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<document id="D82B03CFB9E93EBAE8F0826A71FAD358" ID-CLB-Dataset="298643" ID-DOI="10.5194/fr-24-33-2021" ID-GBIF-Dataset="d7300392-709d-47cc-a2e5-a51acfe8e4cc" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965797" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1712928270136" checkinUser="felipe" docAuthor="Nonsrirach, Thanit, Manitkoon, Sita &amp; Lauprasert, Komsorn" docDate="2021" docId="6D66DD4E6735FFF1FF23FA9770D4FB56" docLanguage="en" docName="FossRec.24.1.33-47.pdf" docOrigin="Fossil Record 24 (1)" docSource="http://dx.doi.org/10.5194/fr-24-33-2021" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Cyclotosaurus posthumus Fraas 1913" docType="treatment" docVersion="6" lastPageNumber="35" masterDocId="915FA5366737FFF3FF92FFF7754EFFBC" masterDocTitle="First occurrence of brachyopid temnospondyls in Southeast Asia and review of the Mesozoic amphibians from Thailand" masterLastPageNumber="47" masterPageNumber="33" pageNumber="35" updateTime="1718740114086" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2309801A34E0B2DA4C3C4B181E89F74B">First occurrence of brachyopid temnospondyls in Southeast Asia and review of the Mesozoic amphibians from Thailand</mods:title>
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<mods:namePart id="AD53B078E9AA88FB1D1B7DF3859B79F1">Nonsrirach, Thanit</mods:namePart>
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<mods:namePart id="B2D65A8706E8F05C5F0AD19F4A1A65BE">Manitkoon, Sita</mods:namePart>
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<mods:namePart id="16F64F3F64A9EEF3037E96D77F8AADF0">Lauprasert, Komsorn</mods:namePart>
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<mods:title id="84492F9429B435CBDA3D8D26F39955FB">Fossil Record</mods:title>
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<mods:date id="2644A4D19DC7EA044DC65062C6E6461B">2021</mods:date>
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<treatment id="6D66DD4E6735FFF1FF23FA9770D4FB56" ID-DOI="http://doi.org/10.5281/zenodo.12110271" ID-GBIF-Taxon="233223873" ID-Zenodo-Dep="12110271" LSID="urn:lsid:plazi:treatment:6D66DD4E6735FFF1FF23FA9770D4FB56" httpUri="http://treatment.plazi.org/id/6D66DD4E6735FFF1FF23FA9770D4FB56" lastPageNumber="35" pageId="2" pageNumber="35">
<subSubSection id="ADD53FD36735FFF1FF23FA9774ABFAC4" box="[177,485,1376,1400]" pageId="2" pageNumber="35" type="nomenclature">
<paragraph id="E5706C586735FFF1FF23FA9774ABFAC4" blockId="2.[177,485,1376,1400]" box="[177,485,1376,1400]" pageId="2" pageNumber="35">
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<taxonomicName id="22CF17DB6735FFF1FF23FA9774ABFAC4" authorityName="Fraas" authorityYear="1913" box="[177,485,1376,1400]" class="Amphibia" family="Capitosauridae" genus="Cyclotosaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Temnospondyli" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="posthumus">
<emphasis id="D7BBB04A6735FFF1FF23FA977405FAC4" box="[177,331,1376,1400]" italics="true" pageId="2" pageNumber="35">Cyclotosaurus</emphasis>
cf.
<emphasis id="D7BBB04A6735FFF1FEE1FA9774ABFAC4" box="[371,485,1376,1400]" italics="true" pageId="2" pageNumber="35">posthumus</emphasis>
</taxonomicName>
</heading>
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<subSubSection id="ADD53FD36735FFF1FFEEFA4B75A0F83D" pageId="2" pageNumber="35" type="materials_examined">
<paragraph id="E5706C586735FFF1FFEEFA4B741AFA68" blockId="2.[124,340,1468,1492]" box="[124,340,1468,1492]" pageId="2" pageNumber="35">
<emphasis id="D7BBB04A6735FFF1FFEEFA4B741AFA68" bold="true" box="[124,340,1468,1492]" pageId="2" pageNumber="35">Reference material</emphasis>
</paragraph>
<paragraph id="E5706C586735FFF1FFEEFA0E74EAF98C" blockId="2.[124,767,1529,1584]" pageId="2" pageNumber="35">
D.M.
<collectionCode id="83DEF49D6735FFF1FF26FA0E7587F9AC" box="[180,201,1529,1552]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="35">R</collectionCode>
. no. Ch.D 001 (
<figureCitation id="7DF470DD6735FFF1FEE6FA0E7492F9AC" box="[372,476,1529,1552]" captionStart="Figure 2" captionStartId="3.[124,191,1031,1053]" captionTargetBox="[226,1359,177,1000]" captionTargetId="figure-13@3.[226,1360,177,1000]" captionTargetPageId="3" captionText="Figure 2. Triassic amphibian remains from Thailand. A replica skull of Cyclotosaurus cf. posthumus (D.M.R. Ch.D 001) in dorsal (a), palatal (b), and posterior views (c). A reconstruction of Thai Cyclotosauridae (d). A dermal bone of Plagiosauroidea indet. (TF 1453) in ventral (e) and dorsal (f) views. A reconstruction of Thai Plagiosauroidea (g). Reconstruction images (d, g) are not to scale (drawn by Sita Manitkoon)." figureDoi="http://doi.org/10.5281/zenodo.10965801" httpUri="https://zenodo.org/record/10965801/files/figure.png" pageId="2" pageNumber="35">Fig. 2ac</figureCitation>
); a posterior part of a skull (
<bibRefCitation id="815E11A96735FFF1FF11F9EE74D6F98C" author="Ingavat, R. &amp; Janvier, P." box="[131,408,1561,1584]" pageId="2" pageNumber="35" pagination="711 - 725" refId="ref9043" refString="Ingavat, R. and Janvier, P.: Cyclotosaurus cf. posthumus Fraas (Capitosauridae, Stereospondyli) from the Huai Hin Lat Formation (Upper Triassic), northeastern Thailand, with a note on capitosaurid biogeography, Geobios, 14, 711 - 725, 1981." type="journal article" year="1981">Ingavat and Janvier, 1981</bibRefCitation>
).
</paragraph>
<paragraph id="E5706C586735FFF1FFEEF9A3746CF9D0" blockId="2.[124,290,1620,1644]" box="[124,290,1620,1644]" pageId="2" pageNumber="35">
<emphasis id="D7BBB04A6735FFF1FFEEF9A3746CF9D0" bold="true" box="[124,290,1620,1644]" pageId="2" pageNumber="35">Formation/age</emphasis>
</paragraph>
<paragraph id="E5706C586735FFF1FFEEF9657706F915" blockId="2.[124,584,1682,1705]" box="[124,584,1682,1705]" pageId="2" pageNumber="35">Huai Hin Lat Formation (CarnianNorian).</paragraph>
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<emphasis id="D7BBB04A6735FFF1FFEEF93A7439F959" bold="true" box="[124,375,1741,1765]" pageId="2" pageNumber="35">Locality and sediment</emphasis>
</paragraph>
<paragraph id="E5706C586735FFF1FFEEF8FD75A0F83D" blockId="2.[124,767,1802,1921]" pageId="2" pageNumber="35">
<materialsCitation id="55A766056735FFF1FFEEF8FD75A4F83D" ID-GBIF-Occurrence="4892258301" location="Khon San District" pageId="2" pageNumber="35" specimenCount="1" stateProvince="Chaiyaphum">
The specimen was discovered in 1981 at an outcrop near the Chulabhorn Dam,
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,
<collectingRegion id="270BA2BA6735FFF1FD9EF8DD77B2F8FD" box="[524,764,1834,1857]" country="Thailand" name="Chaiyaphum" pageId="2" pageNumber="35">Chaiyaphum Province</collectingRegion>
. The fossil assemblage layer consists of a grey to red conglomerate
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.
</paragraph>
</subSubSection>
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<emphasis id="D7BBB04A6735FFF1FCA1FC6F76F8FC0C" bold="true" box="[819,950,920,944]" pageId="2" pageNumber="35">Description</emphasis>
</paragraph>
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D.M.
<collectionCode id="83DEF49D6735FFF1FCFEFC2376CFFC57" box="[876,897,980,1003]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="35">R</collectionCode>
. no. Ch.D 001 is a posterior part of a comparatively large skull (approximately
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wide and
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long), but the lateral part of the right side is missing. However, a pair of otic notches and a pineal foramen are preserved. The pattern of the ornamentation and the shallow interotic embayment of this skull allow an assignment to
<taxonomicName id="22CF17DB6735FFF1FB1EFB847068FB37" authorityName="Fraas" authorityYear="1889" box="[1164,1318,1139,1163]" class="Amphibia" family="Capitosauridae" genus="Cyclotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="D7BBB04A6735FFF1FB1EFB847068FB37" box="[1164,1318,1139,1163]" italics="true" pageId="2" pageNumber="35">Cyclotosaurus</emphasis>
</taxonomicName>
(
<bibRefCitation id="815E11A96735FFF1FAA1FB8470FFFB36" author="Fraas, E." box="[1331,1457,1139,1162]" pageId="2" pageNumber="35" pagination="275 - 294" refId="ref8826" refString="Fraas, E.: Neue Labyrinthodonten aus der Schwabischen Trias, Palaeontographica, 60, 275 - 294, 1913." type="journal article" year="1913">Fraas, 1913</bibRefCitation>
;
<bibRefCitation id="815E11A96735FFF1FCA1FB647101FB16" author="Ingavat, R. &amp; Janvier, P." box="[819,1103,1171,1194]" pageId="2" pageNumber="35" pagination="711 - 725" refId="ref9043" refString="Ingavat, R. and Janvier, P.: Cyclotosaurus cf. posthumus Fraas (Capitosauridae, Stereospondyli) from the Huai Hin Lat Formation (Upper Triassic), northeastern Thailand, with a note on capitosaurid biogeography, Geobios, 14, 711 - 725, 1981." type="journal article" year="1981">Ingavat and Janvier, 1981</bibRefCitation>
;
<bibRefCitation id="815E11A96735FFF1FBCCFB64704EFB16" author="Damiani, R. J." box="[1118,1280,1171,1194]" pageId="2" pageNumber="35" pagination="379 - 482" refId="ref7994" refString="Damiani, R. J.: A systematic revision and phylogenetic analysis of Triassic mastodonsauroids (Temnospondyli: Stereospondyli), Zool. J. Linn. Soc. - Lond., 133, 379 - 482, 2001." type="journal article" year="2001">Damiani, 2001</bibRefCitation>
) and are similar to those of the
<typeStatus id="3A74D2FA6735FFF1FC45FB44717AFB76" box="[983,1076,1203,1226]" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
of
<taxonomicName id="22CF17DB6735FFF1FBC8FB447626FB56" authority="Fraas, 1913" authorityName="Fraas" authorityYear="1913" class="Amphibia" family="Capitosauridae" genus="Cyclotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="35" phylum="Chordata" rank="species" species="posthumus">
<emphasis id="D7BBB04A6735FFF1FBC8FB447020FB77" box="[1114,1390,1203,1227]" italics="true" pageId="2" pageNumber="35">Cyclotosaurus posthumus</emphasis>
<bibRefCitation id="815E11A96735FFF1FAE4FB447626FB56" author="Fraas, E." pageId="2" pageNumber="35" pagination="275 - 294" refId="ref8826" refString="Fraas, E.: Neue Labyrinthodonten aus der Schwabischen Trias, Palaeontographica, 60, 275 - 294, 1913." type="journal article" year="1913">Fraas, 1913</bibRefCitation>
</taxonomicName>
from the Upper Middle Keuper,
<collectingRegion id="270BA2BA6735FFF1FB55FB247056FB56" box="[1223,1304,1235,1258]" country="Germany" name="Bayern" pageId="2" pageNumber="35">Bavaria</collectingRegion>
(
<collectingCountry id="9DD82CC86735FFF1FAB4FB2470C1FB56" box="[1318,1423,1235,1258]" name="Germany" pageId="2" pageNumber="35">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
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