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<document id="61A21AB7715A5B6C6BC319BC12D154D9" ID-CLB-Dataset="298643" ID-DOI="10.5194/fr-24-33-2021" ID-GBIF-Dataset="d7300392-709d-47cc-a2e5-a51acfe8e4cc" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965797" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1712928270136" checkinUser="felipe" docAuthor="Nonsrirach, Thanit, Manitkoon, Sita &amp; Lauprasert, Komsorn" docDate="2021" docId="6D66DD4E6732FFF4FCFBFEAE7407FA6F" docLanguage="en" docName="FossRec.24.1.33-47.pdf" docOrigin="Fossil Record 24 (1)" docSource="http://dx.doi.org/10.5194/fr-24-33-2021" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Stereospondyli" docType="treatment" docVersion="6" lastPageNumber="38" masterDocId="915FA5366737FFF3FF92FFF7754EFFBC" masterDocTitle="First occurrence of brachyopid temnospondyls in Southeast Asia and review of the Mesozoic amphibians from Thailand" masterLastPageNumber="47" masterPageNumber="33" pageNumber="38" updateTime="1718740114086" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9CB2E62430A46E28E4EEA0F7013B89BD">First occurrence of brachyopid temnospondyls in Southeast Asia and review of the Mesozoic amphibians from Thailand</mods:title>
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<mods:namePart id="8250CB556A1204D52B9A3CB5B36A68DE">Nonsrirach, Thanit</mods:namePart>
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<mods:namePart id="404A32330D2A7A5DAF387B2ED9374A6D">Manitkoon, Sita</mods:namePart>
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<treatment id="6D66DD4E6732FFF4FCFBFEAE7407FA6F" ID-DOI="http://doi.org/10.5281/zenodo.12110307" ID-GBIF-Taxon="233223880" ID-Zenodo-Dep="12110307" LSID="urn:lsid:plazi:treatment:6D66DD4E6732FFF4FCFBFEAE7407FA6F" httpUri="http://treatment.plazi.org/id/6D66DD4E6732FFF4FCFBFEAE7407FA6F" lastPageId="7" lastPageNumber="38" pageId="5" pageNumber="38">
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<taxonomicName id="22CF17DB6732FFF6FCFBFEAE715FFECD" box="[873,1041,345,369]" class="Amphibia" isUncertain="true" kingdom="Animalia" pageId="5" pageNumber="46" rank="species" subOrder="Stereospondyli">
<emphasis id="D7BBB04A6732FFF6FCFBFEAE715FFECD" bold="true" box="[873,1041,345,369]" pageId="5" pageNumber="38">Stereospondyli</emphasis>
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indet.
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<emphasis id="D7BBB04A6732FFF6FCA1FE477145FE74" bold="true" box="[819,1035,432,456]" pageId="5" pageNumber="38">Reference material</emphasis>
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<paragraph id="E5706C586732FFF6FCA1FE1B7106FDDF" blockId="5.[819,1462,492,611]" pageId="5" pageNumber="38">
<collectionCode id="83DEF49D6732FFF6FCA1FE1B76CAFDBF" box="[819,900,492,515]" pageId="5" pageNumber="38">CY-HN</collectionCode>
378 (
<figureCitation id="7DF470DD6732FFF6FC5FFE1B7174FDBF" box="[973,1082,492,515]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5ad</figureCitation>
), intercentrum (
<bibRefCitation id="815E11A96732FFF6FB7CFE1B7623FD9F" author="Laojumpon, C. &amp; Deesri, U. &amp; Khamha, S. &amp; Wattanapituksakul, A. &amp; Lauprasert, K. &amp; Suteethorn, S. &amp; Suteethorn, V." pageId="5" pageNumber="38" pagination="335 - 343" refId="ref9321" refString="Laojumpon, C., Deesri, U., Khamha, S., Wattanapituksakul, A., Lauprasert, K., Suteethorn, S., and Suteethorn, V.: New vertebrate-bearing localities in the Triassic of Thailand, J. Sci. Technol. MSU, 4, 335 - 343, 2014." type="journal article" year="2014">Laojumpon et al., 2014</bibRefCitation>
);
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364 (
<figureCitation id="7DF470DD6732FFF6FB87FDFB7135FD9F" box="[1045,1147,524,547]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5ef</figureCitation>
), left clavicle bone; and
<collectionCode id="83DEF49D6732FFF6FA19FDFB7617FDFF" pageId="5" pageNumber="38">CY-HN</collectionCode>
365 (
<figureCitation id="7DF470DD6732FFF6FC07FDDB76B0FDFF" box="[917,1022,556,579]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">
Fig.
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h
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),
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368 (
<figureCitation id="7DF470DD6732FFF6FB0EFDDB71B7FDFF" box="[1180,1273,556,579]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5ij</figureCitation>
), and
<collectionCode id="83DEF49D6732FFF6FAA5FDDB70C6FDFF" box="[1335,1416,556,579]" pageId="5" pageNumber="38">CY-HN</collectionCode>
377 (
<figureCitation id="7DF470DD6732FFF6FCA9FDBB76D1FDDF" box="[827,927,588,611]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5kl</figureCitation>
), dermal bones.
</paragraph>
<paragraph id="E5706C586732FFF6FCA1FD717697FD22" blockId="5.[819,985,646,670]" box="[819,985,646,670]" pageId="5" pageNumber="38">
<emphasis id="D7BBB04A6732FFF6FCA1FD717697FD22" bold="true" box="[819,985,646,670]" pageId="5" pageNumber="38">Formation/age</emphasis>
</paragraph>
<paragraph id="E5706C586732FFF6FCA1FD35704EFD65" blockId="5.[819,1280,706,729]" box="[819,1280,706,729]" pageId="5" pageNumber="38">Huai Hin Lat Formation (CarnianNorian).</paragraph>
<paragraph id="E5706C586732FFF6FCA1FD0B7160FCA8" blockId="5.[819,1070,764,788]" box="[819,1070,764,788]" pageId="5" pageNumber="38">
<emphasis id="D7BBB04A6732FFF6FCA1FD0B7160FCA8" bold="true" box="[819,1070,764,788]" pageId="5" pageNumber="38">Locality and sediment</emphasis>
</paragraph>
<paragraph id="E5706C586732FFF6FCA1FCCF711AFBF2" blockId="5.[819,1462,824,1102]" pageId="5" pageNumber="38">
These specimens were collected by a ThaiFrench paleontological team during an excursion in 2010 at Huai Nam Aun, an outcrop situated near the Nong Yakong village, Khon San District,
<collectingRegion id="270BA2BA6732FFF6FC01FC6071C8FC12" box="[915,1158,919,942]" country="Thailand" name="Chaiyaphum" pageId="5" pageNumber="38">Chaiyaphum Province</collectingRegion>
. This locality contains various beds of limestone and mudstone, deposited in brackish water near a calcium carbonate source (
<bibRefCitation id="815E11A96732FFF6FB62FC207623FBB2" author="Laojumpon, C. &amp; Deesri, U. &amp; Khamha, S. &amp; Wattanapituksakul, A. &amp; Lauprasert, K. &amp; Suteethorn, S. &amp; Suteethorn, V." pageId="5" pageNumber="38" pagination="335 - 343" refId="ref9321" refString="Laojumpon, C., Deesri, U., Khamha, S., Wattanapituksakul, A., Lauprasert, K., Suteethorn, S., and Suteethorn, V.: New vertebrate-bearing localities in the Triassic of Thailand, J. Sci. Technol. MSU, 4, 335 - 343, 2014." type="journal article" year="2014">Laojumpon et al., 2014</bibRefCitation>
). Other fossils such as
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<emphasis id="D7BBB04A6732FFF6FBF3FC0171F1FBB2" box="[1121,1215,1014,1038]" italics="true" pageId="5" pageNumber="38">Hybodus</emphasis>
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teeth, bony fish scales, and coprolites have also been found from this locality (
<bibRefCitation id="815E11A96732FFF6FA10FBE0714AFBF2" author="Laojumpon, C. &amp; Matkhammee, T. &amp; Wathanapitaksakul, A. &amp; Suteethorn, V. &amp; Suteethorn, S. &amp; Lauprasert, K. &amp; Srisuk, S. &amp; Loeuff, J. L. E." pageId="5" pageNumber="38" pagination="207 - 213" refId="ref9240" refString="Laojumpon, C., Matkhammee, T., Wathanapitaksakul, A., Suteethorn, V., Suteethorn, S., Lauprasert, K., Srisuk, S., Loeuff, J. L. E.: Preliminary report on coprolites from the Late Triassic of Thailand, Vertebrate Coprolites, Bull. N. Mex. Mus. Nat. Hist. Sci., 57, 207 - 213, 2012." type="journal article" year="2012">Laojumpon et al., 2012</bibRefCitation>
,
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).
</paragraph>
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<emphasis id="D7BBB04A6732FFF6FCA1FB8676F8FB35" bold="true" box="[819,950,1137,1161]" pageId="5" pageNumber="38">Description</emphasis>
</paragraph>
<paragraph id="E5706C586732FFF6FCA1FB5A76F4F8BD" blockId="5.[819,1463,1197,1921]" pageId="5" pageNumber="38">
The material consists of an intercentrum and several fragments of dermal bones. A large intercentrum (
<collectionCode id="83DEF49D6732FFF6FABEFB3B7031FB5F" box="[1324,1407,1228,1251]" pageId="5" pageNumber="38">CY-HN</collectionCode>
378,
<figureCitation id="7DF470DD6732FFF6FCA1FB1B76D6FABF" box="[819,920,1260,1283]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5ad</figureCitation>
) is preserved over a length of approximately
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and is
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thick.
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378 is a disk-shaped intercentrum with reduced or absent pleurocentra (
<bibRefCitation id="815E11A96732FFF6FA8DFADB76D6FADF" author="Laojumpon, C. &amp; Deesri, U. &amp; Khamha, S. &amp; Wattanapituksakul, A. &amp; Lauprasert, K. &amp; Suteethorn, S. &amp; Suteethorn, V." pageId="5" pageNumber="38" pagination="335 - 343" refId="ref9321" refString="Laojumpon, C., Deesri, U., Khamha, S., Wattanapituksakul, A., Lauprasert, K., Suteethorn, S., and Suteethorn, V.: New vertebrate-bearing localities in the Triassic of Thailand, J. Sci. Technol. MSU, 4, 335 - 343, 2014." type="journal article" year="2014">Laojumpon et al., 2014</bibRefCitation>
). The anterior and posterior faces of
<collectionCode id="83DEF49D6732FFF6FAA1FABB70CAFADF" box="[1331,1412,1356,1379]" pageId="5" pageNumber="38">CY-HN</collectionCode>
378 are concave. Laterally, the parapophyses are located in less than half of the total length. These features share characters with
<taxonomicName id="22CF17DB6732FFF6FC05FA5B7178FA7F" box="[919,1078,1452,1475]" pageId="5" pageNumber="46" rank="subOrder" subOrder="Stereospondyli">Stereospondyli</taxonomicName>
(
<bibRefCitation id="815E11A96732FFF6FBD7FA5B7041FA7F" author="Milner, A. R. &amp; Fraser, N. C. &amp; Sues, H. D." box="[1093,1295,1452,1475]" pageId="5" pageNumber="38" refId="ref10226" refString="Milner, A. R., Fraser, N. C., and Sues, H. D. (Eds.): Late Triassic and Jurassic amphibians: fossil record and phylogeny, in: The shadow of the dinosaurs: Early Mesozoic Tetrapods, Cambridge University Press, USA, 221 pp., 1994." type="book" year="1994">Milner et al., 1994</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FA89FA5B769EFA5E" author="Witzmann, F. &amp; Gassner, T." pageId="5" pageNumber="38" pagination="37 - 51" refId="ref12377" refString="Witzmann, F. and Gassner, T.: Metoposaurid and mastodonsaurid stereospondyls from the Triassic - Jurassic boundary of Portugal, Alcheringa, 32, 37 - 51, 2008." type="journal article" year="2008">Witzmann and Gassner, 2008</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FC4FFA3C71AEFA5E" author="Laojumpon, C. &amp; Deesri, U. &amp; Khamha, S. &amp; Wattanapituksakul, A. &amp; Lauprasert, K. &amp; Suteethorn, S. &amp; Suteethorn, V." box="[989,1248,1483,1506]" pageId="5" pageNumber="38" pagination="335 - 343" refId="ref9321" refString="Laojumpon, C., Deesri, U., Khamha, S., Wattanapituksakul, A., Lauprasert, K., Suteethorn, S., and Suteethorn, V.: New vertebrate-bearing localities in the Triassic of Thailand, J. Sci. Technol. MSU, 4, 335 - 343, 2014." type="journal article" year="2014">Laojumpon et al., 2014</bibRefCitation>
). The shape of
<collectionCode id="83DEF49D6732FFF6FA19FA3C7617F9BE" pageId="5" pageNumber="38">CY-HN</collectionCode>
378 is comparable to those of the Metoposauroidea or Mastodonsauroidea intercentrum (
<bibRefCitation id="815E11A96732FFF6FB30F9FC70FFF99E" author="Moser, M. &amp; Schoch, R." box="[1186,1457,1547,1570]" pageId="5" pageNumber="38" pagination="1245 - 1266" refId="ref10287" refString="Moser, M. and Schoch, R.: Revision of the type material and nomenclature of Mastodonsaurus giganteus (Jaeger) (Temnospondyli) from the Middle Triassic of Germany, Palaeontology, 50, 1245 - 1266, 2007." type="journal article" year="2007">Moser and Schoch, 2007</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FCA1F9DC76F8F9FE" author="Sulej, T." box="[819,950,1579,1602]" pageId="5" pageNumber="38" pagination="29 - 139" refId="ref11520" refString="Sulej, T.: Osteology, variability, and evolution of Metoposaurus, a temnospondyl from the Late Triassic of Poland, Palaeontol. Pol., 64, 29 - 139, 2007." type="journal article" year="2007">Sulej, 2007</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FC55F9DC71E3F9FE" author="Fortuny, J. &amp; Arbez, T. &amp; Mujal, E. &amp; Steyer, J. S." box="[967,1197,1579,1602]" pageId="5" pageNumber="38" pagination="1576701" refId="ref8730" refString="Fortuny, J., Arbez, T., Mujal, E., and Steyer, J. S.: Reappraisal of &quot; Metoposaurus hoffmani &quot; Dutuit, 1978, and description of new temnospondyl specimens from the Middle-Late Triassic of Madagascar (Morondava Basin), J. Vertebr. Paleontol., 39, e 1576701, https: // doi. org / 10.1080 / 02724634.2019.1576701, 2019." type="journal article" year="2019">Fortuny et al., 2019</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FB2CF9DC70E5F9FE" author="Marzola, M. &amp; Mateus, O. &amp; Shubin, N. H. &amp; Clemmensen, L. B." box="[1214,1451,1579,1602]" pageId="5" pageNumber="38" pagination="1303501" refId="ref9974" refString="Marzola, M., Mateus, O., Shubin, N. H., and Clemmensen, L. B.: Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland), J. Vertebr. Paleontol., 37, e 1303501, https: // doi. org / 10.1080 / 02724634.2017.1303501, 2017." type="journal article" year="2017">Marzola et al., 2017</bibRefCitation>
). The Plagiosauroidea possess cylindrical intercentra (
<bibRefCitation id="815E11A96732FFF6FAF4F9BC7692F93E" author="Warren, A. &amp; Snell, N." pageId="5" pageNumber="38" pagination="43 - 64" refId="ref12191" refString="Warren, A. and Snell, N.: The postcranial skeleton of Mesozoic temnospondyl amphibians: A review, Alcheringa, 15, 43 - 64, 1991." type="journal article" year="1991">Warren and Snell, 1991</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FC7AF99C7051F93E" author="Konietzko-Meier, D. &amp; Danto, M." box="[1000,1311,1643,1666]" pageId="5" pageNumber="38" pagination="747 - 764" refId="ref9195" refString="Konietzko-Meier, D., Danto, M., and G ˛ adek, K.: The microstructural variability of the intercentra among temnospondyl amphibians, Biol. J. Linn. Soc., 112, 747 - 764, 2014." type="journal article" year="2014">Konietzko-Meier et al., 2014</bibRefCitation>
). The
<taxonomicName id="22CF17DB6732FFF6FAF2F99C76C9F91E" pageId="5" pageNumber="44" rank="superFamily" superFamily="Brachyopoidea">Brachyopoidea</taxonomicName>
commonly show a wedge-shaped intercentrum in lateral view (
<bibRefCitation id="815E11A96732FFF6FC28F95C7113F97E" author="Shishkin, M. A." box="[954,1117,1707,1730]" pageId="5" pageNumber="38" pagination="78 - 91" refId="ref11274" refString="Shishkin, M. A.: A late Jurassic labyrinthodont from Mongolia, Paleontol. J., 1, 78 - 91, 1991." type="journal article" year="1991">Shishkin, 1991</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FBF4F95C7011F97E" author="Warren, A. &amp; Snell, N." box="[1126,1375,1707,1730]" pageId="5" pageNumber="38" pagination="43 - 64" refId="ref12191" refString="Warren, A. and Snell, N.: The postcranial skeleton of Mesozoic temnospondyl amphibians: A review, Alcheringa, 15, 43 - 64, 1991." type="journal article" year="1991">Warren and Snell, 1991</bibRefCitation>
;
<bibRefCitation id="815E11A96732FFF6FAFAF95C76FEF95D" author="Warren, A. &amp; Rich, T. H. &amp; Vickers-Rich, P." pageId="5" pageNumber="38" pagination="10 - 24" refId="ref12299" refString="Warren, A., Rich, T. H., and Vickers-Rich, P.: The last labyrinthodonts, Palaeontographica Abteilung Stuttgart, 247, 10 - 24, 1997." type="journal article" year="1997">Warren et al., 1997</bibRefCitation>
,
<bibRefCitation id="815E11A96732FFF6FC2EF93D76BAF95D" author="Warren, A. &amp; Rozefelds, A. C. &amp; Bull, S." box="[956,1012,1738,1761]" pageId="5" pageNumber="38" pagination="738 - 753" refId="ref12333" refString="Warren, A., Rozefelds, A. C., and Bull, S.: Tupilakosaur-like vertebrae in Bothriceps australis, an Australian brachyopid stereospondyli, J. Vertebr. Paleontol., 31, 738 - 753, 2011." type="journal article" year="2011">2011</bibRefCitation>
; Warren and Dammiani, 1999;
<bibRefCitation id="815E11A96732FFF6FADBF93D76E0F8BD" author="Averianov, A. O. &amp; Martin, T. &amp; Skutschas, P. P. &amp; Rezvyi, A. S. &amp; Bakirov, A. A." pageId="5" pageNumber="38" pagination="471 - 485" refId="ref6508" refString="Averianov, A. O., Martin, T., Skutschas, P. P., Rezvyi, A. S., and Bakirov, A. A.: Amphibians from the Middle Jurassic Balabansai Svita in the Fergana Depression, Kyrgyzstan (Central Asia), Palaeontol., 51, 471 - 485, 2008." type="journal article" year="2008">Averianov et al., 2008</bibRefCitation>
).
</paragraph>
<paragraph id="E5706C586732FFF4FCDCF8FD7407FA6F" blockId="5.[819,1463,1197,1921]" lastBlockId="7.[124,767,1245,1491]" lastPageId="7" lastPageNumber="40" pageId="5" pageNumber="38">
The other undescribed specimens are dermal bone fragments. The largest fragment (
<collectionCode id="83DEF49D6732FFF6FBE2F8DD718DF8FD" box="[1136,1219,1834,1857]" pageId="5" pageNumber="38">CY-HN</collectionCode>
364,
<figureCitation id="7DF470DD6732FFF6FA90F8DD702BF8FD" box="[1282,1381,1834,1857]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="5" pageNumber="38">Fig. 5ef</figureCitation>
) shows two distinct
<typeStatus id="3A74D2FA6732FFF6FC2FF8BD76BBF8DD" box="[957,1013,1866,1889]" pageId="5" pageNumber="38">types</typeStatus>
of heavy ornamentation in ventral view, i.e., radial and polygonal sculptures. The radial sculpture consists of parallel or radial ridges without transverse ridges, while the polygonal sculpture, located close to the lateral edge of the bone, consists of short ridges connected and forming polygons in a honeycomb or hexagonal shape. The dorsal surface is smooth with a part of the ascending process located along the lateral edge. The polygonal ornamentation located close to the lateral edge and the presence of an ascending process indicate that
<collectionCode id="83DEF49D6731FFF5FB01F91771AAF94B" box="[1171,1252,1760,1783]" pageId="6" pageNumber="39">CY-HN</collectionCode>
364 is a left clavicle bone. The other small dermal fragments, i.e.,
<collectionCode id="83DEF49D6731FFF5FAF7F8F770F8F8AB" box="[1381,1462,1792,1815]" pageId="6" pageNumber="39">CY-HN</collectionCode>
365 (
<figureCitation id="7DF470DD6731FFF5FCFCF8D77693F88B" box="[878,989,1824,1847]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="6" pageNumber="39">
Fig.
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h
</figureCitation>
), CY-HN-368 (
<figureCitation id="7DF470DD6731FFF5FB1BF8D771A5F88B" box="[1161,1259,1824,1847]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="6" pageNumber="39">Fig. 5ij</figureCitation>
), and CY-HN-377 (
<figureCitation id="7DF470DD6731FFF5FCA9F8B776EEF8EB" box="[827,928,1856,1879]" captionStart="Figure 5" captionStartId="7.[124,191,1068,1090]" captionTargetBox="[226,1360,177,1036]" captionTargetId="figure-13@7.[226,1360,177,1036]" captionTargetPageId="7" captionText="Figure 5. New Triassic amphibian remains from Thailand. An intercentrum of Stereospondyli indet. (CY-HN 378) in anterior (a), posterior (b), left (c), and right (d) views. Dermal bone fragments of Stereospondyli indet. of CY-HN 364 (ef), CY-HN 365 (gh), CY-HN- 368 (ij), and CY-HN-377 (kl) in ventral view and dorsal view. Abbreviations: ap, ascending process; pp, parapophyses; ps, polygonal sculpture; rs, radial sculpture." figureDoi="http://doi.org/10.5281/zenodo.10965809" httpUri="https://zenodo.org/record/10965809/files/figure.png" pageId="6" pageNumber="39">Fig. 5kl</figureCitation>
), are covered by radial and polygonal ornamentation in ventral view. These specimens, however, could be assigned as either interclavicle or clavicle bones. Based on the shape of the intercentrum and ornamentation pattern, these specimens could refer to either the late Triassic Mastodonsauroidea (e.g.
<taxonomicName id="22CF17DB6730FFF4FE8FFACB74F9FAE8" authorityName="Fraas" authorityYear="1889" box="[285,439,1340,1364]" class="Amphibia" family="Capitosauridae" genus="Cyclotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="D7BBB04A6730FFF4FE8FFACB74F9FAE8" box="[285,439,1340,1364]" italics="true" pageId="7" pageNumber="40">Cyclotosaurus</emphasis>
</taxonomicName>
) or Metoposauroidea (
<bibRefCitation id="815E11A96730FFF4FD3AFACB75BFFACF" author="Chowdhury, T. R." pageId="7" pageNumber="40" pagination="1 - 52" refId="ref7592" refString="Chowdhury, T. R.: A new metoposaurid amphibian from the Upper Triassic Maleri Formation of Central India, Philos. T. Roy. Soc. B, 250, 1 - 52, 1965." type="journal article" year="1965">Chowdhury, 1965</bibRefCitation>
;
<bibRefCitation id="815E11A96730FFF4FF6FFAAB74B3FACF" author="Warren, A. &amp; Snell, N." box="[253,509,1372,1395]" pageId="7" pageNumber="40" pagination="43 - 64" refId="ref12191" refString="Warren, A. and Snell, N.: The postcranial skeleton of Mesozoic temnospondyl amphibians: A review, Alcheringa, 15, 43 - 64, 1991." type="journal article" year="1991">Warren and Snell, 1991</bibRefCitation>
;
<bibRefCitation id="815E11A96730FFF4FD9BFAAB77B4FACF" author="Sulej, T. &amp; Majer, D." box="[521,762,1372,1395]" pageId="7" pageNumber="40" pagination="157 - 170" refId="ref11556" refString="Sulej, T. and Majer, D.: The temnospondyl amphibian Cyclotosaurus from the Upper Triassic of Poland, Palaeontology, 48, 157 - 170, 2005." type="journal article" year="2005">Sulej and Majer, 2005</bibRefCitation>
;
<bibRefCitation id="815E11A96730FFF4FFEEFA8B75B2FA2F" author="Sulej, T." box="[124,252,1404,1427]" pageId="7" pageNumber="40" pagination="29 - 139" refId="ref11520" refString="Sulej, T.: Osteology, variability, and evolution of Metoposaurus, a temnospondyl from the Late Triassic of Poland, Palaeontol. Pol., 64, 29 - 139, 2007." type="journal article" year="2007">Sulej, 2007</bibRefCitation>
;
<bibRefCitation id="815E11A96730FFF4FE99FA8B74A1FA2F" author="Brusatte, S. L. &amp; Butler, R. J. &amp; Mateus, O. &amp; Steyer, J. S." box="[267,495,1404,1427]" pageId="7" pageNumber="40" pagination="912988" refId="ref6686" refString="Brusatte, S. L., Butler, R. J., Mateus, O., and Steyer, J. S.: A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls, J. Vertebr. Paleontology, 35, e 912988, https: // doi. org / 10.1080 / 02724634.2014.912988, 2015." type="journal article" year="2015">Brusatte et al., 2015</bibRefCitation>
;
<bibRefCitation id="815E11A96730FFF4FE6CFA8B75F8FA0F" author="Antczak, M. &amp; Bodzioch, A." pageId="7" pageNumber="40" pagination="5267" refId="ref6464" refString="Antczak, M. and Bodzioch, A.: Ornamentation of dermal bones of Metoposaurus krasiejowensis and its ecological implications, PeerJ, 6, e 5267, https: // doi. org / 10.7717 / peerj. 5267, 2018." type="journal article" year="2018">Antczak and Bodzioch, 2018</bibRefCitation>
). Therefore, more evidence is required for a familylevel identification.
</paragraph>
</subSubSection>
</treatment>
</document>