125 lines
13 KiB
XML
125 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.250.3715" ID-GBIF-Dataset="4f61fd14-c253-4e81-b31a-b04e06300223" ID-PMC="PMC3558971" ID-Pensoft-Pub="1313-2970-250-1" ID-PubMed="23378803" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-250-1" ModsDocOrigin="ZooKeys 250" ModsDocTitle="Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451247866083" checkinUser="pensoft" docAuthor="Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael" docDate="2012" docId="67ECDBCCAF17497828ACF1623D687300" docLanguage="en" docName="ZooKeys 250: 1-76" docOrigin="ZooKeys 250" docSource="http://dx.doi.org/10.3897/zookeys.250.3715" docTitle="Exocelina brahminensis Shaverdo, Hendrich & Balke, sp. n." docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="FFF3FFABFFA8FF93FFA65605CF65FFB4" masterDocTitle="Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="9" updateTime="1668155011676" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Shaverdo, Helena V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Surbakti, Suriani</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Hendrich, Lars</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Balke, Michael</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2012</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>250</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>76</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.250.3715</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.250.3715</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-250-1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="152039174" LSID="urn:lsid:zoobank.org:act:E31C6D32-6B87-4E19-B676-94F6EE79CA84" httpUri="http://treatment.plazi.org/id/67ECDBCCAF17497828ACF1623D687300" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
|
||
<subSubSection pageId="8" pageNumber="9" type="nomenclature">
|
||
<paragraph pageId="8" pageNumber="9">
|
||
7.
|
||
<taxonomicName LSID="urn:lsid:zoobank.org:act:E31C6D32-6B87-4E19-B676-94F6EE79CA84" authority="Shaverdo, Hendrich & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina brahminensis" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brahminensis">Exocelina brahminensis Shaverdo, Hendrich & Balke</taxonomicName>
|
||
<taxonomicNameLabel pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
|
||
Figs 21
|
||
<normalizedToken originalValue="A–E">A-E</normalizedToken>
|
||
, 47
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="type locality">
|
||
<paragraph pageId="8" pageNumber="9">Type locality.</paragraph>
|
||
<paragraph pageId="8" pageNumber="9">
|
||
Papua New Guinea: Madang Province, Adalbert Mts., near Keki,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-4.705">04°42.30'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="9" value="145.41817">145°25.09'E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="type material">
|
||
<paragraph pageId="8" pageNumber="9">Type material.</paragraph>
|
||
<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
Holotype: male "Papua New Guinea: Madang, Adalbert Mts., creek nr Keki, 790m, 28.xi.1994, 04.42.300S, 145.25.089E, Binatang Boys leg. (PNG 53a)" (ZSM). Paratypes: Madang: 20 males, 27 females with the same label as the holotype (NARI, NHMW, ZSM). 2 males "Papua New Guinea: Madang, Brahmin, 150 m, 26.IX.2002 Balke & Sagata (PNG 024)" (ZSM). 6 males, 1 female "Papua New Guinea: Madang, Akameku-Brahmin, Bismarck Range, 750m, 25.xi.1994, nr 05.49.307S, 145.24.389E, Balke & Kinibel (PNG 114)", one of them additionally with a green label "DNA M.Balke 1524" (NHMW, ZSM). 20 males "Papua New Guinea: Madang, Usino, 260m, 15.iii.2007, 05.31.125S, 145.25.316E, Kinibel (PNG 158)", one of them additionally with a green label "DNA M.Balke 3319" (ZSM). 1 male, 1 female "Papua New Guinea: Madang, Highway nr. Madang, ford, 80m, 26.xi./2.-3.xii.1994, 05.24.405S, 145.38.213E, Binatang Boys, (PNG 117)", the male additionally with a green label: "DNA M.Balke 1523" (ZSM). 1 male, 2 female "Papua New Guinea: Madang, Mt. Tapo, 180 m, ii.2008 5 24.11.00 S 145 36 17 16 E, BRC leg. (PNG 178)" (ZSM). 2 males, 2 females "Papua New Guinea: Madang, Wannang, 230m 3.x.2008, 05.17.235S, 145.06.160E, Posman (PNG188)", one of them additionally with a green label "DNA M.Balke 3760" (ZSM). 3 males, 6 females "Papua New Guinea: Madang, Wannang, 270m, 31.x.2008, 05.15.458S 145.02.389, Posman, (PNG187)", one of them additionally with a green label: "DNA M. Balke 4168" (NHMW, ZSM). 29 males, 24 females "Papua New Guinea: Madang, Adalbert Mts., Sewan- Keki, 700m, 4.v.1994, 04.42.215S, 145.25.154E, Balke & Manaono (PNG 51)" (NHMW, ZSM). 14 males, 11 females "Papua New Guinea: Madang, Adalbert Mts., Keki, 850m, 5.v.1994, nr 04.42.300S, 145.25.089E, Balke & Manaono (PNG 52)" (NHMW, ZSM). 45 males, 29 females "Papua New Guinea: Madang, Adalbert Mts., below Keki, 790m, 5.v.1994, 04.42.300S, 145.25.089E, Balke & Manaono (PNG 53)" (NHMW, ZSM). 2 males, 6 females "Papua New Guinea: Madang, Adalbert Mts., Keki to Sewan, 650m, 7.v.1994, 04.41.802S, 145.25.460E, Balke (PNG 54)", one of them additionally with a green label "DNA M.Balke 1298" (ZSM). 15 males,
|
||
<pageBreakToken pageId="9" pageNumber="10" start="start">16</pageBreakToken>
|
||
females "Papua New Guinea: Madang, Keki, Adalbert Mts., 400m, 29.xi.1994, 04.43.058S, 145.24.437E, Binatang Boys, (PNG 119)" (NHMW, ZSM). 44 males, 35 females "Papua New Guinea: Madang, Keki-Sewan, Adalbert Mts., 700m, 30.xi.1994 nr 04.41.802S, 145.25.460E Binatang Boys (PNG 120)" (NHMW, ZSM). Sandaun: 2 males "Papua New Guinea: Sandaun, Toricelli Mts., village below Sibilanga Stn., 400m, 18.iv.1994, nr 03.26.750S, 142.29.949E, Balke (PNG 42)", one of them additionally with a green label "DNA M.Balke 1293" (ZSM). 13 males, 8 females "Papua New Guinea: Sandaun, Toricelli Mts., 2h walk fr Sibilanga Stn., 350m, 19.iv.1994, 03.39.121S, 142.29.991E, Balke (PNG 44)" (NHMW, ZSM). East Sepik: 2 males, 2 females "Papua New Guinea: East Sepik, Prince Alexander Mts., Wewak, 400m, 21.iv.1994, 03.37.319S, 143.36.764E, Balke (PNG 45)", one of them additionally with a green label "DNA M.Balke 1294" (NHMW, ZSM). Morobe: 1 male "Papua New Guinea: Morobe, Herzog Mts., Bundun, 700-800m, 2.iv.1994, 06.51.598S, 146.37.07E, Balke & Sagata (PNG 27)" (ZSM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="10" type="additional material">
|
||
<paragraph pageId="9" pageNumber="10">Additional material.</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
8 females "Papua New Guinea: Madang, Keki, Adalbert Mts., 400m, 29.xi.1994, 04.43.058S, 145.24.437E, Binatang Boys, (PNG 119)" (NHMW, ZSM). 8 females "Papua New Guinea: Madang, Keki-Sewan, Adalbert Mts., 700m, 30.xi.1994 nr 04.41.802S, 145.25.460E Binatang Boys (PNG 120)" (NHMW, ZSM). These females most likely belong to
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina brahminensis" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brahminensis">Exocelina brahminensis</taxonomicName>
|
||
sp. n. However, the sure separation from
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina larsoni" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="larsoni">Exocelina larsoni</taxonomicName>
|
||
(Balke, 1998) is not possible because of more strongly developed traces of a lateral bead of the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="10" type="diagnosis">
|
||
<paragraph pageId="9" pageNumber="10">Diagnosis.</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
Beetle small, piceous, with reddish brown head and pronotum, shiny; pronotum without lateral bead or with weak traces of lateral bead; male antennomeres simple; male protarsomere 4 with large, thick, strongly curved anterolateral hook; median lobe with weak submedian constriction in ventral view; paramere with strong notch on dorsal side, with notch tip sharply pointed, and subdistal part elongate, with upper setae thin and more numerous and lower setae shorter, thick, and flattend. The species is similar to
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina eme" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="eme">Exocelina eme</taxonomicName>
|
||
sp. n., from which it differs by simple male antennae and the shape of the male genialia, especially by the strong notch of the paramere and its sharply poined tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
|
||
<paragraph pageId="9" pageNumber="10">Description.</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">Size and shape: Beetle small (TL-H 3.15-3.3 mm, TL 3.5-3.65 mm, MW 1.7-1.8 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Head reddish-brown, darker posterior eyes; pronotum reddish-brown, with slightly darker disc; elytra piceous, with dark brown sutural bands; head appendages yellow to yellowish-red, legs distally darker, to dark brown (Fig. 47).</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
Surface sculpture: as in
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina eme" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="eme">Exocelina eme</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">Structures: Pronotum without lateral bead or with weak traces of lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, anteriorly smooth, without anterolateral extensions. Blade of prosternal process lanceolate, relatively broad, convex, with distinct bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal sternite 7 broadly rounded apically.</paragraph>
|
||
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Male: Antenna simple (Fig. 21A). Protarsomere 4 with large, thick, strongly curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 16 short setae and
|
||
<pageBreakToken pageId="10" pageNumber="11" start="start">posterior</pageBreakToken>
|
||
row with 7 short setae (Fig. 21B). Abdominal sternite 7 with 6-13 lateral striae on each side. Median lobe with weak submedian constriction in ventral view (Fig. 21C). Paramere with strong notch on dorsal side, with notch tip sharply pointed, and subdistal part elongate, with upper setae thin and more numerous and lower setae shorter, thick, and flattend (Fig. 21E).
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Female: Traces of the pronotal lateral bead developed stronger in some specimens; abdominal sternite 7 without striae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph pageId="10" pageNumber="11">Distribution.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Papua New Guinea. This species is wide distributed in the Momase Region: Sandaun, East Sepik, Madang, and Morobe Provinces (Fig. 50).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph pageId="10" pageNumber="11">Etymology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">The name refers to the village Brahmin where this species was first discovered. The species name is an adjective in the nominative singular.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |