667">8°40'N</geoCoordinate>
,
<geoCoordinate id="FF2C6818CC5F7E8DE21467B70AA397AF" degrees="79" direction="west" minutes="55" orientation="longitude" precision="925" value="-79.916664">79°55'W</geoCoordinate>
;
<quantity id="B911968DA59CCD80C9DE85BC8C437238" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="BD84B6FCA83A0B0EC3A937543AFA6268" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
</quantity>
; fl;
<collectingDate id="C5CD4DD78766AE499EDAB315363B9FD4" value="1992-05-21">21 May 1992</collectingDate>
;
<emphasis id="C8C6B94F85A0742D985A27778D9DF84D" italics="true" pageId="0" pageNumber="97">
<collectorName id="4DF262B4AC803D0CB67418B855BBE6E5">M.D. Correa</collectorName>
et al. 8944
</emphasis>
(PMA) •
<location id="C318C6CBF6F33F578D63F38D4D58442D" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:C318C6CBF6F33F578D63F38D4D58442D" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
; colectado a
<quantity id="6A40F2BB81B0F9D9C5BA5142FD1C8F8D" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
<elevation id="4FFB84A4B8A1D8E2105FBD0BDDC95C5F" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</elevation>
</quantity>
de la orilla de la carretera dentro del parque;
<geoCoordinate id="8C2AB74DEBEBF7399BC7BAE6CF2D7799" degrees="8" direction="north" minutes="40" orientation="latitude" precision="925" value="8.666667">8°40'N</geoCoordinate>
,
<geoCoordinate id="2AD05526F3E0C9CE804E915F6DE3DA08" degrees="79" direction="west" minutes="55" orientation="longitude" precision="925" value="-79.916664">79°55'W</geoCoordinate>
;
<quantity id="FCBE0A2400D15B54D79BF1280763F932" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="CFA8B99CB56818A182337AB6B6CD8342" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
</quantity>
; fr;
<collectingDate id="96FA478B5F6FF04B0D9F1526C37A6C78" value="1993-06-24">24 June 1993</collectingDate>
;
<emphasis id="035123F2CC8575472C8675D0B9FAB857" italics="true" pageId="0" pageNumber="97">
<collectorName id="7F8BA90312A131393B65ED25287FB1C2">M.D. Correa</collectorName>
&amp; E. Montenegro 9622
</emphasis>
(PMA) •
<location id="4051314858ECFFDACDBF168306FAFDFF" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:4051314858ECFFDACDBF168306FAFDFF" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
; Finca
<normalizedToken id="6344B5A313381FE3C24461C81EB5D2D5" originalValue="García">Garcia</normalizedToken>
;
<geoCoordinate id="F31A4951F69650071D2B25BCDCDB089B" degrees="8" direction="north" minutes="40" orientation="latitude" precision="925" value="8.666667">8°40'N</geoCoordinate>
,
<geoCoordinate id="0B5FE342EC10CC4A27C75D967E9E09DE" degrees="79" direction="west" minutes="55" orientation="longitude" precision="925" value="-79.916664">79°55'W</geoCoordinate>
;
<quantity id="B2F8815F8BF5A166F0EBFFE1D367BE0E" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="4C0DEBF16C051AAF73F8CE1C640AAD04" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
</quantity>
; fr;
<collectingDate id="57D7B55613ADCD5156CFF089471809EF" value="1994-07-28">28 July 1994</collectingDate>
;
<emphasis id="8EFD7A146CE08A0AA2C31C0EBF743500" italics="true" pageId="0" pageNumber="97">
<collectorName id="7D4F0F2D318BFEE23941FFE73BCE0A84">M.D. Correa</collectorName>
&amp; E.
<collectingCountry id="556093425D8A3C8EC04E7BAA012D2A1A" name="Serbia">Montenegro</collectingCountry>
10717
</emphasis>
(PMA) •
<location id="A6FA8688FE41DD29DE3E2F6196A39B93" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:A6FA8688FE41DD29DE3E2F6196A39B93" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
;
<collectingDate id="B08C51776375700090C635961E66844F" value="1995-08-03">3 agosto 1995</collectingDate>
;
<emphasis id="1DD9CDDEF50F0473CB29E94515A433DC" italics="true" pageId="0" pageNumber="97">E. Montenegro 1112</emphasis>
(SCZ) •
<location id="69070FC0736D5B00821A4B27388BC0CF" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:69070FC0736D5B00821A4B27388BC0CF" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
; Las Nubes;
<normalizedToken id="E6FFF3B716164A512A8BFBCB49E3B5C1" originalValue="División">Division</normalizedToken>
Continental entre
<location id="EACDC40230D4921B6D47CD1B5386DE5B" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:EACDC40230D4921B6D47CD1B5386DE5B" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Finca de Tomas Herrera" stateProvince="Oeste Province">
Finca de
<normalizedToken id="3E5FCF5E9DAA7312EC008DF8A8A97698" originalValue="Tomás">Tomas</normalizedToken>
Herrera
</location>
;
<quantity id="EADBF95ACB5772EE3AC2DCA13AA04983" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<elevation id="73A073D1F2F59364D97E3E56575BD8CF" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
</quantity>
;
<collectingDate id="99A5690C96710CB9C451B6B688FD8B7A" value="1997-05-03">3 May 1997</collectingDate>
;
<emphasis id="08BA07A8B3C6C2215D000DFF74C0441D" italics="true" pageId="0" pageNumber="97">
<collectorName id="A214D0FBBD44591588C644532A1F0855">M.D. Correa</collectorName>
et al. 11430
</emphasis>
(F; MO PMA) •
<location id="60FEC090E15530D3C5DB5363297D82D4" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:60FEC090E15530D3C5DB5363297D82D4" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
;
<collectorName id="9656422D79ECD2DC383EE2C63787671E">Sendero El Tigre</collectorName>
; bosque nuboso;
<geoCoordinate id="021E6E9EC2B218E53468157AE27841EA" degrees="8" direction="north" minutes="40" orientation="latitude" precision="925" value="8.666667">8°40'N</geoCoordinate>
,
<geoCoordinate id="A580C7389BC126E5A2B0F03468F3F04C" degrees="79" direction="west" minutes="55" orientation="longitude" precision="925" value="-79.916664">79°55'W</geoCoordinate>
;
<quantity id="0303D9F77B8CB711605907E260C9E8F8" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation id="7099572B36329C2C4453A7861D140748" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
; fr;
<collectingDate id="B58FD003525626EBE3C4DD7359B5E36B" value="1998-07-11">11 Jul 1998</collectingDate>
;
<emphasis id="08B73CE1405C1EC4BFF717B867273011" italics="true" pageId="0" pageNumber="97">
<collectorName id="9A1D03B61547F266FF80B4825E23BB35">C. Galdames</collectorName>
</emphasis>
;
<emphasis id="F4E824A7093C2BC0FB61E88048A0C766" italics="true" pageId="0" pageNumber="97">
E.
<collectingCountry id="7B96F6B54F70166C9E96B5F1A34BF7E1" name="Serbia">Montenegro</collectingCountry>
&amp;
<collectorName id="83B31D50E4B909A10CCD5ED15D8E8157">
H.
<normalizedToken id="9A487BB27F2BDA1D17687A5BB02D176E" originalValue="Valdéz">Valdez</normalizedToken>
</collectorName>
4317
</emphasis>
(F; PMA; SCZ) •
<location id="B294E37F26719A504AE99AC5D0808664" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:B294E37F26719A504AE99AC5D0808664" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Distrito de Capira" stateProvince="Oeste Province">
Distrito
<collectorName id="C78B8F4CF1C6F68D0CA1AE728E19CF54">de Capira</collectorName>
</location>
;
<location id="DED26F37988740ADDB2E62F7E444F2CE" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:DED26F37988740ADDB2E62F7E444F2CE" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
; colectado en el sendero
<collectorName id="3331BCDE365D33CCB7D224B7E3AED412">de la Rana Dorada</collectorName>
; ca. A
<quantity id="6B9FF8A077CF6F811035AD0A678D19B5" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="D27D4F1DC2C0BDB1AE846B48B68E385B" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
de la entrada;
<geoCoordinate id="50A8FE27AD4A0858FD593FF514B00102" degrees="8" direction="north" minutes="41" orientation="latitude" precision="15" seconds="32" value="8.692223">8°41'32&quot;N</geoCoordinate>
,
<geoCoordinate id="EBE93D922A2237F863F02028EB01C0BC" degrees="79" direction="west" minutes="55" orientation="longitude" precision="15" seconds="34" value="-79.92611">79°55'34&quot;W</geoCoordinate>
;
<quantity id="FF19152B85E9EAA989E1E5F1657A2027" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">
<elevation id="73D6135B44DF93832BE923D36C565CB5" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">839 m</elevation>
</quantity>
; fl;
<collectingDate id="9750FB0A6CC804EE9D4EF4C9CF44D540" value="2018-05-03">3 May 2018</collectingDate>
;
<emphasis id="A2519341E7903496EB94FBBC9417C762" italics="true" pageId="0" pageNumber="97">
<collectorName id="C22623470976702E849CBFD25F79C7C3">E. Campos</collectorName>
&amp;
<collectorName id="18EC2E9A83CF3A38506FE82D2ADE6772">C. Galdames</collectorName>
1087
</emphasis>
(B; SCZ) •
<location id="447A519E00DAC46F873E0D1F532C9C5A" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:447A519E00DAC46F873E0D1F532C9C5A" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Distrito de Capira" stateProvince="Oeste Province">
Distrito
<collectorName id="BFBEA11013D32AF0F99714DDB945994A">de Capira</collectorName>
</location>
;
<location id="9CDC15F762564A26E4199E8775296A35" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:9CDC15F762564A26E4199E8775296A35" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
; sendero
<collectorName id="2BA0EE0D8EF2616CE921298399BD6139">La Rana Dorada</collectorName>
; ca.
<quantity id="0F0C7CC8E67D3A46B9C4A1539D650FAE" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="C0A58788CEE70989BE0A8EE03077EDFD" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
desde la entrada;
<geoCoordinate id="8A735D3720254E73D6C11F108AF226BB" degrees="8" direction="north" minutes="41" orientation="latitude" precision="15" seconds="32" value="8.692223">8°41'32&quot;N</geoCoordinate>
,
<geoCoordinate id="A53025D073E75BEC663BE6760CAA90F3" degrees="79" direction="west" minutes="55" orientation="longitude" precision="15" seconds="34" value="-79.92611">79°55'34&quot;W</geoCoordinate>
;
<quantity id="E6632CAA39010F01D8ADE7391A59D72D" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">
<elevation id="A49CBC99CD1F92CBF0F9A39DCE0B9181" metricMagnitude="2" metricUnit="m" metricValue="8.39" unit="m" value="839.0">839 m</elevation>
</quantity>
; fl;
<collectingDate id="9CC5221A2B2E416D9858249318E13E37" value="2022-04-27">27 April 2022</collectingDate>
;
<emphasis id="8903E58C9A26DC98D36C0533476F6BB5" italics="true" pageId="0" pageNumber="97">
<collectorName id="01DB1E5FCDFB0EE07E8BC82C149DDFD1">J. Sumich</collectorName>
&amp;
<collectorName id="C7014F55493349372827D2014C7DAF8B">E. Campos</collectorName>
151
</emphasis>
(SCZ) •
<location id="8A7CACCA5547D523F443F9A43E0D4F25" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:8A7CACCA5547D523F443F9A43E0D4F25" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Distrito de Capira" stateProvince="Oeste Province">
Distrito
<collectorName id="5809A2AD38495875B43B19D95D4C96C1">de Capira</collectorName>
</location>
;
<location id="40451164817F4C89B6CE7FDFD828EDB6" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:40451164817F4C89B6CE7FDFD828EDB6" country="Panama Panama" county="Parque Nacional Altos de Campana" latitude="8.692223" longLatPrecision="21" longitude="-79.92611" municipality="Sendero de Interpretacion" name="Parque Nacional Altos de Campana" stateProvince="Oeste Province">Parque Nacional Altos de Campana</location>
;
<collectingDate id="CFD632A8117287A20F1C22236EA79BE7" value="2022-04-27">27 April 2022</collectingDate>
; sendero
<collectorName id="21318181715839611044DC04DF4D2F1B">La Rana Dorada</collectorName>
;
<emphasis id="03139AD844B903A9DAE6322014023E58" italics="true" pageId="0" pageNumber="97">
<collectorName id="03AD68AEA1991E3C3CF8E6D94744D536">J. Sumich</collectorName>
127
</emphasis>
(SCZ)
</materialsCitation>
.
<materialsCitation id="56AA87C5A36491EB76F5EC6C39C69407" collectingDate="2010-06-29" collectorName="Sr. Rodrigo Coba, C. Galdames, R. Vergara, F. Rodriguez" country="Panama Panama" elevation="825" latitude="9.2080555" location="Cerro Jefe" longLatPrecision="21" longitude="-79.37583" municipality="Cerro Pelon" specimenCount="1" stateProvince="Panama Province">
<emphasis id="72F24856A5096A15440B0769FD96DFB4" bold="true" pageId="0" pageNumber="97">
<collectingRegion id="39198D23346E9E875A3E31BDAF172440">
<normalizedToken id="614E72BA3E0AE6EC91FF9B7AB1FF8C8D" originalValue="Panamá">Panama</normalizedToken>
Province
</collectingRegion>
</emphasis>
:
<collectingMunicipality id="4F0C51477885B4EBA141CFC0C16F9D70">
Cerro
<normalizedToken id="B86C777E88644392590C6247F1774CFC" originalValue="Pelón">Pelon</normalizedToken>
</collectingMunicipality>
; finca del
<collectorName id="E0E93EB7FDC611AE4F2272ACD6F2FABC">Sr. Rodrigo Coba</collectorName>
; reserva boscosa privada; adyacente a
<location id="652B710766C96B2493311DF8BC50F011" LSID="urn:lsid:plazi:treatment:724A0DAA225B5D1F9EF2BD8ECFA63B9B:652B710766C96B2493311DF8BC50F011" country="Panama Panama" latitude="9.2080555" longLatPrecision="21" longitude="-79.37583" municipality="Cerro Pelon" name="Cerro Jefe" stateProvince="Panama Province">Cerro Jefe</location>
;
<geoCoordinate id="6FBA15CD006DAA6058702AFECE4D015B" degrees="9" direction="north" minutes="12" orientation="latitude" precision="15" seconds="29" value="9.2080555">9°12'29&quot;N</geoCoordinate>
,
<geoCoordinate id="A9B7961F68B4C3F75088C9F98ECA2579" degrees="79" direction="west" minutes="22" orientation="longitude" precision="15" seconds="33" value="-79.37583">79°22'33&quot;W</geoCoordinate>
;
<quantity id="4E44F7C9609574A0A06936EF4E931AB8" metricMagnitude="2" metricUnit="m" metricValue="8.25" unit="m" value="825.0">
<elevation id="FB359BDAD1E8F1C2B97869FF04A5CC53" metricMagnitude="2" metricUnit="m" metricValue="8.25" unit="m" value="825.0">825 m</elevation>
</quantity>
; fr;
<collectingDate id="7758849CB8152DE2FEB84A59308A3F6E" value="2010-06-29">29 June 2010</collectingDate>
;
<emphasis id="7A044C88AAFED9415BB9F2342D5ADD7E" italics="true" pageId="0" pageNumber="97">
<collectorName id="F4FD51D89EEC607750BE41253E1ADE8D">C. Galdames</collectorName>
</emphasis>
;
<emphasis id="B3335D0BA541D9FB3DDF9E7A3C24238A" italics="true" pageId="0" pageNumber="97">
<collectorName id="946A82E1608CCDB71295348B59906A9B">R. Vergara</collectorName>
&amp;
<collectorName id="9A336CE2F25A9316622F87B03CE1ABE6">
F.
<normalizedToken id="01E492819060253BCC60A99B824A7475" originalValue="Rodríguez">Rodriguez</normalizedToken>
</collectorName>
6642
</emphasis>
(PMA; SCZ)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>