1131 lines
213 KiB
XML
1131 lines
213 KiB
XML
<document id="11CB0AB246F2F01050536242570C6835" ID-DOI="10.11646/zootaxa.5443.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10997142" ID-ZooBank="E206E545-7A72-4917-97B4-CDC85847DCBD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1713521610830" checkinUser="plazi" docAuthor="Pati, Sameer Kumar, Bajantri, Parashuram Prabhu & Hegde, Gopalkrishna Dattatraya" docDate="2024" docId="6E688161FFDDFFC051B8DEF510B0FCEE" docLanguage="en" docName="zootaxa.5443.1.6.pdf" docOrigin="Zootaxa 5443 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ghatiana dvirupa Pati & Bajantri & Hegde 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="125" masterDocId="9251F919FFDFFFC9512FDE621332FFB3" masterDocTitle="The “ Mr. and Mrs. Ghat crab ”, Ghatiana dvirupa sp. nov. (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India" masterLastPageNumber="126" masterPageNumber="116" pageNumber="118" updateTime="1713792921949" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="A8A4EFD392AD2C45D9600C421ADEB780">The “ Mr. and Mrs. Ghat crab ”, Ghatiana dvirupa sp. nov. (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India</mods:title>
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<mods:name id="FB5F6DB193E4217EB94B33BCCE4C656C" type="personal">
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<mods:roleTerm id="F2BBE7861D18494668CA637483AA6D06">Author</mods:roleTerm>
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<mods:namePart id="3DCC58E323F07B30A9CBFDCAC1EFE641">Pati, Sameer Kumar</mods:namePart>
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<mods:affiliation id="B8BD1EF4B1B658748145EA4C53F97269">Zoological Survey of India, Western Regional Centre, Vidyanagar, Sector 29, P. C. N. T. Post, Akurdi, Pune- 411 044, Maharashtra, India.</mods:affiliation>
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<mods:nameIdentifier id="72290FC7E9028956598A47C515E8A165" type="email">sameer_pati@yahoo.co.in</mods:nameIdentifier>
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<mods:namePart id="B76B59FE4FB159FCE984E8558E4FA520">Bajantri, Parashuram Prabhu</mods:namePart>
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<mods:affiliation id="1161C02E2950784BAE2913B5B2C932E9">Citizen Scientist, Karnataka Forest Department, Kali Tiger Reserve Division, Anshi Wildlife Range, Anshi- 581 187, Karnataka, India.</mods:affiliation>
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<mods:nameIdentifier id="8335817ABEC74747A99A23ED3E5B030E" type="email">parashurambajantrikfd@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3BF80DFD10F12711C139699DD1FB2D88">Hegde, Gopalkrishna Dattatraya</mods:namePart>
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<mods:affiliation id="7839107237DA49AE3759A7F345207790">Citizen Scientist, Honnepal, Bare Post- 581 337, Karnataka, India.</mods:affiliation>
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<mods:nameIdentifier id="31CEC3CC78C5855591048AB0D06E49E2" type="email">gopalkrishna.h@yahoo.com</mods:nameIdentifier>
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<mods:title id="CCF2F8EA6810AB9C94ADD7C42005EB8A">Zootaxa</mods:title>
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<mods:date id="3B94E57A028CE784908CAAD42D54119B">2024</mods:date>
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<mods:number id="C8002A42577D5A4EDDE6F749B20BBDF7">2024-04-19</mods:number>
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<treatment id="6E688161FFDDFFC051B8DEF510B0FCEE" LSID="urn:lsid:plazi:treatment:6E688161FFDDFFC051B8DEF510B0FCEE" httpUri="http://treatment.plazi.org/id/6E688161FFDDFFC051B8DEF510B0FCEE" lastPageId="9" lastPageNumber="125" pageId="2" pageNumber="118">
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<subSubSection id="AEDB63FCFFDDFFCB51B8DEF512E0FF02" box="[151,466,151,178]" pageId="2" pageNumber="118" type="nomenclature">
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<paragraph id="E67E3077FFDDFFCB51B8DEF512E0FF02" blockId="2.[151,569,151,250]" box="[151,466,151,178]" pageId="2" pageNumber="118">
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<heading id="BD36871BFFDDFFCB51B8DEF512E0FF02" bold="true" box="[151,466,151,178]" fontSize="11" level="1" pageId="2" pageNumber="118" reason="1">
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<emphasis id="D4B5EC65FFDDFFCB51B8DEF512E0FF02" bold="true" box="[151,466,151,178]" pageId="2" pageNumber="118">
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<taxonomicName id="21C14BF4FFDDFFCB51B8DEF5125CFF01" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[151,366,151,178]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="118" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
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<emphasis id="D4B5EC65FFDDFFCB51B8DEF5125CFF01" bold="true" box="[151,366,151,178]" italics="true" pageId="2" pageNumber="118">Ghatiana dvirupa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CF86511EFFDDFFCB505ADEF512E0FF02" box="[373,466,151,177]" pageId="2" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AEDB63FCFFDDFFCB51B8DEDE1285FF66" box="[151,439,187,214]" pageId="2" pageNumber="118" type="vernacular_names">
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<paragraph id="E67E3077FFDDFFCB51B8DEDE1285FF66" blockId="2.[151,569,151,250]" box="[151,439,187,214]" pageId="2" pageNumber="118">[Mr. and Mrs. Ghat Crab]</paragraph>
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</subSubSection>
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<subSubSection id="AEDB63FCFFDDFFCB51B8DEBD110BFF4A" box="[151,569,223,250]" pageId="2" pageNumber="118" type="description">
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<paragraph id="E67E3077FFDDFFCB51B8DEBD110BFF4A" blockId="2.[151,569,151,250]" box="[151,569,223,250]" pageId="2" pageNumber="118">
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(
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<figureCitation id="7EFA2CF2FFDDFFCB51B0DEBD1217FF4A" box="[159,293,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2A–C</figureCitation>
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,
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<figureCitation id="7EFA2CF2FFDDFFCB501FDEBD1244FF4A" box="[304,374,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">3A–H</figureCitation>
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,
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||
<figureCitation id="7EFA2CF2FFDDFFCB50ADDEBD12F1FF4A" box="[386,451,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4A–F</figureCitation>
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,
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<figureCitation id="7EFA2CF2FFDDFFCB50E1DEBD1103FF4A" box="[462,561,223,250]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="2" pageNumber="118">5A–D, F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="AEDB63FCFFDDFFCB51B8DF451272FD7E" pageId="2" pageNumber="118" type="materials_examined">
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<emphasis id="D4B5EC65FFDDFFCB51B8DF4512B2FEF2" bold="true" box="[151,384,295,321]" pageId="2" pageNumber="118">Material examined.</emphasis>
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<typeStatus id="397A8ED5FFDDFFCB50A8DF45112DFEF2" box="[391,543,295,321]" pageId="2" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
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:
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–
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<collectingRegion id="2405FE95FFDDFFCB53BDDF451075FEF2" box="[658,839,295,321]" country="India" name="Karnataka" pageId="2" pageNumber="118">Karnataka State</collectingRegion>
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: male adult (
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, CL
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, CH
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, FW
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),
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,
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<collectingMunicipality id="061AAA0DFFDDFFCB5091DF2911B5FED5" box="[446,647,331,358]" pageId="2" pageNumber="118">Castlerock Range</collectingMunicipality>
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,
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,
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(
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), alt.
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<emphasis id="D4B5EC65FFDDFFCB534CDF1211A6FE3A" box="[611,660,367,393]" italics="true" pageId="2" pageNumber="118">et al</emphasis>
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. leg. (
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<collectionCode id="80D0A8B2FFDDFFCB53F5DF0D107DFE3A" box="[730,847,367,393]" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
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C.2443)
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.
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<materialsCitation id="56A93A2AFFDDFFCB52EFDF0D119DFE1E" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="paratype">
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: male adult (
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<collectionCode id="80D0A8B2FFDDFFCB55C1DF0D162EFE3A" box="[1262,1308,367,393]" pageId="2" pageNumber="118">CW</collectionCode>
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, CL
|
||
<quantity id="21399D92FFDDFFCB51ECDFF1120BFE1D" box="[195,313,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="2.005" pageId="2" pageNumber="118" unit="mm" value="20.05">20.05 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB5069DFF1125FFE1E" box="[326,365,403,429]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB5059DFF112DEFE1D" box="[374,492,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="118" unit="mm" value="14.6">14.60 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB5303DFF11196FE1E" box="[556,676,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.072" pageId="2" pageNumber="118" unit="mm" value="10.72">10.72 mm</quantity>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="56A93A2AFFDDFFCB5397DFF1109DFE62" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
female adult (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5279DFF110B6FE1E" box="[854,900,403,429]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB52A2DFF11731FE1D" box="[909,1027,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="2.867" pageId="2" pageNumber="118" unit="mm" value="28.67">28.67 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB5513DFF11780FE1D" box="[1084,1202,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.907" pageId="2" pageNumber="118" unit="mm" value="19.07">19.07 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB55EFDFF117D5FE1E" box="[1216,1255,403,429]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB55C0DFF11657FE1D" box="[1263,1381,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.397" pageId="2" pageNumber="118" unit="mm" value="13.97">13.97 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB51B8DFD5123DFE62" box="[151,271,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.066" pageId="2" pageNumber="118" unit="mm" value="10.66">10.66 mm</quantity>
|
||
), same collection data as for holotype (
|
||
<collectionCode id="80D0A8B2FFDDFFCB53E1DFD51071FE62" box="[718,835,439,465]" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
|
||
C.2444);
|
||
</materialsCitation>
|
||
<materialsCitation id="56A93A2AFFDDFFCB5298DFD512DCFE46" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
male adult (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5511DFD5175EFE62" box="[1086,1132,439,465]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB555BDFD517D8FE61" box="[1140,1258,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="2.606" pageId="2" pageNumber="118" unit="mm" value="26.06">26.06 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB540DDFD516AAFE61" box="[1314,1432,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.759" pageId="2" pageNumber="118" unit="mm" value="17.59">17.59 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB51B8DFB9138CFE46" box="[151,190,475,501]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB51E9DFB91209FE45" box="[198,315,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.236" pageId="2" pageNumber="118" unit="mm" value="12.36">12.36 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB5056DFB912D0FE46" box="[377,482,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="9.87" pageId="2" pageNumber="118" unit="mm" value="9.87">9.87 mm</quantity>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="56A93A2AFFDDFFCB50D9DFB911E8FDA9" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
female adult (
|
||
<collectionCode id="80D0A8B2FFDDFFCB53BDDFB911F2FE46" box="[658,704,475,501]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB53E7DFB9100FFE45" box="[712,829,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="2.725" pageId="2" pageNumber="118" unit="mm" value="27.25">27.25 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB525BDFB910DBFE45" box="[884,1001,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.8539999999999999" pageId="2" pageNumber="118" unit="mm" value="18.54">18.54 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB52D9DFB9172FFE46" box="[1014,1053,475,501]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB550ADFB917A8FE45" box="[1061,1178,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.278" pageId="2" pageNumber="118" unit="mm" value="12.78">12.78 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB55F7DFB9167DFE46" box="[1240,1359,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.014" pageId="2" pageNumber="118" unit="mm" value="10.14">10.14 mm</quantity>
|
||
), same collection data as for holotype (
|
||
<collectionCode id="80D0A8B2FFDDFFCB50D5DF9D115DFDAA" box="[506,623,511,537]" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
|
||
C.2445);
|
||
</materialsCitation>
|
||
<materialsCitation id="56A93A2AFFDDFFCB53CEDF9D123EFD8E" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
male adult (
|
||
<collectionCode id="80D0A8B2FFDDFFCB524ADF9D10A1FDAA" box="[869,915,511,537]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB52B4DF9D173DFDA9" box="[923,1039,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="2.396" pageId="2" pageNumber="118" unit="mm" value="23.96">23.96 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB556ADF9D178BFDA9" box="[1093,1209,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="1.566" pageId="2" pageNumber="118" unit="mm" value="15.66">15.66 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB55EADF9D17DEFDAA" box="[1221,1260,511,537]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB55DBDF9D1655FDA9" box="[1268,1383,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="1.127" pageId="2" pageNumber="118" unit="mm" value="11.27">11.27 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB51B8DC411232FD8E" box="[151,256,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="9.29" pageId="2" pageNumber="118" unit="mm" value="9.29">9.29 mm</quantity>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="56A93A2AFFDDFFCB503BDC4112D3FDD2" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
female adult (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5080DC4112EFFD8E" box="[431,477,547,573]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB50CADC411168FD8D" box="[485,602,547,574]" metricMagnitude="-2" metricUnit="m" metricValue="2.312" pageId="2" pageNumber="118" unit="mm" value="23.12">23.12 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB53BEDC411034FD8D" box="[657,774,547,574]" metricMagnitude="-2" metricUnit="m" metricValue="1.581" pageId="2" pageNumber="118" unit="mm" value="15.81">15.81 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB523DDC41100BFD8E" box="[786,825,547,573]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB526EDC411086FD8D" box="[833,948,547,574]" metricMagnitude="-2" metricUnit="m" metricValue="1.115" pageId="2" pageNumber="118" unit="mm" value="11.15">11.15 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB52DDDC411769FD8E" box="[1010,1115,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="8.93" pageId="2" pageNumber="118" unit="mm" value="8.93">8.93 mm</quantity>
|
||
), same collection data as for holotype (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5028DC25124EFDD2" box="[263,380,583,609]" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
|
||
C.2446)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="56A93A2AFFDDFFCB50C5DC2517DAFD36" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="13" specimenCount-male="13" stateProvince="Karnataka">
|
||
<typeStatus id="397A8ED5FFDDFFCB50C5DC251002FDD2" box="[490,816,583,609]" isOtherTypeStatus="true" pageId="2" pageNumber="118">ADDITIONAL MATERIAL</typeStatus>
|
||
:
|
||
<specimenCount id="F0C7FBFEFFDDFFCB5215DC2510ACFDD2" box="[826,926,583,609]" count="13" pageId="2" pageNumber="118" type="male">13 males</specimenCount>
|
||
(
|
||
<collectionCode id="80D0A8B2FFDDFFCB5283DC2510E8FDD2" box="[940,986,583,609]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB52CEDC251790FDD1" box="[993,1186,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.9929999999999999" metricValueMax="2.376" metricValueMin="1.61" pageId="2" pageNumber="118" unit="mm" value="19.93" valueMax="23.76" valueMin="16.1">16.10–23.76 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB55F8DC2516AAFDD1" box="[1239,1432,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.2615" metricValueMax="1.508" metricValueMin="1.015" pageId="2" pageNumber="118" unit="mm" value="12.615" valueMax="15.08" valueMin="10.15">10.15–15.08 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB51B8DC09138CFD36" box="[151,190,619,645]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB51EADC09124AFD35" box="[197,376,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="8.719999999999999" metricValueMax="10.6" metricValueMin="6.84" pageId="2" pageNumber="118" unit="mm" value="8.719999999999999" valueMax="10.6" valueMin="6.84">6.84–10.60 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB509CDC091168FD36" box="[435,602,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="7.43" metricValueMax="8.78" metricValueMin="6.08" pageId="2" pageNumber="118" unit="mm" value="7.43" valueMax="8.78" valueMin="6.08">6.08–8.78 mm</quantity>
|
||
), same collection data as for holotype (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5521DC0917B1FD36" box="[1038,1155,619,645]" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
|
||
C.2447)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="56A93A2AFFDDFFCB55DCDC09120EFD7E" collectingDate="2023-08-03" collectionCode="CW, ZSI-WRC" collectorName="Md. Jafer Palot" country="Switzerland" county="Atle" elevation="746" latitude="15.419" location="Kali Tiger Reserve" longLatPrecision="76" longitude="74.288" municipality="Castlerock Range" pageId="2" pageNumber="118" specimenCount="8" specimenCount-female="8" stateProvince="Karnataka">
|
||
<specimenCount id="F0C7FBFEFFDDFFCB55DCDC09166DFD36" box="[1267,1375,619,645]" count="8" pageId="2" pageNumber="118" type="female">8 females</specimenCount>
|
||
(
|
||
<collectionCode id="80D0A8B2FFDDFFCB5441DC0916AEFD36" box="[1390,1436,619,645]" pageId="2" pageNumber="118">CW</collectionCode>
|
||
<quantity id="21399D92FFDDFFCB51B8DCED1268FD19" box="[151,346,655,682]" metricMagnitude="-2" metricUnit="m" metricValue="1.939" metricValueMax="2.48" metricValueMin="1.398" pageId="2" pageNumber="118" unit="mm" value="19.39" valueMax="24.8" valueMin="13.98">13.98–24.80 mm</quantity>
|
||
, CL
|
||
<quantity id="21399D92FFDDFFCB50BDDCED1175FD19" box="[402,583,655,682]" metricMagnitude="-2" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.573" metricValueMin="0.917" pageId="2" pageNumber="118" unit="mm" value="12.45" valueMax="15.73" valueMin="9.17">9.17–15.73 mm</quantity>
|
||
,
|
||
<collectingCountry id="9ED670E7FFDDFFCB537BDCED1149FD1A" box="[596,635,655,681]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
<quantity id="21399D92FFDDFFCB53ACDCED1005FD19" box="[643,823,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="8.785" metricValueMax="11.45" metricValueMin="6.12" pageId="2" pageNumber="118" unit="mm" value="8.785" valueMax="11.45" valueMin="6.12">6.12–11.45 mm</quantity>
|
||
, FW
|
||
<quantity id="21399D92FFDDFFCB5259DCED172CFD1A" box="[886,1054,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="7.115" metricValueMax="8.92" metricValueMin="5.31" pageId="2" pageNumber="118" unit="mm" value="7.115" valueMax="8.92" valueMin="5.31">5.31–8.92 mm</quantity>
|
||
), same data as for holotype (
|
||
<collectionCode id="80D0A8B2FFDDFFCB5448DCED13E5FD7E" pageId="2" pageNumber="118">ZSI-WRC</collectionCode>
|
||
C.2448)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFDDFFCB51E8DCB5161AFB86" pageId="2" pageNumber="118" type="diagnosis">
|
||
<paragraph id="E67E3077FFDDFFCB51E8DCB5161AFB86" blockId="2.[151,1437,295,2014]" pageId="2" pageNumber="118">
|
||
<emphasis id="D4B5EC65FFDDFFCB51E8DCB51270FD42" bold="true" box="[199,322,727,753]" pageId="2" pageNumber="118">Diagnosis.</emphasis>
|
||
Carapace in adult proportionately broad (CW/CL = 1.5–1.6), strongly arched (
|
||
<collectingCountry id="9ED670E7FFDDFFCB5583DCB517E1FD42" box="[1196,1235,727,753]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
/CL = 0.7); lateral margins strongly convex; frontal margin some distance from anterior margin of epistome, exposing antennular fossae (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB51C0DD7D1245FC89" box="[239,375,799,826]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB50AADD7D12E2FC8A" box="[389,464,799,825]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4A, D</figureCitation>
|
||
). Eyes relatively small as compared to orbital space; each eye with relatively stouter eyestalk (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB502CDD211269FCEE" box="[259,347,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Fig. 2B</figureCitation>
|
||
). Maxillipeds 1, 2 each with long flagellum on exopod (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB52FCDD21171CFCEE" box="[979,1070,835,861]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">Fig. 3A</figureCitation>
|
||
). Major chela in adult male with palm relatively stouter; ventral margin of fixed finger and distal half of palm strongly concave (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB55F0DD051655FC32" box="[1247,1383,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB545ADD0516A5FC32" box="[1397,1431,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">3C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB51B8DDE913D3FC16" box="[151,225,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4A, D</figureCitation>
|
||
). Ambulatory legs relatively short (P3 length/CL = ca. 2.0–2.2) (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB52E6DDE9171DFC16" box="[969,1071,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5512DDE917B5FC16" box="[1085,1159,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4A, D</figureCitation>
|
||
). Male pleonal somite 6 subquadrate, slightly longer than broad (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5375DDCD118EFC7A" box="[602,700,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB53E7DDCD11DEFC7A" box="[712,748,943,969]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">3D</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB53D7DDCD102EFC7A" box="[760,796,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4B</figureCitation>
|
||
). G1 relatively slender, medially gently curved outwards; ultimate article relatively slender, distally gently curved inwards, relatively long, ca. 0.4 times length of penultimate article; penultimate article relatively slender (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB53B3DD951012FBA2" box="[668,800,1015,1041]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">Figs. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5202DD951060FBA2" box="[813,850,1015,1041]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">4C</figureCitation>
|
||
). Vulvae in adult relatively closely positioned (VD/ SW = ca. 0.2), relatively large (occupying ca. 0.5 times length of s6), positioned close to s5/s6 (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB55E9DA79162EFB86" box="[1222,1308,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="2" pageNumber="118">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFDDFFCF51E8DA5D1032FE62" lastPageId="6" lastPageNumber="122" pageId="2" pageNumber="118" type="description">
|
||
<paragraph id="E67E3077FFDDFFCB51E8DA5D11D1F9BA" blockId="2.[151,1437,295,2014]" pageId="2" pageNumber="118">
|
||
<emphasis id="D4B5EC65FFDDFFCB51E8DA5D1119FBEA" bold="true" box="[199,555,1087,1113]" pageId="2" pageNumber="118">
|
||
Description of male
|
||
<typeStatus id="397A8ED5FFDDFFCB5091DA5D1115FBEA" box="[446,551,1087,1113]" pageId="2" pageNumber="118" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Carapace transversely ovate, distinctly broad (CW/CL = 1.5), conspicuously arched (
|
||
<collectingCountry id="9ED670E7FFDDFFCB51DDDA01122BFBCE" box="[242,281,1123,1149]" name="Switzerland" pageId="2" pageNumber="118">CH</collectingCountry>
|
||
/CL = 0.7); dorsal surface smooth, glabrous, strongly convex in frontal view; lateral margins strongly convex; anterolateral margins entire; front broad (FW/CW = 0.4), strongly deflexed anteriorly, trapezoidal; frontal margin smooth, cristate, some distance from anterior margin of epistome, exposing antennular fossae; frontal medial triangle incomplete, with dorsal margin only, lateral margins indiscernible; epigastric cristae low, only visible as 2 broad, rugose protuberances; postorbital cristae indiscernible; external orbital angle indistinct, low; epibranchial tooth indiscernible; postorbital region shallow; branchial regions inflated, smooth; cervical grooves shallow, narrow, interrupted; mesogastric groove shallow, narrow, long, bifurcated posteriorly; H-shaped groove visible; subhepatic region rugose; suborbital region generally smooth; pterygostomial region anteriorly with distinct tubercles; supraorbital margin almost straight, cristate, smooth; suborbital margin concave, subcristate, almost smooth, continuous with supraorbital margin; epistome posterior margin with well-developed, broadly triangular medial tooth and gently concave lateral parts (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB53B2DBA9102BFA56" box="[669,793,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Fig. 2A–C</figureCitation>
|
||
). Eyes small as compared to orbital space; eyestalk short, stout; cornea moderately large, pigmented (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5351DB8D11E4F9BA" box="[638,726,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFDDFFCB51E8D87111BCF952" blockId="2.[151,1437,295,2014]" pageId="2" pageNumber="118">
|
||
Antennules short, folded in longitudinally broad fossae; antennae conspicuously short (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5585D8711631F99E" box="[1194,1283,1555,1581]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Fig. 2B</figureCitation>
|
||
). Mandibular palp with 2 articles; terminal article bilobed, anterior lobe stout, short, ca. 0.5 times length of posterior lobe (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5440D855138FF9C6" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">Fig. 3A</figureCitation>
|
||
). Maxillipeds 1, 2 each with long flagellum on exopod (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5263D839109BF9C6" box="[844,937,1627,1653]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">Fig. 3A</figureCitation>
|
||
). Maxilliped 3 cover most of buccal cavity when closed; ischium subrectangular, longer than broad, longitudinal groove indiscernible; merus subquadrate, slightly broader than long, sunken; exopod stout, distally narrow, reaching beyond anterolateral corner of ischium, completely lacking flagellum (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB50C0D8A51163F952" box="[495,593,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5372D8A511B2F952" box="[605,640,1735,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFDDFFCB51E8D88917FEF826" blockId="2.[151,1437,295,2014]" pageId="2" pageNumber="118">
|
||
Chelipeds smooth, glabrous, unequal, right chela conspicuously larger (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB52D7D8891744F8B6" box="[1016,1142,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Fig. 2A, C</figureCitation>
|
||
). Major chela with 3 or 4 low, blunt teeth on each finger, remaining teeth inconspicuous, fingertips pointed, distinct gape when fingers closed; dactylus curved, stout, longer than upper margin of palm; palm stout, as long as high, smooth, distal half of palm and ventral margin of fixed finger strongly concave; carpus smooth, gently inflated, with low, blunt inner distal tooth; merus smooth except for low tubercles on margins, lacking subterminal spine (
|
||
<figureCitation id="7EFA2CF2FFDDFFCB5524D91917BDF826" box="[1035,1167,1915,1941]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="2" pageNumber="118">Figs. 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDDFFCB55B4D919178DF826" box="[1179,1215,1915,1941]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="2" pageNumber="118">3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFDDFFCA51E8D9FD10F4FF66" blockId="2.[151,1437,295,2014]" lastBlockId="3.[151,1436,150,213]" lastPageId="3" lastPageNumber="119" pageId="2" pageNumber="118">
|
||
Ambulatory legs slender, shorter than major cheliped, P3 length/CL = ca. 2.2, margins with tuft of 2–6 short brown setae with yellowish tips; merus (P2–P5) elongated, lacking subdistal spine; propodus (P2–P5) with distinct, sharp chitinous spines on lower margins; dactylus (P2–P5) gently recurved, subequal in length to propodus, with distinct, sharp chitinous spines on upper and lower margins (
|
||
<figureCitation id="7EFA2CF2FFDCFFCA5210DED9108BFF66" box="[831,953,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="3" pageNumber="119">Fig. 2A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B2BE60FFFFDCFFCA51B8D9CE10ACF85B" ID-DOI="http://doi.org/10.5281/zenodo.10997146" ID-Zenodo-Dep="10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="3" pageNumber="119" startId="3.[151,250,1964,1988]" targetBox="[193,1394,237,1941]" targetPageId="3" targetType="figure">
|
||
<paragraph id="E67E3077FFDCFFCA51B8D9CE10ACF85B" blockId="3.[151,1437,1964,2024]" pageId="3" pageNumber="119">
|
||
<emphasis id="D4B5EC65FFDCFFCA51B8D9CE1227F877" bold="true" box="[151,277,1964,1988]" pageId="3" pageNumber="119">FIGURE 2.</emphasis>
|
||
<taxonomicName id="21C14BF4FFDCFFCA5031D9CE12E7F877" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[286,469,1964,1988]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="119" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFDCFFCA5031D9CE12E7F877" box="[286,469,1964,1988]" italics="true" pageId="3" pageNumber="119">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFDCFFCA50F1D9CE1103F877" bold="true" box="[478,561,1964,1988]" pageId="3" pageNumber="119">
|
||
<taxonomicNameLabel id="CF86511EFFDCFFCA50F1D9CE1103F877" box="[478,561,1964,1988]" pageId="3" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E67E3077FFDBFFCD51E8DEF51739FE3A" blockId="4.[151,1436,151,393]" pageId="4" pageNumber="120">
|
||
Thoracic sternites smooth, generally glabrous (
|
||
<figureCitation id="7EFA2CF2FFDBFFCD53C2DEF51078FF02" box="[749,842,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="4" pageNumber="120">Fig. 2C</figureCitation>
|
||
); s1–s4 completely fused forming single structure, lacking suture or groove (
|
||
<figureCitation id="7EFA2CF2FFDBFFCD50EDDED9112FFF66" box="[450,541,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="4" pageNumber="120">Fig. 2C</figureCitation>
|
||
); s4/s5 shallow, narrow, medially interrupted; s5/s6 shallow, narrow, medially interrupted, each half extending anteriorly to join s4/s5; s6/s7 shallow, narrow, medially joining with longitudinal groove on s6; s7/s8 shallow, narrow, medially interrupted by longitudinal groove on s7, lacking transverse ridge; s8 almost covered by pleon (
|
||
<figureCitation id="7EFA2CF2FFDBFFCD5090DF45112BFEF2" box="[447,537,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="4" pageNumber="120">Fig. 2C</figureCitation>
|
||
), narrowed medially, longitudinal medial groove indiscernible. Pleonal locking mechanism with prominent tubercle on distal quarter of s5. Sternopleonal cavity deep, conspicuously long, reaching beyond imaginary line joining bases of ischium of maxilliped 3 (
|
||
<figureCitation id="7EFA2CF2FFDBFFCD5242DF0D10FDFE3A" box="[877,975,367,393]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="4" pageNumber="120">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFDBFFCD52F4DF0D10CCFE3A" box="[987,1022,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="4" pageNumber="120">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B2BE60FFFFDBFFCD51B8D95917D3F828" ID-DOI="http://doi.org/10.5281/zenodo.10997148" ID-Zenodo-Dep="10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="4" pageNumber="120" startId="4.[151,250,1851,1875]" targetBox="[151,1435,417,1828]" targetPageId="4" targetType="figure">
|
||
<paragraph id="E67E3077FFDBFFCD51B8D95917D3F828" blockId="4.[151,1436,1851,1947]" pageId="4" pageNumber="120">
|
||
<emphasis id="D4B5EC65FFDBFFCD51B8D9591221F8E0" bold="true" box="[151,275,1851,1875]" pageId="4" pageNumber="120">FIGURE 3.</emphasis>
|
||
<taxonomicName id="21C14BF4FFDBFFCD5036D95912FCF8E0" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[281,462,1851,1875]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="120" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFDBFFCD5036D95912FCF8E0" box="[281,462,1851,1875]" italics="true" pageId="4" pageNumber="120">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFDBFFCD50FBD9591116F8E0" bold="true" box="[468,548,1851,1875]" pageId="4" pageNumber="120">
|
||
<taxonomicNameLabel id="CF86511EFFDBFFCD50FBD9591116F8E0" box="[468,548,1851,1875]" pageId="4" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B2BE60FFFFDAFFCC51B8D8D916A0F8A8" ID-DOI="http://doi.org/10.5281/zenodo.10997150" ID-Zenodo-Dep="10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="5" pageNumber="121" startId="5.[151,249,1723,1747]" targetBox="[151,1435,181,1699]" targetPageId="5" targetType="figure">
|
||
<paragraph id="E67E3077FFDAFFCC51B8D8D916A0F8A8" blockId="5.[151,1437,1723,1819]" pageId="5" pageNumber="121">
|
||
<emphasis id="D4B5EC65FFDAFFCC51B8D8D91226F960" bold="true" box="[151,276,1723,1747]" pageId="5" pageNumber="121">FIGURE 4.</emphasis>
|
||
<taxonomicName id="21C14BF4FFDAFFCC5032D8D912E1F960" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[285,467,1723,1747]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="121" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFDAFFCC5032D8D912E1F960" box="[285,467,1723,1747]" italics="true" pageId="5" pageNumber="121">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFDAFFCC50F3D8D9111DF960" bold="true" box="[476,559,1723,1747]" pageId="5" pageNumber="121">
|
||
<taxonomicNameLabel id="CF86511EFFDAFFCC50F3D8D9111DF960" box="[476,559,1723,1747]" pageId="5" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E67E3077FFDAFFCF51E8D90F11D8FF66" blockId="5.[151,1436,1901,2036]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="122" pageId="5" pageNumber="121">
|
||
Pleon narrow, T-shaped, with strongly concave lateral margins; pleonal somites 1–6 progressively longer; pleonal somites 1, 2 subrectangular, broadest; pleonal somites 3, 4 trapezoidal, with straight lateral margins; pleonal somite 5 subtrapezoidal, with concave lateral margins; pleonal somite 6 subquadrate, slightly longer than broad, medial length ca. 1.1 times proximal width, longest among pleonal somites, distinctly shorter than telson, with sinuous lateral margins (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5082DEF51122FF02" box="[429,528,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="6" pageNumber="122">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5332DEF51171FF02" box="[541,579,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="6" pageNumber="122">3D</figureCitation>
|
||
). Telson elongated, medial length ca. 1.3 times proximal width, with gently concave lateral margins, apex narrow (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5366DED91199FF66" box="[585,683,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="6" pageNumber="122">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5398DED911EFFF66" box="[695,733,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="6" pageNumber="122">3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD9FFCF51E8DEBD17E2FE3A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
G1 slender, medially gently curved outwards, short, tip reaching half-length of s
|
||
<quantity id="21399D92FFD9FFCF551ADEBC176DFF4A" box="[1077,1119,222,249]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="122" unit="in" value="5.0">5 in</quantity>
|
||
situ; ultimate article slender, conical, long, ca. 0.4 times length of penultimate article, distally gently curved inwards, tip blunt; penultimate article stouter than ultimate article, broad proximally, relatively narrow distally, outer margin sinuous, inner margin weakly sinuous; groove for G2 ventral in position (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF53C6DF291054FED6" box="[745,870,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="6" pageNumber="122">Fig. 3E–G</figureCitation>
|
||
). G2 distinctly shorter than G1, ca. 0.6 times G1 length; ultimate article conspicuously short, ca. 0.1 times length of penultimate article (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5546DF0D17F1FE3A" box="[1129,1219,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="6" pageNumber="122">Fig. 3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD9FFCF51E8DFF01032FE62" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
<emphasis id="D4B5EC65FFD9FFCF51E8DFF0125FFE1F" bold="true" box="[199,365,402,428]" pageId="6" pageNumber="122">Colour in life.</emphasis>
|
||
Male crabs have pink carapace, ambulatory legs and chelipeds with blanched fingers (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5411DFF1139EFE62" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5A, B</figureCitation>
|
||
). Female crabs are dark purplish brown (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5357DFD511C1FE62" box="[632,755,438,465]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFD9FFCF51E8DFB912CEFDD2" pageId="6" pageNumber="122" type="etymology">
|
||
<paragraph id="E67E3077FFD9FFCF51E8DFB916AFFDAA" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
<emphasis id="D4B5EC65FFD9FFCF51E8DFB9127CFE46" bold="true" box="[199,334,475,501]" pageId="6" pageNumber="122">Etymology.</emphasis>
|
||
The species name is derived from the Sanskrit meaning ‘having two forms’, alluding to the difference in colour between male and female crabs in life. The name is to be treated as a Latin noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD9FFCF51E8DC4112CEFDD2" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">The common name of the new species is proposed herein as “Mr. and Mrs. Ghat Crab” as males and females possess different colours in life.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFD9FFCF51E8DC081712FC16" pageId="6" pageNumber="122" type="biology_ecology">
|
||
<paragraph id="E67E3077FFD9FFCF51E8DC081712FC16" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
<emphasis id="D4B5EC65FFD9FFCF51E8DC0812BFFD36" bold="true" box="[199,397,618,645]" pageId="6" pageNumber="122">Ecological notes.</emphasis>
|
||
<taxonomicName id="21C14BF4FFD9FFCF50BCDC09116BFD36" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[403,601,619,645]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF50BCDC09116BFD36" box="[403,601,619,645]" italics="true" pageId="6" pageNumber="122">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF5370DC09118AFD36" bold="true" box="[607,696,619,645]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF5370DC09118AFD36" box="[607,696,619,645]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is found on a high mountain laterite plateau (approximately
|
||
<quantity id="21399D92FFD9FFCF547CDC0916B9FD36" box="[1363,1419,619,645]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="122" unit="km" value="3.0">3 km</quantity>
|
||
<superScript id="11B49D3FFFD9FFCF54A4DC0816A1FDCB" attach="left" box="[1419,1427,618,632]" fontSize="6" pageId="6" pageNumber="122">2</superScript>
|
||
) of the Central Western Ghats of
|
||
<collectingCountry id="9ED670E7FFD9FFCF5328DCED1173FD1A" box="[519,577,655,681]" name="India" pageId="6" pageNumber="122">India</collectingCountry>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD9FFCF537EDCED11A6FD1A" box="[593,660,655,681]" captionStart="FIGURE 1" captionStartId="1.[151,250,1589,1613]" captionTargetBox="[151,1436,378,1561]" captionTargetId="figure-106@1.[151,1436,378,1563]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the distribution of Ghatiana dvirupa sp. nov. A, Western Ghats; B, Kali Tiger Reserve; C, Atle (type locality). This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997144" httpUri="https://zenodo.org/record/10997144/files/figure.png" pageId="6" pageNumber="122">Fig. 1</figureCitation>
|
||
). The mountain plateau (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF529BDCED1738FD1A" box="[948,1034,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5E</figureCitation>
|
||
) is almost devoid of vegetation but surrounded by the dense forest of a protected area, i.e., the Kali Tiger Reserve. Crabs of this new species are most active during the monsoon (June-September) and are often seen during the daytime when rain halts momentarily. These semiterrestrial crabs inhabit the holes and crevices of the laterite boulders (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5512DC9917A3FCA6" box="[1085,1169,762,789]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5F</figureCitation>
|
||
) clearly away from the streams. Crabs of the new species can also be seen underneath stones and small laterite boulders. Although the new species is currently restricted to a small area with specific habitat requirements, the population of the species appears to be high within the area.
|
||
<taxonomicName id="21C14BF4FFD9FFCF5337DD0510E3FC32" authority="Pati & Thackeray, 2021" authorityName="Pati & Thackeray" authorityYear="2021" box="[536,977,871,897]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="D4B5EC65FFD9FFCF5337DD0511F0FC33" box="[536,706,871,897]" italics="true" pageId="6" pageNumber="122">Ghatiana rouxi</emphasis>
|
||
<bibRefCitation id="82504D86FFD9FFCF53E7DD0510E3FC32" author="Pati, S. K. & Thackeray, T." box="[712,977,871,897]" pageId="6" pageNumber="122" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, is also found on the same laterite plateau but generally dwells under the stones and small boulders adjacent to the streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFD9FFC051E8DDCD11ABFD42" lastPageId="9" lastPageNumber="125" pageId="6" pageNumber="122" type="discussion">
|
||
<paragraph id="E67E3077FFD9FFCF51E8DDCD121AFBCE" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
<emphasis id="D4B5EC65FFD9FFCF51E8DDCD1209FC7A" bold="true" box="[199,315,943,969]" pageId="6" pageNumber="122">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="397A8ED5FFD9FFCF5052DDCD12EFFC7A" box="[381,477,943,969]" pageId="6" pageNumber="122" type="paratype">paratype</typeStatus>
|
||
males (ZSI-WRC C.2444–2446) of
|
||
<taxonomicName id="21C14BF4FFD9FFCF52A3DDCD173BFC7A" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[908,1033,943,969]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF52A3DDCD173BFC7A" box="[908,1033,943,969]" italics="true" pageId="6" pageNumber="122">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF553ADDCD1741FC7A" bold="true" box="[1045,1139,943,969]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF553ADDCD1741FC7A" box="[1045,1139,943,969]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are adults and similar to the
|
||
<typeStatus id="397A8ED5FFD9FFCF51EADDB11215FC5E" box="[197,295,979,1005]" pageId="6" pageNumber="122" type="holotype">holotype</typeStatus>
|
||
in most of the diagnostic features, including the gently inwardly curved distal portion of the G1 ultimate article (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF507EDD9512FCFBA2" box="[337,462,1015,1041]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4A–C</figureCitation>
|
||
). The subadult males, however, have the cristate anterolateral margins of the carapace (versus entire anterolateral margins of the carapace in adult males), the visible epibranchial tooth (versus indistinct epibranchial tooth in adult males) and the rugose branchial regions (versus generally smooth branchial regions in adult males).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD9FFCF51E8DAE517FAF92A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
The female
|
||
<typeStatus id="397A8ED5FFD9FFCF5068DAE51280FB12" box="[327,434,1159,1185]" pageId="6" pageNumber="122" type="paratype">paratypes</typeStatus>
|
||
(C.2444–2446) of
|
||
<taxonomicName id="21C14BF4FFD9FFCF53AFDAE511C4FB12" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[640,758,1159,1185]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF53AFDAE511C4FB12" box="[640,758,1159,1185]" italics="true" pageId="6" pageNumber="122">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF53D4DAE51063FB12" bold="true" box="[763,849,1159,1185]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF53D4DAE51063FB12" box="[763,849,1159,1185]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are adults and share most of the non-sexual diagnostic character states, including the proportionately broader carapace (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF52A6DAC910D5FB76" box="[905,999,1195,1221]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4D</figureCitation>
|
||
), the noticeable distance between the frontal margin and the anterior margin of the epistome exposing the antennular fossae, the relatively small eyes, and the long flagellum on the exopod of each of the maxillipeds 1, 2. As in subadult males, adult and subadult females of the new species possess a relatively rugose carapace on the branchial regions, with the cristate anterolateral margins and the visible but low epibranchial tooth (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF53CEDB59100EFAE6" box="[737,828,1339,1365]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4D</figureCitation>
|
||
). The heterochely is less pronounced in adult females (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF51B0DB3D13CAFACA" box="[159,248,1375,1401]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4D</figureCitation>
|
||
) than in adult males (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF50DFDB3D1145FACA" box="[496,631,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="6" pageNumber="122">Figs. 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD9FFCF53ABDB3D1198FACA" box="[644,682,1375,1401]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">4D</figureCitation>
|
||
). More importantly, female crabs of the new species have a darker colour in life (purplish brown) (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF50D8DBE1115FFA2E" box="[503,621,1410,1437]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5C, D</figureCitation>
|
||
) in contrast to the pink colouration of live male crabs (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF55E3DBE11676FA2E" box="[1228,1348,1410,1437]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5A, B</figureCitation>
|
||
). While live colouration of crabs may help identify a species of
|
||
<taxonomicName id="21C14BF4FFD9FFCF522ADBC5105DFA72" authorityName="Pati & Sharma" authorityYear="2014" box="[773,879,1447,1473]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4B5EC65FFD9FFCF522ADBC5105DFA72" box="[773,879,1447,1473]" italics="true" pageId="6" pageNumber="122">Ghatiana</emphasis>
|
||
</taxonomicName>
|
||
[e.g.,
|
||
<taxonomicName id="21C14BF4FFD9FFCF5299DBC513FDFA56" authority="Pati, Thackeray & Pawar, 2023" authorityName="Pati, Thackeray & Pawar" authorityYear="2023" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="sanguinolenta">
|
||
<emphasis id="D4B5EC65FFD9FFCF5299DBC51745FA72" box="[950,1143,1447,1473]" italics="true" pageId="6" pageNumber="122">G. sanguinolenta</emphasis>
|
||
<bibRefCitation id="82504D86FFD9FFCF5552DBC513FDFA56" author="Pati, S. K. & Thackeray, T. & Pawar, S." pageId="6" pageNumber="122" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">Pati, Thackeray & Pawar, 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="82504D86FFD9FFCF5021DBA912FCFA56" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[270,462,1483,1509]" pageId="6" pageNumber="122" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD9FFCF506DDBA91246FA56" box="[322,372,1483,1509]" italics="true" pageId="6" pageNumber="122">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
)], it should not be considered a key character. In a species of
|
||
<taxonomicName id="21C14BF4FFD9FFCF55BEDBA917C9FA56" authorityName="Pati & Sharma" authorityYear="2014" box="[1169,1275,1483,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4B5EC65FFD9FFCF55BEDBA917C9FA56" box="[1169,1275,1483,1509]" italics="true" pageId="6" pageNumber="122">Ghatiana</emphasis>
|
||
</taxonomicName>
|
||
, the colour of live crabs may vary between males and females [e.g.,
|
||
<taxonomicName id="21C14BF4FFD9FFCF53DDDB8D1765F9BA" authority="Pati & Thackeray, 2018" authorityName="Pati & Thackeray" authorityYear="2018" box="[754,1111,1519,1545]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD9FFCF53DDDB8D1075F9BA" box="[754,839,1519,1545]" italics="true" pageId="6" pageNumber="122">G. botti</emphasis>
|
||
<bibRefCitation id="82504D86FFD9FFCF5261DB8D1765F9BA" author="Pati, S. K. & Thackeray, T." box="[846,1111,1519,1545]" pageId="6" pageNumber="122" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="82504D86FFD9FFCF55BEDB8D16AAF9BA" author="Pati, S. K. & Thackeray, T." box="[1169,1432,1519,1545]" pageId="6" pageNumber="122" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 2A, B)]. As in the present new species, the contrasting colour in life between males and females may lead to wrong identification. The colour of the crabs may also vary between wet and dry seasons [e.g.,
|
||
<taxonomicName id="21C14BF4FFD9FFCF55E4D855127CF9C6" authority="Pati & Sharma, 2014" authorityName="Pati & Sharma" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="aurantiaca">
|
||
<emphasis id="D4B5EC65FFD9FFCF55E4D855165BF9E3" box="[1227,1385,1591,1617]" italics="true" pageId="6" pageNumber="122">G. aurantiaca</emphasis>
|
||
<bibRefCitation id="82504D86FFD9FFCF545ED855127CF9C6" author="Pati, S. K. & Sharma, R. M." pageId="6" pageNumber="122" pagination="1279 - 1298" refId="ref6612" refString="Pati, S. K. & Sharma, R. M. (2014) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48, 1279 - 1298. https: // doi. org / 10.1080 / 00222933.2013.859315" type="journal article" year="2014">Pati & Sharma, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5072D83912E8F9C6" box="[349,474,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5G, H</figureCitation>
|
||
)]. Irrespective of gender and season, the difference in live colouration is also observed among the individuals of
|
||
<taxonomicName id="21C14BF4FFD9FFCF509AD81D1120F92B" authorityName="Pati & Thackeray" authorityYear="2021" box="[437,530,1663,1689]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="D4B5EC65FFD9FFCF509AD81D1120F92B" box="[437,530,1663,1689]" italics="true" pageId="6" pageNumber="122">G. rouxi</emphasis>
|
||
</taxonomicName>
|
||
, which varies from deep purple to purplish brown (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF557CD81D1788F92A" box="[1107,1210,1662,1689]" captionStart="FIGURE 5" captionStartId="7.[151,247,1775,1799]" captionTargetBox="[260,1328,181,1751]" captionTargetId="figure-20@7.[260,1328,181,1751]" captionTargetPageId="7" captionText="FIGURE 5. Ghatiana dvirupa sp. nov., colour in life:A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of Ghatiana dvirupa sp. nov.: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder. Ghatiana aurantiaca Pati & Sharma, 2014, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011). Ghatiana rouxi Pati & Thackeray, 2021, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="6" pageNumber="122">Fig. 5I, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD9FFCF51E8D8C1102FF86E" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="122">
|
||
In the adult females of
|
||
<taxonomicName id="21C14BF4FFD9FFCF50E8D8C11172F90E" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[455,576,1699,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF50E8D8C11172F90E" box="[455,576,1699,1725]" italics="true" pageId="6" pageNumber="122">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF5369D8C111ADF90E" bold="true" box="[582,671,1699,1725]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF5369D8C111ADF90E" box="[582,671,1699,1725]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the pleon and telson together form a broadly subtriangular structure that completely covers the sternopleonal cavity when closed, the remaining portion of the thoracic sternum being exposed (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF502BD889126CF8B6" box="[260,350,1771,1797]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4E</figureCitation>
|
||
). The pleonal somite 1 of adult females of the new species is the shortest; pleonal somites 2–5 are progressively longer; and pleonal somite 6 is the longest, distinctly broader than long, shorter than the telson, with the convex lateral margins (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5325D9511151F8FE" box="[522,611,1843,1869]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4E</figureCitation>
|
||
). The female telson in adults of
|
||
<taxonomicName id="21C14BF4FFD9FFCF52E1D951177AF8FE" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[974,1096,1843,1869]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF52E1D951177AF8FE" box="[974,1096,1843,1869]" italics="true" pageId="6" pageNumber="122">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF557FD9511798F8FE" bold="true" box="[1104,1194,1843,1869]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF557FD9511798F8FE" box="[1104,1194,1843,1869]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is triangular, broader than long, with the strongly concave lateral margins and a broad apex (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5298D935173DF8C2" box="[951,1039,1879,1905]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4E</figureCitation>
|
||
). The vulvae in adult females of
|
||
<taxonomicName id="21C14BF4FFD9FFCF54AED93513DCF826" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="122" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD9FFCF54AED93513DCF826" italics="true" pageId="6" pageNumber="122">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD9FFCF51DAD919127DF826" bold="true" box="[245,335,1915,1941]" pageId="6" pageNumber="122">
|
||
<taxonomicNameLabel id="CF86511EFFD9FFCF51DAD919127DF826" box="[245,335,1915,1941]" pageId="6" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are located close to each other (VD/SW = ca. 0.2) on s6; each vulva is subovate, large (occupying ca. 0.5 times the length of s6), positioned close to s5/s6 (not touching s5/s6), covered with a soft membranous operculum, and surrounded by a low, broad rim (
|
||
<figureCitation id="7EFA2CF2FFD9FFCF5394D9A11023F86E" box="[699,785,1987,2013]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="6" pageNumber="122">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B2BE60FFFFD8FFCE51B8D88D1200F86C" ID-DOI="http://doi.org/10.5281/zenodo.10997152" ID-Zenodo-Dep="10997152" httpUri="https://zenodo.org/record/10997152/files/figure.png" pageId="7" pageNumber="123" startId="7.[151,247,1775,1799]" targetBox="[260,1328,181,1751]" targetPageId="7" targetType="figure">
|
||
<paragraph id="E67E3077FFD8FFCE51B8D88D1200F86C" blockId="7.[151,1437,1775,2015]" pageId="7" pageNumber="123">
|
||
<emphasis id="D4B5EC65FFD8FFCE51B8D88D123CF8B4" bold="true" box="[151,270,1775,1799]" pageId="7" pageNumber="123">FIGURE 5.</emphasis>
|
||
<taxonomicName id="21C14BF4FFD8FFCE503CD88D12F3F8B4" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[275,449,1775,1799]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="123" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD8FFCE503CD88D12F3F8B4" box="[275,449,1775,1799]" italics="true" pageId="7" pageNumber="123">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD8FFCE50E8D88D1126F8B4" bold="true" box="[455,532,1775,1799]" pageId="7" pageNumber="123">
|
||
<taxonomicNameLabel id="CF86511EFFD8FFCE50E8D88D1126F8B4" box="[455,532,1775,1799]" pageId="7" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, colour in life: A, holotype male, ZSI-WRC C.2443; B, male, ZSI-WRC C.2447; C, paratype female, ZSI-WRC C.2444; D, paratype female, ZSI-WRC C.2446. Habitats of
|
||
<taxonomicName id="21C14BF4FFD8FFCE52B5D971177BF898" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[922,1097,1811,1835]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="123" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD8FFCE52B5D971177BF898" box="[922,1097,1811,1835]" italics="true" pageId="7" pageNumber="123">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD8FFCE5560D97117AFF898" bold="true" box="[1103,1181,1811,1835]" pageId="7" pageNumber="123">
|
||
<taxonomicNameLabel id="CF86511EFFD8FFCE5560D97117AFF898" box="[1103,1181,1811,1835]" pageId="7" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: E, general habitat at type locality; F, male, ZSI-WRC C.2447, in crevice of laterite boulder.
|
||
<taxonomicName id="21C14BF4FFD8FFCE5232D95517F3F8FC" authority="Pati & Sharma, 2014" authorityName="Pati & Sharma" authorityYear="2014" box="[797,1217,1847,1871]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="123" phylum="Arthropoda" rank="species" species="aurantiaca">
|
||
<emphasis id="D4B5EC65FFD8FFCE5232D95510DEF8FC" box="[797,1004,1847,1871]" italics="true" pageId="7" pageNumber="123">Ghatiana aurantiaca</emphasis>
|
||
<bibRefCitation id="82504D86FFD8FFCE52DED95517F3F8FC" author="Pati, S. K. & Sharma, R. M." box="[1009,1217,1847,1871]" pageId="7" pageNumber="123" pagination="1279 - 1298" refId="ref6612" refString="Pati, S. K. & Sharma, R. M. (2014) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48, 1279 - 1298. https: // doi. org / 10.1080 / 00222933.2013.859315" type="journal article" year="2014">Pati & Sharma, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, colour in life: G, crab in its natural habitat during wet season (31 August 2013); H, crab outside of its habitat during dry season (28 November 2011).
|
||
<taxonomicName id="21C14BF4FFD8FFCE51B8D91D112DF824" authority="Pati & Thackeray, 2021" authorityName="Pati & Thackeray" authorityYear="2021" box="[151,543,1919,1943]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="123" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="D4B5EC65FFD8FFCE51B8D91D121FF824" box="[151,301,1919,1943]" italics="true" pageId="7" pageNumber="123">Ghatiana rouxi</emphasis>
|
||
<bibRefCitation id="82504D86FFD8FFCE501BD91D112DF824" author="Pati, S. K. & Thackeray, T." box="[308,543,1919,1943]" pageId="7" pageNumber="123" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, colour in life. I, male, ZSI-WRC C.2177; J, male, ZSI-WRC C.2176. Figure G is credited to Satyen Mehta, which is modified from https://www.inaturalist.org/observations/51882255. This figure is available in colour at
|
||
<emphasis id="D4B5EC65FFD8FFCE51B8D9AA13DAF86C" box="[151,232,1992,2015]" italics="true" pageId="7" pageNumber="123">Zootaxa</emphasis>
|
||
online.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E67E3077FFD7FFC151E8DEF511A6FD36" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="124">
|
||
<taxonomicName id="21C14BF4FFD7FFC151E8DEF512A6FF02" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[199,404,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC151E8DEF512A6FF02" box="[199,404,151,177]" italics="true" pageId="8" pageNumber="124">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC1508FDEF512CCFF02" bold="true" box="[416,510,151,177]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC1508FDEF512CCFF02" box="[416,510,151,177]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely resembles
|
||
<taxonomicName id="21C14BF4FFD7FFC1520CDEF510DBFF02" authorityName="Pati, Thackeray & Pawar" authorityYear="2023" box="[803,1001,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="sanguinolenta">
|
||
<emphasis id="D4B5EC65FFD7FFC1520CDEF510DBFF02" box="[803,1001,151,177]" italics="true" pageId="8" pageNumber="124">G. sanguinolenta</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="2405FE95FFD7FFC15519DEF5179BFF02" box="[1078,1193,151,177]" country="India" name="Karnataka" pageId="8" pageNumber="124">Karnataka</collectingRegion>
|
||
in having a broader carapace (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15026DED9125EFF66" box="[265,364,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15056DED912F2FF66" box="[377,448,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC150D8DED91180FF66" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[503,690,187,213]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC15305DED91169FF66" box="[554,603,187,213]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: figs. 2A, 3A, C); the relatively smaller eyes (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15599DED9162BFF66" box="[1206,1305,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC1540ADED9165EFF66" box="[1317,1388,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC151B8DEBD1266FF4A" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[151,340,223,249]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC151E5DEBD13CEFF4A" box="[202,252,223,249]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2B); relatively stouter palm of the major chela in adult males, with the ventral margin of the distal half of the palm and the fixed finger strongly concave (
|
||
<figureCitation id="7EFA2CF2FFD7FFC1527FDF611098FEAE" box="[848,938,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Fig. 3C</figureCitation>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC152CDDF611792FEAE" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[994,1184,259,285]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC15539DF61177AFEAE" box="[1046,1096,259,285]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2F); the medially gently outwardly curved G1, with the relatively slenderer and longer ultimate article, ca. 0.4 times the length of the penultimate article (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15055DF2912CCFED6" box="[378,510,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15324DF29111CFED6" box="[523,558,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC15348DF291017FED6" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[615,805,331,357]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC153B4DF2911FFFED6" box="[667,717,331,357]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2H, I); and the strongly concave lateral margins of the female telson (
|
||
<figureCitation id="7EFA2CF2FFD7FFC1504CDF0D128AFE3A" box="[355,440,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">Fig. 4E</figureCitation>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC150C2DF0D1196FE3A" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[493,676,367,393]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC15331DF0D117DFE3A" box="[542,591,367,393]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 3D). The G1 ultimate article, however, is gently curved inwards distally in
|
||
<taxonomicName id="21C14BF4FFD7FFC1503FDFF112B8FE1E" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[272,394,403,429]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC1503FDFF112B8FE1E" box="[272,394,403,429]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC150BCDFF112DFFE1E" bold="true" box="[403,493,403,429]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC150BCDFF112DFFE1E" box="[403,493,403,429]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD7FFC150D1DFF111B0FE1E" box="[510,642,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC153A0DFF11183FE1E" box="[655,689,403,429]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
) (versus G1 ultimate article distally gently curved outwards in
|
||
<taxonomicName id="21C14BF4FFD7FFC154AEDFF11204FE62" authorityName="Pati, Thackeray & Pawar" authorityYear="2023" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="sanguinolenta">
|
||
<emphasis id="D4B5EC65FFD7FFC154AEDFF11204FE62" italics="true" pageId="8" pageNumber="124">G. sanguinolenta</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC15040DFD5111BFE62" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[367,553,439,465]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC1508DDFD512E1FE62" box="[418,467,439,465]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2H, I). Other features also differentiate them: the frontal margin is located some distance from the anterior margin of the epistome, exposing the antennular fossae in
|
||
<taxonomicName id="21C14BF4FFD7FFC155A8DFB91632FE46" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[1159,1280,475,501]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC155A8DFB91632FE46" box="[1159,1280,475,501]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC15428DFB91652FE46" bold="true" box="[1287,1376,475,501]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC15428DFB91652FE46" box="[1287,1376,475,501]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD7FFC15440DFB9138BFDAA" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Fig. 2B</figureCitation>
|
||
) (versus frontal margin relatively close to the anterior margin of the epistome, almost hiding the antennular fossae in
|
||
<taxonomicName id="21C14BF4FFD7FFC151D0DC41128DFD8E" authorityName="Pati, Thackeray & Pawar" authorityYear="2023" box="[255,447,547,573]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="sanguinolenta">
|
||
<emphasis id="D4B5EC65FFD7FFC151D0DC41128DFD8E" box="[255,447,547,573]" italics="true" pageId="8" pageNumber="124">G. sanguinolenta</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC150D9DC41119CFD8E" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[502,686,547,573]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC15307DC41116AFD8E" box="[552,600,547,573]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2B); maxillipeds 1, 2 each with a long flagellum on the exopod in
|
||
<taxonomicName id="21C14BF4FFD7FFC15199DC251203FDD2" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[182,305,583,609]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC15199DC251203FDD2" box="[182,305,583,609]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC15015DC2512A7FDD2" bold="true" box="[314,405,583,609]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC15015DC2512A7FDD2" box="[314,405,583,609]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD7FFC15089DC251132FDD2" box="[422,512,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Fig. 3A</figureCitation>
|
||
) (versus each of the maxillipeds 1, 2 with a short flagellum on the exopod in
|
||
<taxonomicName id="21C14BF4FFD7FFC154AEDC251204FD36" authorityName="Pati, Thackeray & Pawar" authorityYear="2023" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="sanguinolenta">
|
||
<emphasis id="D4B5EC65FFD7FFC154AEDC251204FD36" italics="true" pageId="8" pageNumber="124">G. sanguinolenta</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC15040DC091118FD36" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[367,554,619,645]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC1508DDC0912E1FD36" box="[418,467,619,645]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
: fig. 2D).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD7FFC151E8DCED120CFBA2" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="124">
|
||
<taxonomicName id="21C14BF4FFD7FFC151E8DCED1243FD1B" authorityName="Pati & Thackeray" authorityYear="2021" box="[199,369,655,681]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="D4B5EC65FFD7FFC151E8DCED1243FD1B" box="[199,369,655,681]" italics="true" pageId="8" pageNumber="124">Ghatiana rouxi</emphasis>
|
||
</taxonomicName>
|
||
was also collected from the
|
||
<typeStatus id="397A8ED5FFD7FFC15382DCED11EFFD1A" box="[685,733,655,681]" pageId="8" pageNumber="124">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="21C14BF4FFD7FFC15274DCED10E1FD1A" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[859,979,655,681]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC15274DCED10E1FD1A" box="[859,979,655,681]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC152F6DCED1700FD1A" bold="true" box="[985,1074,655,681]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC152F6DCED1700FD1A" box="[985,1074,655,681]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The new species,
|
||
<taxonomicName id="21C14BF4FFD7FFC155D3DCED1646FD1A" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[1276,1396,655,681]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC155D3DCED1646FD1A" box="[1276,1396,655,681]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC15454DCED13FAFD7E" bold="true" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC15454DCED13FAFD7E" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, can easily be differentiated from
|
||
<taxonomicName id="21C14BF4FFD7FFC15361DCD11199FD7F" authorityName="Pati & Thackeray" authorityYear="2021" box="[590,683,691,717]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="D4B5EC65FFD7FFC15361DCD11199FD7F" box="[590,683,691,717]" italics="true" pageId="8" pageNumber="124">G. rouxi</emphasis>
|
||
</taxonomicName>
|
||
mainly by its proportionately broader carapace, CW/CL = 1.5–1.6 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC151B0DCB51237FD42" box="[159,261,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC1503DDCB5126AFD42" box="[274,344,727,753]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
) (versus carapace proportionately narrower, CW/CL = 1.2–1.3; see
|
||
<bibRefCitation id="82504D86FFD7FFC15571DCB51658FD42" author="Pati, S. K. & Thackeray, T." box="[1118,1386,727,753]" pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 4A, B); the strongly convex lateral margins of the carapace (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15210DC991091FCA6" box="[831,931,763,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15280DC9910C6FCA6" box="[943,1012,763,789]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
) (versus carapace with gently convex lateral margins; see
|
||
<bibRefCitation id="82504D86FFD7FFC15056DD7D11B1FC8A" author="Pati, S. K. & Thackeray, T." box="[377,643,799,825]" pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 4A, B); the relatively small eyes (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15515DD7D17A3FC8A" box="[1082,1169,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Fig. 2B</figureCitation>
|
||
) (versus eyes relatively larger; see
|
||
<bibRefCitation id="82504D86FFD7FFC1503EDD21112AFCEE" author="Pati, S. K. & Thackeray, T." box="[273,536,835,861]" pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 4C); the relatively shorter ultimate article of the G1, ca. 0.4 times the length of the penultimate article (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15090DD05112DFC32" box="[447,543,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15304DD05117FFC32" box="[555,589,871,897]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
) (versus G1 ultimate article relatively longer, ca. 0.5 times the length of the penultimate article; see
|
||
<bibRefCitation id="82504D86FFD7FFC150B0DDE91196FC16" author="Pati, S. K. & Thackeray, T." box="[415,676,907,933]" pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 5D); the relatively slenderer penultimate article of the G1 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC1544BDDE913E9FC7A" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E, G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC151C9DDCD1235FC7A" box="[230,263,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
) (versus G1 penultimate article relatively stouter; see
|
||
<bibRefCitation id="82504D86FFD7FFC1524CDDCD1758FC7A" author="Pati, S. K. & Thackeray, T." box="[867,1130,943,969]" pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 5D, E); and the vulvae being located close to the s5/s6 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC150D0DDB11160FC5E" box="[511,594,979,1005]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">Fig. 4F</figureCitation>
|
||
) (versus vulvae located some distance from the s5/s6; see
|
||
<bibRefCitation id="82504D86FFD7FFC155F6DDB113E1FBA2" author="Pati, S. K. & Thackeray, T." pageId="8" pageNumber="124" pagination="627 - 647" refId="ref6761" refString="Pati, S. K. & Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati & Thackeray 2021</bibRefCitation>
|
||
: fig. 5H).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD7FFC151E8DA7917D6FA82" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="124">
|
||
Concerning other species of
|
||
<taxonomicName id="21C14BF4FFD7FFC1533ADA79114DFB86" authorityName="Pati & Sharma" authorityYear="2014" box="[533,639,1051,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4B5EC65FFD7FFC1533ADA79114DFB86" box="[533,639,1051,1077]" italics="true" pageId="8" pageNumber="124">Ghatiana</emphasis>
|
||
</taxonomicName>
|
||
occurring in
|
||
<collectingRegion id="2405FE95FFD7FFC1520FDA7910A1FB86" box="[800,915,1051,1077]" country="India" name="Karnataka" pageId="8" pageNumber="124">Karnataka</collectingRegion>
|
||
[
|
||
<taxonomicName id="21C14BF4FFD7FFC15288DA7913E0FBEA" authority="Pati, Thackeray & Khaire, 2016" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="atropurpurea">
|
||
<emphasis id="D4B5EC65FFD7FFC15288DA791753FB87" box="[935,1121,1051,1077]" italics="true" pageId="8" pageNumber="124">G. atropurpurea</emphasis>
|
||
<bibRefCitation id="82504D86FFD7FFC15543DA7913E0FBEA" author="Pati, S. K. & Thackeray, T. & Khaire, A." pageId="8" pageNumber="124" pagination="569 - 586" refId="ref7050" refString="Pati, S. K., Thackeray, T. & Khaire, A. (2016) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India. Zootaxa, 4083 (4), 569 - 586. https: // doi. org / 10.11646 / zootaxa. 4083.4.7" type="journal article" year="2016">Pati, Thackeray & Khaire, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="21C14BF4FFD7FFC151F2DA5D11EDFBEA" authority="Klaus, Fernandez & Yeo, 2014" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[221,735,1087,1113]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC151F2DA5D124EFBEA" box="[221,380,1087,1113]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
<bibRefCitation id="82504D86FFD7FFC150ACDA5D11EDFBEA" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[387,735,1087,1113]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">Klaus, Fernandez & Yeo, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="21C14BF4FFD7FFC15236DA5D165DFBEA" authority="Pati, Thackeray, Bajantri & Hegde, 2022" authorityName="Pati, Thackeray, Bajantri & Hegde" authorityYear="2022" box="[793,1391,1087,1113]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvivarna">
|
||
<emphasis id="D4B5EC65FFD7FFC15236DA5D1092FBEA" box="[793,928,1087,1113]" italics="true" pageId="8" pageNumber="124">G. dvivarna</emphasis>
|
||
<bibRefCitation id="82504D86FFD7FFC15289DA5D165DFBEA" author="Pati, S. K. & Thackeray, T. & Bajantri, P. P. & Hegde, G. D." box="[934,1391,1087,1113]" pageId="8" pageNumber="124" pagination="2022019" refId="ref6973" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. & Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">Pati, Thackeray, Bajantri & Hegde, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="82504D86FFD7FFC151B8DA011267FBCE" author="Pati, S. K. & Thackeray, T. & Pawar, S." box="[151,341,1123,1149]" pageId="8" pageNumber="124" pagination="372 - 378" refId="ref7132" refString="Pati, S. K., Thackeray, T. & Pawar, S. (2023 a) Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5353 (4), 372 - 378. https: // doi. org / 10.11646 / zootaxa. 5353.4.4" type="journal article" year="2023">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC151E5DA0113CEFBCE" box="[202,252,1123,1149]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2023a
|
||
</bibRefCitation>
|
||
)],
|
||
<taxonomicName id="21C14BF4FFD7FFC1505FDA0112D8FBCE" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[368,490,1123,1149]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC1505FDA0112D8FBCE" box="[368,490,1123,1149]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC150DDDA01117EFBCE" bold="true" box="[498,588,1123,1149]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC150DDDA01117EFBCE" box="[498,588,1123,1149]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from them by the relatively stouter palm of the major chela in adult males, with the ventral margin of the distal half of the palm and the fixed finger strongly concave (
|
||
<figureCitation id="7EFA2CF2FFD7FFC155C9DAE5160EFB12" box="[1254,1340,1159,1185]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Fig. 3C</figureCitation>
|
||
) (versus palm of the major chela in adult males relatively slenderer, with the ventral margin of the distal half of the palm and the fixed finger gently concave; see
|
||
<bibRefCitation id="82504D86FFD7FFC15301DAAD11C0FB5A" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[558,754,1231,1257]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC15358DAAD119AFB5A" box="[631,680,1231,1257]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 3C;
|
||
<bibRefCitation id="82504D86FFD7FFC15275DAAD173AFB5A" author="Pati, S. K. & Thackeray, T. & Khaire, A." box="[858,1032,1231,1257]" pageId="8" pageNumber="124" pagination="569 - 586" refId="ref7050" refString="Pati, S. K., Thackeray, T. & Khaire, A. (2016) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India. Zootaxa, 4083 (4), 569 - 586. https: // doi. org / 10.11646 / zootaxa. 4083.4.7" type="journal article" year="2016">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC152A2DAAD108CFB5A" box="[909,958,1231,1257]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
: fig. 3D; 2022a: fig. 4B) in addition to the relatively elongated male pleonal somite 6 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC153E3DA91101DFABE" box="[716,815,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15214DA911052FABE" box="[827,864,1267,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">3D</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15243DA9110BCFABE" box="[876,910,1267,1293]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4B</figureCitation>
|
||
) (versus male pleonal somite 6 subquadrate or slightly broader than long; see
|
||
<bibRefCitation id="82504D86FFD7FFC150DEDB751187FA82" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[497,693,1303,1329]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC15315DB751159FA82" box="[570,619,1303,1329]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 3C;
|
||
<bibRefCitation id="82504D86FFD7FFC15233DB7510FBFA82" author="Pati, S. K. & Thackeray, T. & Khaire, A." box="[796,969,1303,1329]" pageId="8" pageNumber="124" pagination="569 - 586" refId="ref7050" refString="Pati, S. K., Thackeray, T. & Khaire, A. (2016) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India. Zootaxa, 4083 (4), 569 - 586. https: // doi. org / 10.11646 / zootaxa. 4083.4.7" type="journal article" year="2016">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD7FFC15261DB7510B2FA82" box="[846,896,1303,1329]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
: fig. 3G; 2022a: fig. 4C).
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD7FFC151E8DB591193F8B6" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="124">
|
||
Among these
|
||
<taxonomicName id="21C14BF4FFD7FFC15047DB59111EFAE6" box="[360,556,1339,1365]" pageId="8" pageNumber="124">
|
||
<emphasis id="D4B5EC65FFD7FFC15047DB5912E0FAE6" box="[360,466,1339,1365]" italics="true" pageId="8" pageNumber="124">Ghatiana</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of
|
||
<collectingRegion id="2405FE95FFD7FFC15379DB5911FEFAE6" box="[598,716,1339,1365]" country="India" name="Karnataka" pageId="8" pageNumber="124">Karnataka</collectingRegion>
|
||
,
|
||
<taxonomicName id="21C14BF4FFD7FFC153F6DB591048FAE6" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[729,890,1339,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC153F6DB591048FAE6" box="[729,890,1339,1365]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
</taxonomicName>
|
||
is known from an area that is located some
|
||
<quantity id="21399D92FFD7FFC154AFDB581389FACA" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="124" unit="km" value="15.0">15 km</quantity>
|
||
north of the
|
||
<typeStatus id="397A8ED5FFD7FFC1507FDB3D12B2FACA" box="[336,384,1375,1401]" pageId="8" pageNumber="124">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="21C14BF4FFD7FFC1532BDB3D114CFACA" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[516,638,1375,1401]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC1532BDB3D114CFACA" box="[516,638,1375,1401]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC153A9DB3D11D3FACA" bold="true" box="[646,737,1375,1401]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC153A9DB3D11D3FACA" box="[646,737,1375,1401]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see
|
||
<bibRefCitation id="82504D86FFD7FFC15231DB3D10D7FACA" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[798,997,1375,1401]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC15247DB3D10A8FACA" box="[872,922,1375,1401]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="82504D86FFD7FFC152DEDB3D17CFFACA" author="Pati, S. K. & Thackeray, T." box="[1009,1277,1375,1401]" pageId="8" pageNumber="124" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
). As such it is possible that one may encounter both species in the same locality. In
|
||
<taxonomicName id="21C14BF4FFD7FFC152B3DBE11727FA2E" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[924,1045,1411,1437]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC152B3DBE11727FA2E" box="[924,1045,1411,1437]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC15533DBE11747FA2E" bold="true" box="[1052,1141,1411,1437]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC15533DBE11747FA2E" box="[1052,1141,1411,1437]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the carapace, however, is relatively less broad, CW/CL = 1.5–1.6 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15370DBC511F1FA72" box="[607,707,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC153E0DBC51026FA72" box="[719,788,1447,1473]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
) (versus carapace distinctly broader, CW/CL =
|
||
<quantity id="21399D92FFD7FFC15404DBC516AEFA72" box="[1323,1436,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="8" pageNumber="124" unit="in" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 in</quantity>
|
||
<taxonomicName id="21C14BF4FFD7FFC151B8DBA91206FA56" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[151,308,1483,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC151B8DBA91206FA56" box="[151,308,1483,1509]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC15043DBA91119FA56" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[364,555,1483,1509]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC1509CDBA912D1FA56" box="[435,483,1483,1509]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 3B); the eyes are relatively small as compared to the orbit (
|
||
<figureCitation id="7EFA2CF2FFD7FFC155C7DBA9160FFA56" box="[1256,1341,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Fig. 2B</figureCitation>
|
||
) (versus eyes relatively large in relation to the orbit in
|
||
<taxonomicName id="21C14BF4FFD7FFC15386DB8D107BF9BA" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[681,841,1519,1545]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC15386DB8D107BF9BA" box="[681,841,1519,1545]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC152A8DB8D1762F9BA" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[903,1104,1519,1545]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC152FEDB8D1736F9BA" box="[977,1028,1519,1545]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 3A); the G1 is medially gently curved outwards, with the relatively slenderer, distally gently curved inwards and relatively longer ultimate article, ca. 0.4 times the length of the penultimate article (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15202D8551083F9E2" box="[813,945,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15291D85510D2F9E2" box="[958,992,1591,1617]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
) (versus G1 medially distinctly curved outwards, with the relatively stouter, straight and relatively shorter ultimate article, ca. 0.3 times the length of the penultimate article in
|
||
<taxonomicName id="21C14BF4FFD7FFC150A2D81D1119F92A" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[397,555,1663,1689]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC150A2D81D1119F92A" box="[397,555,1663,1689]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC1534AD81D101BF92A" author="Klaus, S. & Fernandez, K. & Yeo, D. C. J." box="[613,809,1663,1689]" pageId="8" pageNumber="124" pagination="651 - 660" refId="ref6464" refString="Klaus, S., Fernandez, K. & Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoologica Scripta, 43, 651 - 660. https: // doi. org / 10.1111 / zsc. 12078" type="journal article" year="2014">
|
||
Klaus
|
||
<emphasis id="D4B5EC65FFD7FFC15381D81D11EDF92A" box="[686,735,1663,1689]" italics="true" pageId="8" pageNumber="124">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 3D, E); and the female telson has strongly concave lateral margins (
|
||
<figureCitation id="7EFA2CF2FFD7FFC1507DD8C1129BF90E" box="[338,425,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">Fig. 4E</figureCitation>
|
||
) (versus female telson with almost straight lateral margins in
|
||
<taxonomicName id="21C14BF4FFD7FFC1555CD8C11621F90E" authorityName="Klaus, Fernandez & Yeo" authorityYear="2014" box="[1139,1299,1699,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="basalticola">
|
||
<emphasis id="D4B5EC65FFD7FFC1555CD8C11621F90E" box="[1139,1299,1699,1725]" italics="true" pageId="8" pageNumber="124">G. basalticola</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="82504D86FFD7FFC1547DD8C11260F952" author="Pati, S. K. & Thackeray, T." pageId="8" pageNumber="124" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 5G). The additional differences in the major chela and pleonal somite 6 of adult males are already mentioned in the preceding paragraph.
|
||
</paragraph>
|
||
<paragraph id="E67E3077FFD7FFC051E8D96D11ABFD42" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,150,861]" lastPageId="9" lastPageNumber="125" pageId="8" pageNumber="124">
|
||
The strongly convex lateral margins of the carapace (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15204D96D10A3F89A" box="[811,913,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC152B0D96D10DAF89A" box="[927,1000,1807,1833]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4A, D</figureCitation>
|
||
), the long flagellum on the exopod of each of the maxillipeds 1, 2 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC150CED951110EF8FE" box="[481,572,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Fig. 3A</figureCitation>
|
||
), the elongated male pleonal somite 6 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC152D7D9511769F8FE" box="[1016,1115,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="8" pageNumber="124">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15547D95117BEF8FE" box="[1128,1164,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">3D</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC155B6D951178FF8FE" box="[1177,1213,1843,1869]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4B</figureCitation>
|
||
), the distally gently inwardly curved G1 ultimate article (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15319D935118BF8C2" box="[566,697,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC153EAD93511DAF8C2" box="[709,744,1879,1905]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
), the basally narrow G1 penultimate article (
|
||
<figureCitation id="7EFA2CF2FFD7FFC155F2D9351652F8C2" box="[1245,1376,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="8" pageNumber="124">Figs. 3E, G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD7FFC15444D93516BDF8C2" box="[1387,1423,1879,1905]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">4C</figureCitation>
|
||
), and the relatively closely positioned vulvae, VD/SW = ca. 0.2 (
|
||
<figureCitation id="7EFA2CF2FFD7FFC15246D919108CF826" box="[873,958,1915,1941]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="8" pageNumber="124">Fig. 4F</figureCitation>
|
||
) of
|
||
<taxonomicName id="21C14BF4FFD7FFC152C1D919175AF826" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[1006,1128,1915,1941]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD7FFC152C1D919175AF826" box="[1006,1128,1915,1941]" italics="true" pageId="8" pageNumber="124">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD7FFC1555ED91917FEF826" bold="true" box="[1137,1228,1915,1941]" pageId="8" pageNumber="124">
|
||
<taxonomicNameLabel id="CF86511EFFD7FFC1555ED91917FEF826" box="[1137,1228,1915,1941]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resemble those of
|
||
<taxonomicName id="21C14BF4FFD7FFC151B8D9FD13DFF80A" authorityName="Pati & Thackeray" authorityYear="2018" box="[151,237,1951,1977]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD7FFC151B8D9FD13DFF80A" box="[151,237,1951,1977]" italics="true" pageId="8" pageNumber="124">G. botti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="21C14BF4FFD7FFC15009D9FD1015F80A" authority="Pati, Thackeray & Khaire, 2016" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" box="[294,807,1951,1977]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="D4B5EC65FFD7FFC15009D9FD1284F80A" box="[294,438,1951,1977]" italics="true" pageId="8" pageNumber="124">G. splendida</emphasis>
|
||
<bibRefCitation id="82504D86FFD7FFC15091D9FD1015F80A" author="Pati, S. K. & Thackeray, T. & Khaire, A." box="[446,807,1951,1977]" pageId="8" pageNumber="124" pagination="569 - 586" refId="ref7050" refString="Pati, S. K., Thackeray, T. & Khaire, A. (2016) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India. Zootaxa, 4083 (4), 569 - 586. https: // doi. org / 10.11646 / zootaxa. 4083.4.7" type="journal article" year="2016">Pati, Thackeray & Khaire, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="82504D86FFD7FFC1524BD9FD175CF80A" author="Pati, S. K. & Thackeray, T." box="[868,1134,1951,1977]" pageId="8" pageNumber="124" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: figs. 6A, G, I–K, N, 10A, C–E, H). Both
|
||
<taxonomicName id="21C14BF4FFD7FFC15011D9A112A0F86E" authorityName="Pati & Thackeray" authorityYear="2018" box="[318,402,1987,2013]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD7FFC15011D9A112A0F86E" box="[318,402,1987,2013]" italics="true" pageId="8" pageNumber="124">G. botti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="21C14BF4FFD7FFC150E7D9A11164F86E" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" box="[456,598,1987,2013]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="D4B5EC65FFD7FFC150E7D9A11164F86E" box="[456,598,1987,2013]" italics="true" pageId="8" pageNumber="124">G. splendida</emphasis>
|
||
</taxonomicName>
|
||
are found in the Sindhudurg district of the neighbouring
|
||
<collectingRegion id="2405FE95FFD7FFC155FFD9A116AAF86E" box="[1232,1432,1987,2013]" country="India" name="Maharashtra" pageId="8" pageNumber="124">Maharashtra state</collectingRegion>
|
||
, approximately
|
||
<quantity id="21399D92FFD6FFC0506DDEF412EEFF02" box="[322,476,150,177]" metricMagnitude="4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="125" unit="km" value="75.0" valueMax="100.0" valueMin="50.0">50 to 100 km</quantity>
|
||
away from the
|
||
<typeStatus id="397A8ED5FFD6FFC053BDDEF511F0FF02" box="[658,706,151,177]" pageId="9" pageNumber="125">type</typeStatus>
|
||
locality of the present new species (see
|
||
<bibRefCitation id="82504D86FFD6FFC055BCDEF51674FF02" author="Pati, S. K. & Thackeray, T. & Khaire, A." box="[1171,1350,151,177]" pageId="9" pageNumber="125" pagination="569 - 586" refId="ref7050" refString="Pati, S. K., Thackeray, T. & Khaire, A. (2016) Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India. Zootaxa, 4083 (4), 569 - 586. https: // doi. org / 10.11646 / zootaxa. 4083.4.7" type="journal article" year="2016">
|
||
Pati
|
||
<emphasis id="D4B5EC65FFD6FFC055E8DEF517C8FF02" box="[1223,1274,151,177]" italics="true" pageId="9" pageNumber="125">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="82504D86FFD6FFC0547DDEF51261FF66" author="Pati, S. K. & Thackeray, T." pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
).
|
||
<taxonomicName id="21C14BF4FFD6FFC05049DED9111DFF66" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[358,559,187,213]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD6FFC05049DED9111DFF66" box="[358,559,187,213]" italics="true" pageId="9" pageNumber="125">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD6FFC05317DED911A1FF66" bold="true" box="[568,659,187,213]" pageId="9" pageNumber="125">
|
||
<taxonomicNameLabel id="CF86511EFFD6FFC05317DED911A1FF66" box="[568,659,187,213]" pageId="9" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is nevertheless differentiated from
|
||
<taxonomicName id="21C14BF4FFD6FFC05509DED9174EFF66" authorityName="Pati & Thackeray" authorityYear="2018" box="[1062,1148,187,213]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD6FFC05509DED9174EFF66" box="[1062,1148,187,213]" italics="true" pageId="9" pageNumber="125">G. botti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="21C14BF4FFD6FFC05599DED91674FF66" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" box="[1206,1350,187,213]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="D4B5EC65FFD6FFC05599DED91674FF66" box="[1206,1350,187,213]" italics="true" pageId="9" pageNumber="125">G. splendida</emphasis>
|
||
</taxonomicName>
|
||
mainly by the proportionately broader carapace, CW/CL = 1.5–1.6 (
|
||
<figureCitation id="7EFA2CF2FFD6FFC05278DEBD108CFF4A" box="[855,958,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="9" pageNumber="125">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD6FFC052E2DEBD1727FF4A" box="[973,1045,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">4A, D</figureCitation>
|
||
) (versus carapace proportionately narrower, CW/CL = 1.3–1.4; see
|
||
<bibRefCitation id="82504D86FFD6FFC0533FDF611028FEAE" author="Pati, S. K. & Thackeray, T." box="[528,794,259,285]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: figs. 6A, 10A); the relatively small eyes as compared to the orbits (
|
||
<figureCitation id="7EFA2CF2FFD6FFC0503CDF451258FEF2" box="[275,362,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1964,1988]" captionTargetBox="[193,1394,237,1941]" captionTargetId="figure-48@3.[193,1394,237,1941]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, habitus, dorsal view; B, frontal view of cephalothorax; C, habitus, ventral view. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10997146" httpUri="https://zenodo.org/record/10997146/files/figure.png" pageId="9" pageNumber="125">Fig. 2B</figureCitation>
|
||
) (versus eyes relatively large; see
|
||
<bibRefCitation id="82504D86FFD6FFC053D9DF451730FEF2" author="Pati, S. K. & Thackeray, T." box="[758,1026,295,321]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: figs. 6B, 10B); the medially gently outwardly curved G1 (
|
||
<figureCitation id="7EFA2CF2FFD6FFC050BCDF291127FED6" box="[403,533,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="9" pageNumber="125">Figs. 3E, G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD6FFC0530FDF291173FED6" box="[544,577,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">4C</figureCitation>
|
||
) (versus G1 almost straight; see
|
||
<bibRefCitation id="82504D86FFD6FFC05281DF291786FED6" author="Pati, S. K. & Thackeray, T." box="[942,1204,331,357]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: figs. 6I, K, 10D, E); and the strongly concave lateral margins of the female telson (
|
||
<figureCitation id="7EFA2CF2FFD6FFC05273DF0D1080FE3A" box="[860,946,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">Fig. 4E</figureCitation>
|
||
) (versus female telson with almost straight to gently concave lateral margins; see
|
||
<bibRefCitation id="82504D86FFD6FFC0536DDFF11078FE1E" author="Pati, S. K. & Thackeray, T." box="[578,842,403,429]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: figs. 6M, 10G). The relatively slenderer G1 ultimate article (
|
||
<figureCitation id="7EFA2CF2FFD6FFC051C2DFD51241FE62" box="[237,371,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="9" pageNumber="125">Figs. 3E, G</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD6FFC05050DFD51293FE62" box="[383,417,439,465]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">4C</figureCitation>
|
||
) and the relatively large vulvae (occupying ca. 0.5 times the length of the s6) (
|
||
<figureCitation id="7EFA2CF2FFD6FFC0540EDFD51647FE62" box="[1313,1397,439,465]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">Fig. 4F</figureCitation>
|
||
) of
|
||
<taxonomicName id="21C14BF4FFD6FFC051B8DFB91223FE46" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[151,273,475,501]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD6FFC051B8DFB91223FE46" box="[151,273,475,501]" italics="true" pageId="9" pageNumber="125">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD6FFC05037DFB91243FE46" bold="true" box="[280,369,475,501]" pageId="9" pageNumber="125">
|
||
<taxonomicNameLabel id="CF86511EFFD6FFC05037DFB91243FE46" box="[280,369,475,501]" pageId="9" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
further distinguish the new species from
|
||
<taxonomicName id="21C14BF4FFD6FFC0526CDFB910AAFE46" authorityName="Pati & Thackeray" authorityYear="2018" box="[835,920,475,501]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD6FFC0526CDFB910AAFE46" box="[835,920,475,501]" italics="true" pageId="9" pageNumber="125">G. botti</emphasis>
|
||
</taxonomicName>
|
||
, in which, the G1 ultimate article is relatively stouter (see
|
||
<bibRefCitation id="82504D86FFD6FFC0500EDF9D111CFDAA" author="Pati, S. K. & Thackeray, T." box="[289,558,511,537]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 6I–K) and the vulvae are relatively smaller (occupying ca. 0.3 times the length of the s6) (see
|
||
<bibRefCitation id="82504D86FFD6FFC050ABDC4111BBFD8E" author="Pati, S. K. & Thackeray, T." box="[388,649,547,573]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 6N). Moreover, the vulva is much closer to the s5/s
|
||
<quantity id="21399D92FFD6FFC055DCDC40162CFD8E" box="[1267,1310,546,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="125" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="21C14BF4FFD6FFC0540BDC4116AEFD8E" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[1316,1436,547,573]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD6FFC0540BDC4116AEFD8E" box="[1316,1436,547,573]" italics="true" pageId="9" pageNumber="125">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD6FFC051B8DC2513C2FDD2" bold="true" box="[151,240,583,609]" pageId="9" pageNumber="125">
|
||
<taxonomicNameLabel id="CF86511EFFD6FFC051B8DC2513C2FDD2" box="[151,240,583,609]" pageId="9" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD6FFC051D0DC251266FDD2" box="[255,340,583,609]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">Fig. 4F</figureCitation>
|
||
), while the said structure is located some distance from the s5/s
|
||
<quantity id="21399D92FFD6FFC05538DC241770FDD2" box="[1047,1090,582,609]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="125" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="21C14BF4FFD6FFC05566DC2517AFFDD2" authorityName="Pati & Thackeray" authorityYear="2018" box="[1097,1181,583,609]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="botti">
|
||
<emphasis id="D4B5EC65FFD6FFC05566DC2517AFFDD2" box="[1097,1181,583,609]" italics="true" pageId="9" pageNumber="125">G. botti</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="82504D86FFD6FFC055F7DC2513E1FD36" author="Pati, S. K. & Thackeray, T." pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 6N). Additional difference in the G1 structure separates the new species from
|
||
<taxonomicName id="21C14BF4FFD6FFC0558CDC091607FD36" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" box="[1187,1333,619,645]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="D4B5EC65FFD6FFC0558CDC091607FD36" box="[1187,1333,619,645]" italics="true" pageId="9" pageNumber="125">G. splendida</emphasis>
|
||
</taxonomicName>
|
||
, i.e., the ultimate article is relatively shorter, ca. 0.4 times the length of the penultimate article in
|
||
<taxonomicName id="21C14BF4FFD6FFC05556DCED17C1FD1A" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[1145,1267,655,681]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD6FFC05556DCED17C1FD1A" box="[1145,1267,655,681]" italics="true" pageId="9" pageNumber="125">G. dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD6FFC055D5DCED1666FD1A" bold="true" box="[1274,1364,655,681]" pageId="9" pageNumber="125">
|
||
<taxonomicNameLabel id="CF86511EFFD6FFC055D5DCED1666FD1A" box="[1274,1364,655,681]" pageId="9" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD6FFC0544BDCED138BFD7E" captionStart="FIGURE 3" captionStartId="4.[151,250,1851,1875]" captionTargetBox="[151,1435,417,1828]" captionTargetId="figure-119@4.[151,1435,417,1828]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana dvirupa sp. nov., holotype male (26.11 × 17.52 mm), ZSI-WRC C.2443. A, mouth parts exposed; B, left maxilliped 3; C, outer view of major or right chela; D, pleon; E, dorsal view of left G1; F, dorsal view of left G1 ultimate article; G, ventral view of left G1; H, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B), 1 mm (E, G, H), 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10997148" httpUri="https://zenodo.org/record/10997148/files/figure.png" pageId="9" pageNumber="125">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="7EFA2CF2FFD6FFC051E9DCD113D8FD7E" box="[198,234,691,717]" captionStart="FIGURE 4" captionStartId="5.[151,249,1723,1747]" captionTargetBox="[151,1435,181,1699]" captionTargetId="figure-20@5.[151,1435,181,1699]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana dvirupa sp. nov.: A–C, paratype male (29.99 × 20.05 mm), ZSI-WRC C.2444; D–F, paratype female (28.67 × 19.07 mm), ZSI-WRC C.2444. A, D, habitus, dorsal view; B, pleonal somites 3–6 and telson; C, dorsal view of left G1; E, pleonal somites 4–6 and telson; F, thoracic sternites showing vulvae. Scale bars = 10 mm (A, D), 5 mm (B, E, F), 1 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10997150" httpUri="https://zenodo.org/record/10997150/files/figure.png" pageId="9" pageNumber="125">4C</figureCitation>
|
||
); whereas
|
||
<taxonomicName id="21C14BF4FFD6FFC0504BDCD112C7FD7E" authorityName="Pati, Thackeray & Khaire" authorityYear="2016" box="[356,501,691,717]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="D4B5EC65FFD6FFC0504BDCD112C7FD7E" box="[356,501,691,717]" italics="true" pageId="9" pageNumber="125">G. splendida</emphasis>
|
||
</taxonomicName>
|
||
has a relatively longer ultimate article, ca. 0.5 times the length of the penultimate article (see
|
||
<bibRefCitation id="82504D86FFD6FFC05037DCB51112FD42" author="Pati, S. K. & Thackeray, T." box="[280,544,727,753]" pageId="9" pageNumber="125" pagination="1 - 73" refId="ref6687" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018</bibRefCitation>
|
||
: fig. 10D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEDB63FCFFD6FFC051E8DC9810B0FCEE" pageId="9" pageNumber="125" type="distribution">
|
||
<paragraph id="E67E3077FFD6FFC051E8DC9810B0FCEE" blockId="9.[151,1437,150,861]" pageId="9" pageNumber="125">
|
||
<emphasis id="D4B5EC65FFD6FFC051E8DC981136FCA7" bold="true" box="[199,516,762,788]" pageId="9" pageNumber="125">Geographical distribution.</emphasis>
|
||
<taxonomicName id="21C14BF4FFD6FFC05325DC9911E3FCA6" authority="Pati & Bajantri & Hegde, 2024" authorityName="Pati & Bajantri & Hegde" authorityYear="2024" box="[522,721,763,789]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="125" phylum="Arthropoda" rank="species" species="dvirupa" status="sp. nov.">
|
||
<emphasis id="D4B5EC65FFD6FFC05325DC9911E3FCA6" box="[522,721,763,789]" italics="true" pageId="9" pageNumber="125">Ghatiana dvirupa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D4B5EC65FFD6FFC053F7DC991002FCA6" bold="true" box="[728,816,763,789]" pageId="9" pageNumber="125">
|
||
<taxonomicNameLabel id="CF86511EFFD6FFC053F7DC991002FCA6" box="[728,816,763,789]" pageId="9" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is presently known only from Atle (
|
||
<typeStatus id="397A8ED5FFD6FFC055EEDC9917C1FCA6" box="[1217,1267,763,789]" pageId="9" pageNumber="125">type</typeStatus>
|
||
locality) of the
|
||
<location id="E31E66ACFFD6FFC051B8DD7D12A9FC8A" LSID="urn:lsid:plazi:treatment:6E688161FFDDFFC051B8DEF510B0FCEE:E31E66ACFFD6FFC051B8DD7D12A9FC8A" box="[151,411,799,825]" country="India" county="Central Western Ghats" municipality="Kali Tiger Reserve" name="Uttara Kannada district" pageId="9" pageNumber="125" stateProvince="Karnataka">Uttara Kannada district</location>
|
||
of
|
||
<collectingRegion id="2405FE95FFD6FFC05091DD7D115FFC8A" box="[446,621,799,825]" country="India" name="Karnataka" pageId="9" pageNumber="125">Karnataka state</collectingRegion>
|
||
,
|
||
<collectingCountry id="9ED670E7FFD6FFC05358DD7D1183FC8A" box="[631,689,799,825]" name="India" pageId="9" pageNumber="125">India</collectingCountry>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD6FFC05390DD7D11CDFC8A" box="[703,767,799,825]" captionStart="FIGURE 1" captionStartId="1.[151,250,1589,1613]" captionTargetBox="[151,1436,378,1561]" captionTargetId="figure-106@1.[151,1436,378,1563]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the distribution of Ghatiana dvirupa sp. nov. A, Western Ghats; B, Kali Tiger Reserve; C, Atle (type locality). This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997144" httpUri="https://zenodo.org/record/10997144/files/figure.png" pageId="9" pageNumber="125">Fig. 1</figureCitation>
|
||
). The
|
||
<typeStatus id="397A8ED5FFD6FFC05267DD7D104AFC8A" box="[840,888,799,825]" pageId="9" pageNumber="125">type</typeStatus>
|
||
locality is situated in the
|
||
<location id="E31E66ACFFD6FFC055BEDD7D1665FC8A" LSID="urn:lsid:plazi:treatment:6E688161FFDDFFC051B8DEF510B0FCEE:E31E66ACFFD6FFC055BEDD7D1665FC8A" box="[1169,1367,799,825]" country="India" county="Central Western Ghats" municipality="Kali Tiger Reserve" name="Castlerock Range" pageId="9" pageNumber="125" stateProvince="Karnataka">Castlerock Range</location>
|
||
of the
|
||
<collectingMunicipality id="061AAA0DFFD6FFC051B8DD21125CFCEE" box="[151,366,835,861]" pageId="9" pageNumber="125">Kali Tiger Reserve</collectingMunicipality>
|
||
, which is in the
|
||
<collectingCounty id="0F1F48FBFFD6FFC0530BDD211012FCEE" box="[548,800,835,861]" pageId="9" pageNumber="125">Central Western Ghats</collectingCounty>
|
||
(
|
||
<figureCitation id="7EFA2CF2FFD6FFC05200DD211043FCEE" box="[815,881,835,861]" captionStart="FIGURE 1" captionStartId="1.[151,250,1589,1613]" captionTargetBox="[151,1436,378,1561]" captionTargetId="figure-106@1.[151,1436,378,1563]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the distribution of Ghatiana dvirupa sp. nov. A, Western Ghats; B, Kali Tiger Reserve; C, Atle (type locality). This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.10997144" httpUri="https://zenodo.org/record/10997144/files/figure.png" pageId="9" pageNumber="125">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |