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<document id="D480B81A86FE0426B41CB7479AEEE531" ID-CLB-Dataset="282875" ID-DOI="10.5852/ejt.2023.906.2329" ID-GBIF-Dataset="c2f9b726-1083-40ff-ada7-97e6df2733ae" ID-ISSN="2118-9773" ID-Zenodo-Dep="10424508" ID-ZooBank="01194EAD-7129-4876-82F9-2173E49C1B0A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1699963422416" checkinUser="plazi" docAuthor="Delgado, Juan A. &amp; Palma, Ricardo L." docDate="2023" docId="6703879E4D240473F72FB78964DEFDC6" docLanguage="en" docName="EJT.2023.906.1-71.pdf" docOrigin="European Journal of Taxonomy 906" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2329/10133" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cholovocera attae" docType="treatment" docVersion="6" lastPageNumber="38" masterDocId="9B3AFFE64D050456F57CB11E677FFFA5" masterDocTitle="A revision of the genus Cholovocera Victor, 1838 (Coleoptera: Endomychidae)" masterLastPageNumber="71" masterPageNumber="1" pageNumber="34" updateTime="1703259287614" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<taxonomicName id="28AA4D0B4D240477F72FB789649FF916" authority="(Kraatz, 1858)" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[595,992,1687,1715]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D240477F72FB7896451F917" bold="true" box="[595,814,1687,1714]" italics="true" pageId="33" pageNumber="34">Cholovocera attae</emphasis>
(
<bibRefCitation id="8B3B4B794D240477F642B78664A7F916" author="Kraatz G." box="[830,984,1688,1715]" pageId="33" pageNumber="34" pagination="123 - 148" refId="ref39933" refString="Kraatz G. 1858. Beitrag zur Kaferfauna Griechenlands. Staphylinidae (Schlufs), Trichopterygia, Histeridae, Phalacridae, Nitidularia, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrrhii. Berliner Entomologische Zeitschrift 2: 123 - 148." type="journal article" year="1858">Kraatz, 1858</bibRefCitation>
)
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<subSubSection id="A7B065034D240477F750B7A26511F8BD" pageId="33" pageNumber="34" type="description">
<paragraph id="EF1536884D240477F750B7A26378F973" blockId="33.[556,1031,1687,1750]" box="[556,1031,1724,1750]" pageId="33" pageNumber="34">
<figureCitation id="77912A0D4D240477F750B7A265F5F973" box="[556,650,1724,1750]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[264,1318,1324,1816]" captionTargetId="figure-432@10.[187,1400,1277,1877]" captionTargetPageId="10" captionText="Fig. 4. Prosternal processes. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845)." figureDoi="http://doi.org/10.5281/zenodo.10424520" httpUri="https://zenodo.org/record/10424520/files/figure.png" pageId="33" pageNumber="34">Figs 4B</figureCitation>
, 
<figureCitation id="77912A0D4D240477F7EBB7A265C4F973" box="[663,699,1724,1750]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[303,1293,291,1782]" captionTargetId="figure-70@18.[247,1340,265,1810]" captionTargetPageId="18" captionText="Fig. 9. Known geographical distribution of the species ofCholovocera Victor, 1838.A. Ch. formiceticola (Rosenhauer, 1856) (red dots) and Ch. gallica (Schaufuss, 1876) (yellow dots). B. Ch. punctata (Märkel, 1845) (red dots) and Ch. occulta sp. nov. (yellow dots). C. Ch. balcanica (Karaman, 1936) (red dots), Ch. attae (Kraatz, 1858) (yellow dots), Ch. formicaria (dark blue dots) and Ch. afghana Johnson, 1977 (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10424533" httpUri="https://zenodo.org/record/10424533/files/figure.png" pageId="33" pageNumber="34">9C</figureCitation>
, 
<figureCitation id="77912A0D4D240477F7B4B7A26584F973" box="[712,763,1724,1750]" captionStart="Fig" captionStartId="35.[189,232,1963,1989]" captionTargetBox="[294,1312,465,1855]" captionTargetId="figure-99@35.[269,1318,454,1909]" captionTargetPageId="35" captionText="Fig. 10. Habitus, dorsal view. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♂. C. Ch. balcanica (Karaman, 1936), ♂. D. Ch. gallica (Schaufuss, 1876), ♂. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10424535" httpUri="https://zenodo.org/record/10424535/files/figure.png" pageId="33" pageNumber="34">10B</figureCitation>
, 
<figureCitation id="77912A0D4D240477F67BB7A26445F973" box="[775,826,1724,1750]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[341,1310,800,1791]" captionTargetId="figure-179@42.[221,1367,721,1840]" captionTargetPageId="42" captionText="Fig. 12. Terminal female antennomeres. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424539" httpUri="https://zenodo.org/record/10424539/files/figure.png" pageId="33" pageNumber="34">12B</figureCitation>
, 
<figureCitation id="77912A0D4D240477F63BB7A26403F973" box="[839,892,1724,1750]" captionStart="Fig" captionStartId="44.[189,232,1928,1954]" captionTargetBox="[247,1330,760,1834]" captionTargetId="figure-280@44.[217,1371,719,1875]" captionTargetPageId="44" captionText="Fig. 13. Terminal male antennomeres.A. Cholovocera attae (Kraatz, 1858). B. Ch. balcánica(Karaman, 1936). C. Ch. gallica (Schaufuss, 1876). D. Ch. formicaria Victor, 1838. E. Ch. formiceticola (Rosenhauer, 1856). F. Ch. occulta sp. nov. G. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424541" httpUri="https://zenodo.org/record/10424541/files/figure.png" pageId="33" pageNumber="34">13A</figureCitation>
, 
<figureCitation id="77912A0D4D240477F6F4B7A264A2F973" box="[904,989,1724,1750]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="33" pageNumber="34">14B–C</figureCitation>
, 
<figureCitation id="77912A0D4D240477F696B7A26378F973" box="[1002,1031,1724,1750]" captionStart="Fig" captionStartId="55.[189,232,975,1001]" captionTargetBox="[475,1078,315,913]" captionTargetId="figure-56@55.[426,1132,265,950]" captionTargetPageId="55" captionText="Fig. 19. Aedeagus and paramere of Cholovocera attae (Kraatz, 1858) (Greece). A–B. Ventral views. C. Lateral view.. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424555" httpUri="https://zenodo.org/record/10424555/files/figure.png" pageId="33" pageNumber="34">19</figureCitation>
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<emphasis id="DDDEEA9A4D240477F5C1B7E066F3F8BD" box="[189,396,1790,1816]" italics="true" pageId="33" pageNumber="34">Choluocera Attae</emphasis>
<bibRefCitation id="8B3B4B794D240477F4E8B7E06515F8BD" author="Kraatz G." box="[404,618,1789,1816]" pageId="33" pageNumber="34" pagination="123 - 148" refId="ref39933" refString="Kraatz G. 1858. Beitrag zur Kaferfauna Griechenlands. Staphylinidae (Schlufs), Trichopterygia, Histeridae, Phalacridae, Nitidularia, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrrhii. Berliner Entomologische Zeitschrift 2: 123 - 148." type="journal article" year="1858">Kraatz, 1858: 140</bibRefCitation>
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<emphasis id="DDDEEA9A4D240477F5C1B6216604F8FC" box="[189,379,1855,1881]" italics="true" pageId="33" pageNumber="34">Coluocera attae</emphasis>
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– 
<treatmentCitation id="6E0B10994D240477F4E4B6216462F8FF" author="Gemminger M. &amp; Harold B. de" box="[408,797,1855,1882]" page="905" pageId="33" pageNumber="34" year="1868">
<bibRefCitation id="8B3B4B794D240477F4E4B6216462F8FF" author="Gemminger M. &amp; Harold B. de" box="[408,797,1855,1882]" pageId="33" pageNumber="34" refId="ref39469" refString="Gemminger M. &amp; Harold B. de 1868. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. III. Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydidae, Rhysodidae, Cucujidae, Cryptophagidae, Derodontidae, Latrididae, Othnidae, Mycetophagidae, Thorictidae, Dermestidae, Byrrhidae, Georyssidae, Parnidae, Heteroceridae, Lucanidae. Sumptu E. H. Gummi, Monachii, Paris, London." type="book" year="1868">Gemminger &amp; Harold 1868: 905</bibRefCitation>
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<treatmentCitationGroup id="CFBA11A64D240477F5C1B67D650CF8D8" box="[189,627,1891,1918]" pageId="33" pageNumber="34">
<taxonomicName id="28AA4D0B4D240477F5C1B67D66F4F8DB" ID-CoL="3SKC" authority="Attae" authorityName="Attae" box="[189,395,1891,1918]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D240477F5C1B67D66F4F8DB" box="[189,395,1891,1918]" italics="true" pageId="33" pageNumber="34">Colovocera Attae</emphasis>
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– 
<treatmentCitation id="6E0B10994D240477F4D4B67A6510F8D8" author="Lucas H." box="[424,623,1891,1918]" page="239" pageId="33" pageNumber="34" year="1874">
<bibRefCitation id="8B3B4B794D240477F4D4B67A6510F8D8" author="Lucas H." box="[424,623,1891,1918]" pageId="33" pageNumber="34" pagination="239" refId="ref40549" refString="Lucas H. 1874. Note. Seance 25 Novembre 1874. Annales de la Societe entomologique de France (Bulletin des Seances et Bulletin bibliographique) 5 (4): 239." type="journal article" year="1874">Lucas 1874: 239</bibRefCitation>
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<taxonomicName id="28AA4D0B4D240477F5C1B69966C2F804" authorityName="Victor" authorityYear="1838" box="[189,445,1927,1953]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D240477F5C1B69966C2F804" box="[189,445,1927,1953]" italics="true" pageId="33" pageNumber="34">Coluocera formicaria</emphasis>
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– 
<treatmentCitation id="6E0B10994D240477F4A7B69965D7F804" author="Reitter E." box="[475,680,1927,1954]" page="301" pageId="33" pageNumber="34" year="1875">
<bibRefCitation id="8B3B4B794D240477F4A7B69965D7F804" author="Reitter E." box="[475,680,1927,1954]" pageId="33" pageNumber="34" pagination="297 - 340" refId="ref41208" refString="Reitter E. 1875. Revision der europaischen Lathridiidae. Entomologische Zeitung 36: 297 - 340." type="journal article" year="1875">Reitter 1875: 301</bibRefCitation>
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(in part).
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<taxonomicName id="28AA4D0B4D240477F5C1B6B566B5F860" authorityName="Motschulsky" authorityYear="1838" box="[189,458,1963,1989]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D240477F5C1B6B566B5F860" box="[189,458,1963,1989]" italics="true" pageId="33" pageNumber="34">Colovocera formicaria</emphasis>
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– 
<treatmentCitation id="6E0B10994D240477F494B6B565D3F860" author="Belon M. - J." box="[488,684,1963,1989]" page="192" pageId="33" pageNumber="34" year="1879">
<bibRefCitation id="8B3B4B794D240477F494B6B565D3F860" author="Belon M. - J." box="[488,684,1963,1989]" pageId="33" pageNumber="34" pagination="157 - 365" refId="ref38155" refString="Belon M. - J. 1879. Famille des Lathridiens, 1 re partie. In: Mulsant E. (ed.) Histoire naturelle des coleopteres de France. Annales de la Societe linneenne de Lyon, n. s. 26: 157 - 365." type="book chapter" year="1879">Belon 1879: 192</bibRefCitation>
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(in part).
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</paragraph>
<paragraph id="EF1536884D240477F5C1B6D065D3F84D" blockId="33.[189,800,1855,2025]" box="[189,684,1998,2025]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="CFBA11A64D240477F5C1B6D065D3F84D" box="[189,684,1998,2025]" pageId="33" pageNumber="34">
<emphasis id="DDDEEA9A4D240477F5C1B6D06646F84D" box="[189,313,1998,2024]" italics="true" pageId="33" pageNumber="34">Colnocera</emphasis>
<taxonomicName id="28AA4D0B4D240477F43CB6D066C8F84D" authorityName="Kraatz" authorityYear="1858" box="[320,439,1998,2024]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="attae">
[sic] 
<emphasis id="DDDEEA9A4D240477F400B6D166C8F84D" box="[380,439,1999,2024]" italics="true" pageId="33" pageNumber="34">attae</emphasis>
</taxonomicName>
– 
<treatmentCitation id="6E0B10994D240477F4AFB6D065D7F84D" author="Walker J. J." box="[467,680,1998,2024]" page="181" pageId="33" pageNumber="34" year="1888">
<bibRefCitation id="8B3B4B794D240477F4AFB6D065D7F84D" author="Walker J. J." box="[467,680,1998,2024]" pageId="33" pageNumber="34" pagination="175 - 187" refId="ref42952" refString="Walker J. J. 1888. A year's insect-hunting at Gibraltar. Entomologist's Monthly Magazine 24: 175 - 187." type="journal article" year="1888">Walker 1888: 181</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF1536884D270474F5C1B017636EFE86" blockId="34.[189,1041,265,292]" box="[189,1041,265,292]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="CFBA11A64D270474F5C1B017636EFE86" box="[189,1041,265,292]" pageId="34" pageNumber="35">
<taxonomicName id="28AA4D0B4D270474F5C1B01766E8FE86" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[189,407,265,291]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F5C1B01766E8FE86" box="[189,407,265,291]" italics="true" pageId="34" pageNumber="35">Cholovocera attae</emphasis>
</taxonomicName>
– 
<treatmentCitation id="6E0B10994D270474F4C8B01765F7FE86" author="Rucker H. W." box="[436,648,265,291]" page="143" pageId="34" pageNumber="35" year="1980">
<bibRefCitation id="8B3B4B794D270474F4C8B01765F7FE86" author="Rucker H. W." box="[436,648,265,291]" pageId="34" pageNumber="35" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker 1980: 143</bibRefCitation>
</treatmentCitation>
, fig. 22 — 
<treatmentCitation id="6E0B10994D270474F676B0176374FE86" author="Audisio P. &amp; Canepari C. &amp; De Biase A. &amp; Poggi R. &amp; Ratti E. &amp; Zampetti M. F." box="[778,1035,265,292]" page="9" pageId="34" pageNumber="35" year="1995">
<bibRefCitation id="8B3B4B794D270474F676B0176374FE86" author="Audisio P. &amp; Canepari C. &amp; De Biase A. &amp; Poggi R. &amp; Ratti E. &amp; Zampetti M. F." box="[778,1035,265,292]" pageId="34" pageNumber="35" pagination="1 - 19" refId="ref38021" refString="Audisio P., Canepari C., De Biase A., Poggi R., Ratti E. &amp; Zampetti M. F. 1995. Coleoptera Polyphaga XI (Clavicornia II). In: Minelli A., Ruffo S. &amp; La Posta S. (eds) Checklist delle specie della fauna italiana 56: 1 - 19. Calderini, Bologna." type="journal article" year="1995">
Audisio 
<emphasis id="DDDEEA9A4D270474F612B01464D6FE86" box="[878,937,265,291]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
1995: 9
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D270474F5C1B05466E1FE5B" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="EF1536884D270474F5C1B05466B6FEC1" blockId="34.[189,457,330,356]" box="[189,457,330,356]" pageId="34" pageNumber="35">
<heading id="B45D81E44D270474F5C1B05466B6FEC1" bold="true" box="[189,457,330,356]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DDDEEA9A4D270474F5C1B05466B6FEC1" bold="true" box="[189,457,330,356]" pageId="34" pageNumber="35">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D270474F5C1B06766E1FE5B" blockId="34.[189,1399,377,510]" pageId="34" pageNumber="35">
<taxonomicName id="28AA4D0B4D270474F5C1B06766E7FE36" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[189,408,377,403]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F5C1B06766E7FE36" box="[189,408,377,403]" italics="true" pageId="34" pageNumber="35">Cholovocera attae</emphasis>
</taxonomicName>
and 
<taxonomicName id="28AA4D0B4D270474F4AAB0676521FE36" authority="Delgado &amp; Palma, 2023" authorityName="Delgado &amp; Palma" authorityYear="2023" box="[470,606,377,403]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="DDDEEA9A4D270474F4AAB0676521FE36" box="[470,606,377,403]" italics="true" pageId="34" pageNumber="35">Ch. occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6ED57E14D270474F71AB06465BDFE31" box="[614,706,378,404]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
are the smallest species in the genus. Size and short wide legs would separate 
<taxonomicName id="28AA4D0B4D270474F4C8B083655CFE13" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[436,547,413,439]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F4C8B083655CFE13" box="[436,547,413,439]" italics="true" pageId="34" pageNumber="35">Ch. attae</emphasis>
</taxonomicName>
from its sympatric 
<taxonomicName id="28AA4D0B4D270474F66FB08364C4FE12" box="[787,955,413,439]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D270474F66FB08364C4FE12" box="[787,955,413,439]" italics="true" pageId="34" pageNumber="35">Ch. balcanica</emphasis>
</taxonomicName>
. However, the most useful character to distinguish 
<taxonomicName id="28AA4D0B4D270474F415B0DE66A7FE7F" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[361,472,448,474]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F415B0DE66A7FE7F" box="[361,472,448,474]" italics="true" pageId="34" pageNumber="35">Ch. attae</emphasis>
</taxonomicName>
from 
<taxonomicName id="28AA4D0B4D270474F759B0DE65B2FE7F" box="[549,717,448,474]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D270474F759B0DE65B2FE7F" box="[549,717,448,474]" italics="true" pageId="34" pageNumber="35">Ch. balcanica</emphasis>
</taxonomicName>
and from all the other species is the morphology of the aedeagus (
<figureCitation id="77912A0D4D270474F445B0FD66F0FE5B" box="[313,399,483,510]" captionStart="Fig" captionStartId="55.[189,232,975,1001]" captionTargetBox="[475,1078,315,913]" captionTargetId="figure-56@55.[426,1132,265,950]" captionTargetPageId="55" captionText="Fig. 19. Aedeagus and paramere of Cholovocera attae (Kraatz, 1858) (Greece). A–B. Ventral views. C. Lateral view.. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424555" httpUri="https://zenodo.org/record/10424555/files/figure.png" pageId="34" pageNumber="35">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D270474F5C1B33B64FFFC9D" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="EF1536884D270474F5C1B33B6613FD9A" blockId="34.[189,364,549,575]" box="[189,364,549,575]" pageId="34" pageNumber="35">
<heading id="B45D81E44D270474F5C1B33B6613FD9A" bold="true" box="[189,364,549,575]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DDDEEA9A4D270474F5C1B33B6613FD9A" bold="true" box="[189,364,549,575]" pageId="34" pageNumber="35">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D270474F5C1B34A622FFDCB" blockId="34.[189,1360,596,622]" box="[189,1360,596,622]" pageId="34" pageNumber="35">
<taxonomicName id="28AA4D0B4D270474F5C1B34A66E8FDCB" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[189,407,596,622]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F5C1B34A66E8FDCB" box="[189,407,596,622]" italics="true" pageId="34" pageNumber="35">Cholovocera attae</emphasis>
</taxonomicName>
: 
<typeStatus id="3011882A4D270474F4DAB34A656DFDCB" box="[422,530,596,622]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
male and 
<typeStatus id="3011882A4D270474F7F7B34A6456FDCB" box="[651,809,596,622]" pageId="34" pageNumber="35" type="paralectotype">paralectotype</typeStatus>
female in the Kraatz Collection, held in 
<collectionCode id="89BBAE4D4D270474F07BB34A6233FDCB" box="[1287,1356,596,622]" pageId="34" pageNumber="35">SDEI</collectionCode>
.
</paragraph>
<materialsCitation id="5FC23CD54D270474F596B38B625EFD76" collectionCode="SDEI" country="Greece" determinerName="Rucker" location="Kraatz Collection" pageId="34" pageNumber="35" specimenCode="SDEI 11869" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<paragraph id="EF1536884D270474F596B38B6527FD15" blockId="34.[189,1313,661,723]" box="[234,600,661,688]" pageId="34" pageNumber="35">
<emphasis id="DDDEEA9A4D270474F596B38B6527FD15" bold="true" box="[234,600,661,688]" pageId="34" pageNumber="35">
<typeStatus id="3011882A4D270474F596B38B6619FD0A" box="[234,358,661,687]" pageId="34" pageNumber="35" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="28AA4D0B4D270474F4F1B38B6527FD15" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[397,600,661,688]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F4F1B38B6527FD15" bold="true" box="[397,600,661,688]" italics="true" pageId="34" pageNumber="35">Colovocera attae</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="EF1536884D270474F5C1B3A7625EFD76" blockId="34.[189,1313,661,723]" box="[189,1313,697,723]" pageId="34" pageNumber="35">
<collectingCountry id="97BD76184D270474F5C1B3A76650FD76" box="[189,303,697,723]" name="Greece" pageId="34" pageNumber="35">GREECE</collectingCountry>
• 
<specimenCount id="F9ACFD014D270474F434B3A7660BFD76" box="[328,372,697,723]" count="1" pageId="34" pageNumber="35" type="male">
1 
<emphasis id="DDDEEA9A4D270474F422B3A7660BFD76" bold="true" box="[350,372,697,723]" pageId="34" pageNumber="35">♂</emphasis>
</specimenCount>
; “Graecia”; 
<location id="EA7560534D270474F774B3A765A3FD76" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F774B3A765A3FD76" box="[520,732,697,723]" country="Greece" name="Kraatz Collection" pageId="34" pageNumber="35">Kraatz Collection</location>
; 
<specimenCode id="BF0C9EF34D270474F79BB3A76403FD76" box="[743,892,697,723]" pageId="34" pageNumber="35">SDEI 11869</specimenCode>
. Designated by 
<bibRefCitation id="8B3B4B794D270474F14BB3A76262FD76" author="Rucker W. H." box="[1079,1309,697,723]" pageId="34" pageNumber="35" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">
<determinerName id="1AC9DA4C4D270474F14BB3A763F3FD76" box="[1079,1164,697,723]" pageId="34" pageNumber="35">Rucker</determinerName>
(2011a: 13)
</bibRefCitation>
.
</paragraph>
</materialsCitation>
<materialsCitation id="5FC23CD54D270474F596B3E46403FC9D" collectionCode="SDEI" country="Greece" location="Kraatz Collection" pageId="34" pageNumber="35" specimenCode="SDEI 11870" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<paragraph id="EF1536884D270474F596B3E465F6FCB0" blockId="34.[189,896,762,824]" box="[234,649,762,789]" pageId="34" pageNumber="35">
<emphasis id="DDDEEA9A4D270474F596B3E465F6FCB0" bold="true" box="[234,649,762,789]" pageId="34" pageNumber="35">
<typeStatus id="3011882A4D270474F596B3E466E8FCB1" box="[234,407,762,788]" pageId="34" pageNumber="35" type="paralectotype">Paralectotype</typeStatus>
of 
<taxonomicName id="28AA4D0B4D270474F4C2B3E465F6FCB0" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[446,649,762,789]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D270474F4C2B3E465F6FCB0" bold="true" box="[446,649,762,789]" italics="true" pageId="34" pageNumber="35">Colovocera attae</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="EF1536884D270474F5C1B20064FFFC9D" blockId="34.[189,896,762,824]" box="[189,896,798,824]" pageId="34" pageNumber="35">
<collectingCountry id="97BD76184D270474F5C1B2006650FC9D" box="[189,303,798,824]" name="Greece" pageId="34" pageNumber="35">GREECE</collectingCountry>
• 
<specimenCount id="F9ACFD014D270474F434B200660BFC9D" box="[328,372,798,824]" count="1" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
; “Graecia”; 
<location id="EA7560534D270474F774B20065A3FC9D" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F774B20065A3FC9D" box="[520,732,798,824]" country="Greece" name="Kraatz Collection" pageId="34" pageNumber="35">Kraatz Collection</location>
; 
<specimenCode id="BF0C9EF34D270474F79BB2006403FC9D" box="[743,892,798,824]" collectionCode="SDEI" pageId="34" pageNumber="35">SDEI 11870</specimenCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="A7B065034D270474F596B2416304FBC9" pageId="34" pageNumber="35" type="discussion">
<paragraph id="EF1536884D270474F596B241664FFCDC" blockId="34.[189,1399,863,1067]" box="[234,304,863,889]" pageId="34" pageNumber="35">
<emphasis id="DDDEEA9A4D270474F596B241664FFCDC" bold="true" box="[234,304,863,889]" pageId="34" pageNumber="35">Notes</emphasis>
</paragraph>
<paragraph id="EF1536884D270474F5C1B29D6568FB8E" blockId="34.[189,1399,863,1067]" pageId="34" pageNumber="35">
As it can be seen in 
<bibRefCitation id="8B3B4B794D270474F4DEB29D653BFC38" author="Rucker W. H." box="[418,580,899,925]" pageId="34" pageNumber="35" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker 2011a</bibRefCitation>
(fig. 12), there is a label reading “ 
<typeStatus id="3011882A4D270474F6B5B29D6337FC38" box="[969,1096,899,925]" pageId="34" pageNumber="35" type="holotype">Holotypus</typeStatus>
” attached to the specimen that Rücker designated as the 
<typeStatus id="3011882A4D270474F756B2B865E6FC65" box="[554,665,934,960]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
. However, this specimen cannot be regarded as the 
<typeStatus id="3011882A4D270474F06CB2B86208FC65" box="[1296,1399,934,960]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
because it was not designated in the original description, which included more than 
<specimenCount id="F9ACFD014D270474F1E7B2D46240FC46" box="[1179,1343,969,996]" count="1" pageId="34" pageNumber="35" type="generic">one specimen</specimenCount>
, i.e., 
<typeStatus id="3011882A4D270474F5C1B2F36658FBA2" box="[189,295,1005,1031]" pageId="34" pageNumber="35" type="syntype">syntypes</typeStatus>
. Examining the printed 
<typeStatus id="3011882A4D270474F73DB2F365C4FBA2" box="[577,699,1005,1031]" pageId="34" pageNumber="35" type="holotype">Holotypus</typeStatus>
label, we conclude that it was added at a later date than the description by 
<bibRefCitation id="8B3B4B794D270474F410B50F656CFB8E" author="Kraatz G." box="[364,531,1040,1067]" pageId="34" pageNumber="35" pagination="123 - 148" refId="ref39933" refString="Kraatz G. 1858. Beitrag zur Kaferfauna Griechenlands. Staphylinidae (Schlufs), Trichopterygia, Histeridae, Phalacridae, Nitidularia, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrrhii. Berliner Entomologische Zeitschrift 2: 123 - 148." type="journal article" year="1858">Kraatz (1858)</bibRefCitation>
.
</paragraph>
<paragraph id="EF1536884D270474F5C1B54C6304FBC9" blockId="34.[189,1147,1106,1132]" box="[189,1147,1106,1132]" pageId="34" pageNumber="35">
The same comment given below under the 
<typeStatus id="3011882A4D270474F7CBB54C6459FBC9" box="[695,806,1106,1132]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
, applies to the 
<typeStatus id="3011882A4D270474F6A9B54C6308FBC9" box="[981,1143,1106,1132]" pageId="34" pageNumber="35" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D270474F5C1B58D66A0F962" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="EF1536884D270474F5C1B58D6542FB08" blockId="34.[189,573,1171,1197]" box="[189,573,1171,1197]" pageId="34" pageNumber="35">
<heading id="B45D81E44D270474F5C1B58D6542FB08" bold="true" box="[189,573,1171,1197]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DDDEEA9A4D270474F5C1B58D6542FB08" bold="true" box="[189,573,1171,1197]" pageId="34" pageNumber="35">Additional material, non-types</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D270474F5C1B5DC6486FACF" blockId="34.[189,1399,1216,1386]" pageId="34" pageNumber="35">
<materialsCitation id="5FC23CD54D270474F5C1B5DC6517FA86" collectingDate="2013-05-10" collectionCode="NHMW" collectorName="Schuh &amp; Mountain" country="Greece" elevation="1380" latitude="37.679585" location="Vitina" longLatPrecision="1" longitude="22.247972" municipality="Menalon mts." pageId="34" pageNumber="35" specimenCount="2" specimenCount-female="1" stateProvince="Peloponnisos">
<collectingCountry id="97BD76184D270474F5C1B5DC6650FB79" box="[189,303,1218,1244]" name="Greece" pageId="34" pageNumber="35">GREECE</collectingCountry>
– 
<emphasis id="DDDEEA9A4D270474F42EB5DC6692FB79" bold="true" box="[338,493,1218,1244]" pageId="34" pageNumber="35">
<collectingRegion id="2D6EF86A4D270474F42EB5DC6692FB79" box="[338,493,1218,1244]" country="Greece" name="Peloponnisos" pageId="34" pageNumber="35">Peloponnese</collectingRegion>
</emphasis>
• 2 ♂♂, 
<specimenCount id="F9ACFD014D270474F720B5DC65F5FB7A" box="[604,650,1218,1247]" count="1" pageId="34" pageNumber="35" type="female">
1 
<emphasis id="DDDEEA9A4D270474F708B5DD65F5FB7A" bold="true" box="[628,650,1219,1247]" pageId="34" pageNumber="35">♀</emphasis>
</specimenCount>
; 
<collectingMunicipality id="0F71ACF24D270474F7E7B5DC643EFB78" box="[667,833,1218,1245]" pageId="34" pageNumber="35">Menalon mts.</collectingMunicipality>
, 
<quantity id="28529B6D4D270474F633B5DC64F3FB79" box="[847,908,1218,1245]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="km" value="5.0">5 km</quantity>
E. 
<location id="EA7560534D270474F6C4B5DC637CFB79" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F6C4B5DC637CFB79" box="[952,1027,1218,1244]" country="Greece" latitude="37.679585" longLatPrecision="1" longitude="22.247972" municipality="Menalon mts." name="Vitina" pageId="34" pageNumber="35" stateProvince="Peloponnisos">Vitina</location>
; 
<geoCoordinate id="8A9E504F4D270474F16DB5DE63C3FB78" box="[1041,1212,1216,1245]" degrees="37" direction="north" minutes="40" orientation="latitude" pageId="34" pageNumber="35" precision="1" seconds="46.5" value="37.679585">37º40′46.5″ N</geoCoordinate>
, 
<geoCoordinate id="8A9E504F4D270474F1B5B5DE620EFB78" box="[1225,1393,1216,1245]" degrees="22" direction="east" minutes="14" orientation="longitude" pageId="34" pageNumber="35" precision="1" seconds="52.7" value="22.247972">22º14′52.7″ E</geoCoordinate>
; 
<elevation id="6487D1BB4D270474F5C1B5F86626FB5A" box="[189,345,1253,1280]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="34" pageNumber="35" unit="m" value="1380.0">
<quantity id="28529B6D4D270474F5C1B5F86667FAA5" box="[189,280,1254,1280]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="34" pageNumber="35" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
; 
<date id="9B1410484D270474F415B5F86573FAA5" box="[361,524,1254,1280]" pageId="34" pageNumber="35" value="2013-05-10">
<collectingDate id="8B50E9A04D270474F415B5F86573FAA5" box="[361,524,1254,1280]" pageId="34" pageNumber="35" value="2013-05-10">10 May 2013</collectingDate>
</date>
; 
<collectorName id="425F535E4D270474F766B5FB651CFB5A" box="[538,611,1253,1279]" pageId="34" pageNumber="35">Schuh</collectorName>
leg.; “Almweide” [
<collectorName id="425F535E4D270474F631B5FB64BDFB5A" box="[845,962,1253,1279]" pageId="34" pageNumber="35">Mountain</collectorName>
meadow], [
<specimenCount id="F9ACFD014D270474F12CB5F8638AFB5A" box="[1104,1269,1253,1280]" count="1" pageId="34" pageNumber="35" type="generic">one specimen</specimenCount>
associated with a 
<taxonomicName id="28AA4D0B4D270474F46CB4146619FA86" authorityName="Forel" authorityYear="1890" box="[272,358,1290,1315]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D270474F46CB4146619FA86" box="[272,358,1290,1315]" italics="true" pageId="34" pageNumber="35">Messor</emphasis>
</taxonomicName>
worker 
<vernacularName id="61A946A64D270474F4B6B417668CFA86" authorityName="Latreille" authorityYear="1802" box="[458,499,1289,1315]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">ant</vernacularName>
]; 
<collectionCode id="89BBAE4D4D270474F774B4176517FA86" box="[520,616,1289,1315]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35" type="Museum">NHMW</collectionCode>
</materialsCitation>
– 
<materialsCitation id="5FC23CD54D270474F7F5B4166625FAE2" collectingDate="1984-04-13" collectionCode="NHMW" collectorName="H. Fulscher" country="Greece" location="Vai-Itanos" municipality="West Crete" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Kriti">
<emphasis id="DDDEEA9A4D270474F7F5B41665B1FA87" bold="true" box="[649,718,1288,1314]" pageId="34" pageNumber="35">
<collectingRegion id="2D6EF86A4D270474F7F5B41665B1FA87" box="[649,718,1288,1314]" country="Greece" name="Kriti" pageId="34" pageNumber="35">Crete</collectingRegion>
</emphasis>
• 1 ♂; 
<collectingMunicipality id="0F71ACF24D270474F65AB41764D3FA86" box="[806,940,1289,1315]" pageId="34" pageNumber="35">West Crete</collectingMunicipality>
, 
<location id="EA7560534D270474F6C4B417634BFA86" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F6C4B417634BFA86" box="[952,1076,1289,1315]" country="Greece" municipality="West Crete" name="Vai-Itanos" pageId="34" pageNumber="35" stateProvince="Kriti">Vai-Itanos</location>
; 
<date id="9B1410484D270474F13DB41763A0FA86" box="[1089,1247,1289,1315]" pageId="34" pageNumber="35" value="1984-04-13">
<collectingDate id="8B50E9A04D270474F13DB41763A0FA86" box="[1089,1247,1289,1315]" pageId="34" pageNumber="35" value="1984-04-13">13 Apr. 1984</collectingDate>
</date>
; 
<collectorName id="425F535E4D270474F190B4176209FA86" box="[1260,1398,1289,1315]" pageId="34" pageNumber="35">H. Fülscher</collectorName>
leg.; 
<collectionCode id="89BBAE4D4D270474F586B4336625FAE2" box="[250,346,1325,1351]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35" type="Museum">NHMW</collectionCode>
</materialsCitation>
• 
<materialsCitation id="5FC23CD54D270474F409B4326599FACF" collectingDate="2010-06-02" collectionCode="NHMW" collectorName="Schuh" country="Greece" county="West Crete" elevation="1150" location="Hania" municipality="Levka Ori" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Kriti">
4 ♂♂, 
<specimenCount id="F9ACFD014D270474F4B5B432668AFAEC" box="[457,501,1324,1353]" count="1" pageId="34" pageNumber="35" type="female">
1 
<emphasis id="DDDEEA9A4D270474F4A3B433668AFAEC" bold="true" box="[479,501,1325,1353]" pageId="34" pageNumber="35">♀</emphasis>
</specimenCount>
; 
<collectingCounty id="06744E044D270474F779B43365F4FAE3" box="[517,651,1324,1351]" pageId="34" pageNumber="35">West Crete</collectingCounty>
, 
<collectingMunicipality id="0F71ACF24D270474F7E4B432646CFAE3" box="[664,787,1324,1350]" pageId="34" pageNumber="35">Levka Ori</collectingMunicipality>
, r[oa]d. 
<location id="EA7560534D270474F604B43264C1FAE3" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F604B43264C1FAE3" box="[888,958,1324,1350]" country="Greece" county="West Crete" municipality="Levka Ori" name="Hania" pageId="34" pageNumber="35" stateProvince="Kriti">Hania</location>
to 
<location id="EA7560534D270474F69AB432633BFAE3" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F69AB432633BFAE3" box="[998,1092,1324,1350]" country="Greece" county="West Crete" municipality="Levka Ori" name="Omalos" pageId="34" pageNumber="35" stateProvince="Kriti">Omalos</location>
, 
<locationDeviation id="FEA72FAA4D270474F12EB432620DFAE3" box="[1106,1394,1324,1351]" location="Omalos" pageId="34" pageNumber="35">
<quantity id="28529B6D4D270474F12EB43263DBFAE3" box="[1106,1188,1324,1351]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="35" unit="km" value="2.5">2.5 km</quantity>
NNE of Omalos
</locationDeviation>
; 
<elevation id="6487D1BB4D270474F5C2B4516629FACC" box="[190,342,1359,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="34" pageNumber="35" unit="m" value="1150.0">
<quantity id="28529B6D4D270474F5C2B4516669FACF" box="[190,278,1359,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="34" pageNumber="35" unit="m" value="1150.0">1150 m</quantity>
a.s.l.
</elevation>
; 
<date id="9B1410484D270474F41FB44E6690FACF" box="[355,495,1360,1386]" pageId="34" pageNumber="35" value="2010-06-02">
<collectingDate id="8B50E9A04D270474F41FB44E6690FACF" box="[355,495,1360,1386]" pageId="34" pageNumber="35" value="2010-06-02">2 Jun. 2010</collectingDate>
</date>
; 
<collectorName id="425F535E4D270474F487B451653BFACC" box="[507,580,1359,1385]" pageId="34" pageNumber="35">Schuh</collectorName>
leg.; 
<collectionCode id="89BBAE4D4D270474F7FAB44E6599FACF" box="[646,742,1360,1386]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35" type="Museum">NHMW</collectionCode>
</materialsCitation>
• 
<materialsCitation id="5FC23CD54D270474F783B44E648AFACF" box="[767,1013,1357,1386]" pageId="34" pageNumber="35">
2 
<emphasis id="DDDEEA9A4D270474F668B44E6455FACF" bold="true" box="[788,810,1360,1386]" pageId="34" pageNumber="35">♂</emphasis>
♂; Crete; 
<collectionCode id="89BBAE4D4D270474F6E2B44E648AFACF" box="[926,1013,1360,1386]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="34" pageNumber="35">NMPC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="EF1536884D270474F5C1B48C6626FA6A" blockId="34.[189,1398,1423,1487]" pageId="34" pageNumber="35">
<materialsCitation id="5FC23CD54D270474F5C1B48C6629FA6A" collectingDate="1975-05-02" collectorName="Besuchet &amp; Lobl" country="Turkey" location="Gocek" pageId="34" pageNumber="35" specimenCount="2" specimenCount-female="2" stateProvince="Mugla">
<collectingCountry id="97BD76184D270474F5C1B48C664CFA09" box="[189,307,1426,1452]" name="Turkey" pageId="34" pageNumber="35">TURKEY</collectingCountry>
– 
<emphasis id="DDDEEA9A4D270474F42FB48F65E7FA0E" bold="true" box="[339,664,1425,1451]" pageId="34" pageNumber="35">Western Anatolia, Aegean</emphasis>
• 1 ♂, 
<specimenCount id="F9ACFD014D270474F791B48F644FFA0B" box="[749,816,1425,1454]" count="2" pageId="34" pageNumber="35" type="female">
2 
<emphasis id="DDDEEA9A4D270474F678B48C644FFA0B" bold="true" box="[772,816,1426,1454]" pageId="34" pageNumber="35">♀♀</emphasis>
</specimenCount>
; 
<collectingRegion id="2D6EF86A4D270474F63DB48F64EEFA0E" box="[833,913,1425,1451]" country="Turkey" name="Mugla" pageId="34" pageNumber="35">Muğla</collectingRegion>
, 
<location id="EA7560534D270474F6E1B48F6492FA0E" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F6E1B48F6492FA0E" box="[925,1005,1425,1451]" country="Turkey" name="Gocek" pageId="34" pageNumber="35" stateProvince="Mugla">Göcek</location>
; 
<date id="9B1410484D270474F685B48F63F3FA09" box="[1017,1164,1425,1452]" pageId="34" pageNumber="35" value="1975-05-02">
<collectingDate id="8B50E9A04D270474F685B48F63F3FA09" box="[1017,1164,1425,1452]" pageId="34" pageNumber="35" value="1975-05-02">2 May 1975</collectingDate>
</date>
; 
<collectorName id="425F535E4D270474F1E5B48F627AFA0E" box="[1177,1285,1425,1451]" pageId="34" pageNumber="35">Besuchet</collectorName>
and 
<collectorName id="425F535E4D270474F043B48F6209FA0E" box="[1343,1398,1425,1451]" pageId="34" pageNumber="35">Löbl</collectorName>
leg.; 
<collectionCode id="89BBAE4D4D270474F584B4AB6629FA6A" box="[248,342,1461,1487]" pageId="34" pageNumber="35">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="EF1536884D270474F596B4E865DBF9B5" blockId="34.[189,1398,1526,1623]" box="[234,676,1526,1553]" pageId="34" pageNumber="35">
<taxonomicName id="28AA4D0B4D270474F596B4E865DBF9B5" authority="Wasmann, 1896" authorityName="Wasmann" authorityYear="1896" box="[234,676,1526,1553]" class="Insecta" family="Curculionidae" genus="Reitteria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="escherichi">
<emphasis id="DDDEEA9A4D270474F596B4E865DBF9B5" bold="true" box="[234,676,1526,1553]" pageId="34" pageNumber="35">
<emphasis id="DDDEEA9A4D270474F596B4E866A9F9B4" bold="true" box="[234,470,1526,1553]" italics="true" pageId="34" pageNumber="35">Reitteria escherichi</emphasis>
Wasmann, 1896
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="EF1536884D270474F5C1B7046626F9F2" blockId="34.[189,1398,1526,1623]" pageId="34" pageNumber="35">
<materialsCitation id="5FC23CD54D270474F5C1B7046629F9F2" collectingDate="1975-05-02" collectorName="Besuchet &amp; Lobl" country="Turkey" location="Gocek" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Mugla">
<collectingCountry id="97BD76184D270474F5C1B704664CF991" box="[189,307,1562,1588]" name="Turkey" pageId="34" pageNumber="35">TURKEY</collectingCountry>
– 
<emphasis id="DDDEEA9A4D270474F42AB70465E1F996" bold="true" box="[342,670,1561,1588]" pageId="34" pageNumber="35">Western Anatolia, Aegean</emphasis>
• 1 ♂, 
<specimenCount id="F9ACFD014D270474F784B7046459F993" box="[760,806,1562,1590]" count="1" pageId="34" pageNumber="35" type="female">
1 
<emphasis id="DDDEEA9A4D270474F66CB7046459F993" bold="true" box="[784,806,1562,1590]" pageId="34" pageNumber="35">♀</emphasis>
</specimenCount>
; 
<collectingRegion id="2D6EF86A4D270474F64BB70764F8F996" box="[823,903,1561,1587]" country="Turkey" name="Mugla" pageId="34" pageNumber="35">Muğla</collectingRegion>
, 
<location id="EA7560534D270474F6E9B707649AF996" LSID="urn:lsid:plazi:treatment:6703879E4D240473F72FB78964DEFDC6:EA7560534D270474F6E9B707649AF996" box="[917,997,1561,1587]" country="Turkey" name="Gocek" pageId="34" pageNumber="35" stateProvince="Mugla">Göcek</location>
; 
<date id="9B1410484D270474F68FB70463F7F991" box="[1011,1160,1561,1588]" pageId="34" pageNumber="35" value="1975-05-02">
<collectingDate id="8B50E9A04D270474F68FB70463F7F991" box="[1011,1160,1561,1588]" pageId="34" pageNumber="35" value="1975-05-02">2 May 1975</collectingDate>
</date>
; 
<collectorName id="425F535E4D270474F1EAB707627DF996" box="[1174,1282,1561,1587]" pageId="34" pageNumber="35">Besuchet</collectorName>
and 
<collectorName id="425F535E4D270474F043B7076209F996" box="[1343,1398,1561,1587]" pageId="34" pageNumber="35">Löbl</collectorName>
leg.; 
<collectionCode id="89BBAE4D4D270474F584B7236629F9F2" box="[248,342,1597,1623]" pageId="34" pageNumber="35">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="EF1536884D270474F5C1B7616622F93D" blockId="34.[189,349,1662,1689]" box="[189,349,1662,1689]" pageId="34" pageNumber="35">
<heading id="B45D81E44D270474F5C1B7616622F93D" bold="true" box="[189,349,1662,1689]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DDDEEA9A4D270474F5C1B7616622F93D" bold="true" box="[189,349,1662,1689]" pageId="34" pageNumber="35">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D270474F5C1B7B366A0F962" blockId="34.[189,479,1709,1735]" box="[189,479,1709,1735]" pageId="34" pageNumber="35">
“ 
<collectingCountry id="97BD76184D270474F5BBB7B36612F962" box="[199,365,1709,1735]" name="Greece" pageId="34" pageNumber="35">Griechenland</collectingCountry>
”, 
<collectingCountry id="97BD76184D270474F4F8B7B366A4F962" box="[388,475,1709,1735]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D270472F5C1B7F06678FE3D" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="EF1536884D270474F5C1B7F06631F8AD" blockId="34.[189,334,1774,1800]" box="[189,334,1774,1800]" pageId="34" pageNumber="35">
<heading id="B45D81E44D270474F5C1B7F06631F8AD" bold="true" box="[189,334,1774,1800]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DDDEEA9A4D270474F5C1B7F06631F8AD" bold="true" box="[189,334,1774,1800]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D270474F5C1B603669EF84C" blockId="34.[187,1400,1821,2025]" pageId="34" pageNumber="35">
Male as in 
<figureCitation id="77912A0D4D270474F442B60366D7F89D" box="[318,424,1821,1848]" captionStart="Fig" captionStartId="35.[189,232,1963,1989]" captionTargetBox="[294,1312,465,1855]" captionTargetId="figure-99@35.[269,1318,454,1909]" captionTargetPageId="35" captionText="Fig. 10. Habitus, dorsal view. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♂. C. Ch. balcanica (Karaman, 1936), ♂. D. Ch. gallica (Schaufuss, 1876), ♂. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10424535" httpUri="https://zenodo.org/record/10424535/files/figure.png" pageId="34" pageNumber="35">Fig. 10B</figureCitation>
. Body length: 
<quantity id="28529B6D4D270474F72EB60365C5F89D" box="[594,698,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="34" pageNumber="35" unit="mm" value="1.36">1.36 mm</quantity>
average, range 
<quantity id="28529B6D4D270474F60FB6006362F89D" box="[883,1053,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="34" pageNumber="35" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.20–1.40 mm</quantity>
(N = 10, males and females). Shape of body oval, with the lateral margins of the pronotum slightly widened anteriorly. Elytral apex rounded. Terminal antennomere subtriangular, with round angles (
<figureCitation id="77912A0D4D270474F6BDB67A6351F8DB" box="[961,1070,1892,1918]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[341,1310,800,1791]" captionTargetId="figure-179@42.[221,1367,721,1840]" captionTargetPageId="42" captionText="Fig. 12. Terminal female antennomeres. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424539" httpUri="https://zenodo.org/record/10424539/files/figure.png" pageId="34" pageNumber="35">Figs 12B</figureCitation>
, 
<figureCitation id="77912A0D4D270474F146B67A630EF8DB" box="[1082,1137,1892,1918]" captionStart="Fig" captionStartId="44.[189,232,1928,1954]" captionTargetBox="[247,1330,760,1834]" captionTargetId="figure-280@44.[217,1371,719,1875]" captionTargetPageId="44" captionText="Fig. 13. Terminal male antennomeres.A. Cholovocera attae (Kraatz, 1858). B. Ch. balcánica(Karaman, 1936). C. Ch. gallica (Schaufuss, 1876). D. Ch. formicaria Victor, 1838. E. Ch. formiceticola (Rosenhauer, 1856). F. Ch. occulta sp. nov. G. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424541" httpUri="https://zenodo.org/record/10424541/files/figure.png" pageId="34" pageNumber="35">13A</figureCitation>
). Metatibiae relatively short, as in 
<figureCitation id="77912A0D4D270474F43DB69966B2F807" box="[321,461,1927,1954]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="34" pageNumber="35">Fig. 14B–C</figureCitation>
. Prosternal process slightly keeled anteriorly, with a marked median constriction and subquadrangular distally (
<figureCitation id="77912A0D4D270474F766B6B5650AF860" box="[538,629,1963,1989]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[264,1318,1324,1816]" captionTargetId="figure-432@10.[187,1400,1277,1877]" captionTargetPageId="10" captionText="Fig. 4. Prosternal processes. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845)." figureDoi="http://doi.org/10.5281/zenodo.10424520" httpUri="https://zenodo.org/record/10424520/files/figure.png" pageId="34" pageNumber="35">Fig. 4B</figureCitation>
). Male last visible ventrite with a slight emargination and bordered by a brush of short setae.
</paragraph>
<paragraph id="EF1536884D260475F5C1B0176655FE29" blockId="35.[187,1399,265,396]" pageId="35" pageNumber="36">
Median lobe of aedeagus curved towards the right in ventral view, tapering to a pointed apex (
<figureCitation id="77912A0D4D260475F181B0176215FE86" box="[1277,1386,265,291]" captionStart="Fig" captionStartId="55.[189,232,975,1001]" captionTargetBox="[475,1078,315,913]" captionTargetId="figure-56@55.[426,1132,265,950]" captionTargetPageId="55" captionText="Fig. 19. Aedeagus and paramere of Cholovocera attae (Kraatz, 1858) (Greece). A–B. Ventral views. C. Lateral view.. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424555" httpUri="https://zenodo.org/record/10424555/files/figure.png" pageId="35" pageNumber="36">Fig. 19A</figureCitation>
). Aedeagus in lateral view as in 
<figureCitation id="77912A0D4D260475F758B03265EFFEE3" box="[548,656,300,326]" captionStart="Fig" captionStartId="55.[189,232,975,1001]" captionTargetBox="[475,1078,315,913]" captionTargetId="figure-56@55.[426,1132,265,950]" captionTargetPageId="55" captionText="Fig. 19. Aedeagus and paramere of Cholovocera attae (Kraatz, 1858) (Greece). A–B. Ventral views. C. Lateral view.. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424555" httpUri="https://zenodo.org/record/10424555/files/figure.png" pageId="35" pageNumber="36">Fig. 19C</figureCitation>
. Distal portion of paramere conical, with five or six long apical setae (
<figureCitation id="77912A0D4D260475F47AB05166ECFECC" box="[262,403,335,361]" captionStart="Fig" captionStartId="55.[189,232,975,1001]" captionTargetBox="[475,1078,315,913]" captionTargetId="figure-56@55.[426,1132,265,950]" captionTargetPageId="55" captionText="Fig. 19. Aedeagus and paramere of Cholovocera attae (Kraatz, 1858) (Greece). A–B. Ventral views. C. Lateral view.. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424555" httpUri="https://zenodo.org/record/10424555/files/figure.png" pageId="35" pageNumber="36">Fig. 19B–C</figureCitation>
). The spermatheca was not dissected because of the small number of females available for study.
</paragraph>
<caption id="BBD566004D260475F5C1B6B56230F84C" ID-DOI="http://doi.org/10.5281/zenodo.10424535" ID-Zenodo-Dep="10424535" httpUri="https://zenodo.org/record/10424535/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,232,1963,1989]" targetBox="[294,1312,465,1855]" targetPageId="35">
<paragraph id="EF1536884D260475F5C1B6B56230F84C" blockId="35.[189,1399,1963,2025]" pageId="35" pageNumber="36">
<emphasis id="DDDEEA9A4D260475F5C1B6B56665F860" bold="true" box="[189,282,1963,1989]" pageId="35" pageNumber="36">Fig. 10.</emphasis>
Habitus, dorsal view. 
<emphasis id="DDDEEA9A4D260475F758B6B56546F860" bold="true" box="[548,569,1963,1989]" pageId="35" pageNumber="36">A</emphasis>
. 
<taxonomicName id="28AA4D0B4D260475F734B6B56447F860" box="[584,824,1963,1989]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="afghan">
<emphasis id="DDDEEA9A4D260475F734B6B56447F860" box="[584,824,1963,1989]" italics="true" pageId="35" pageNumber="36">Cholovocera afghan</emphasis>
</taxonomicName>
a 
<bibRefCitation id="8B3B4B794D260475F632B6B56485F860" author="Johnson C." box="[846,1018,1963,1989]" pageId="35" pageNumber="36" pagination="123 - 132" refId="ref39707" refString="Johnson C. 1977. Beitrage zur Kenntnis der fauna Afghanistans. Merophysiidae and Lathriidae, Coleoptera. Acta Musei Moraviae 62: 123 - 132." type="journal article" year="1977">Johnson, 1977</bibRefCitation>
, ♀. 
<emphasis id="DDDEEA9A4D260475F150B6B5633FF860" bold="true" box="[1068,1088,1963,1989]" pageId="35" pageNumber="36">B</emphasis>
. 
<taxonomicName id="28AA4D0B4D260475F133B6B5620CF863" authority="(Kraatz, 1858)" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1103,1395,1963,1990]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D260475F133B6B563C3F860" box="[1103,1212,1963,1989]" italics="true" pageId="35" pageNumber="36">Ch. attae</emphasis>
(
<bibRefCitation id="8B3B4B794D260475F1B0B6B56216F863" author="Kraatz G." box="[1228,1385,1963,1990]" pageId="35" pageNumber="36" pagination="123 - 148" refId="ref39933" refString="Kraatz G. 1858. Beitrag zur Kaferfauna Griechenlands. Staphylinidae (Schlufs), Trichopterygia, Histeridae, Phalacridae, Nitidularia, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrrhii. Berliner Entomologische Zeitschrift 2: 123 - 148." type="journal article" year="1858">Kraatz, 1858</bibRefCitation>
)
</taxonomicName>
, ♂. 
<emphasis id="DDDEEA9A4D260475F59EB6D06788F84D" bold="true" box="[226,247,1998,2024]" pageId="35" pageNumber="36">C</emphasis>
. 
<taxonomicName id="28AA4D0B4D260475F479B6D066D5F84D" box="[261,426,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D260475F479B6D066D5F84D" box="[261,426,1998,2024]" italics="true" pageId="35" pageNumber="36">Ch. balcanica</emphasis>
</taxonomicName>
(
<bibRefCitation id="8B3B4B794D260475F4C5B6D0650CF84D" author="Karaman Z." box="[441,627,1998,2024]" pageId="35" pageNumber="36" pagination="131 - 133" refId="ref39734" refString="Karaman Z. v. 1936. Ein balkanischer Vertreter des Koleopteren-Genus Reitteria, Reit. balcanica n. sp. Glasnik (Bulletin) de la Societe scientifique de Skopje 17: 131 - 133." type="journal article" year="1936">Karaman, 1936</bibRefCitation>
), ♂. 
<emphasis id="DDDEEA9A4D260475F7D1B6D065BDF84D" bold="true" box="[685,706,1998,2024]" pageId="35" pageNumber="36">D</emphasis>
. 
<taxonomicName id="28AA4D0B4D260475F7ADB6D0634DF84D" authority="(Schaufuss, 1876)" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[721,1074,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D260475F7ADB6D0642CF84D" box="[721,851,1998,2024]" italics="true" pageId="35" pageNumber="36">Ch. gallica</emphasis>
(Schaufuss, 1876)
</taxonomicName>
, ♂. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="EF1536884D210472F5C1B0166694FE87" blockId="36.[189,491,264,290]" box="[189,491,264,290]" pageId="36" pageNumber="37">
<heading id="B45D81E44D210472F5C1B0166694FE87" bold="true" box="[189,491,264,290]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="DDDEEA9A4D210472F5C1B0166694FE87" bold="true" box="[189,491,264,290]" pageId="36" pageNumber="37">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D210472F5C1B0296678FE3D" blockId="36.[189,1399,311,408]" pageId="36" pageNumber="37">
The known distribution of 
<taxonomicName id="28AA4D0B4D210472F77FB029659FFEF4" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[515,736,311,337]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F77FB029659FFEF4" box="[515,736,311,337]" italics="true" pageId="36" pageNumber="37">Cholovocera attae</emphasis>
</taxonomicName>
is in 
<collectingCountry id="97BD76184D210472F655B0296403FEF4" box="[809,892,311,337]" name="Greece" pageId="36" pageNumber="37">Greece</collectingCountry>
(
<collectingRegion id="2D6EF86A4D210472F6F2B0296358FEF4" box="[910,1063,311,337]" country="Greece" name="Peloponnisos" pageId="36" pageNumber="37">Peloponnese</collectingRegion>
, 
<collectingRegion id="2D6EF86A4D210472F14AB0296308FEF4" box="[1078,1143,311,337]" country="Greece" name="Kriti" pageId="36" pageNumber="37">Crete</collectingRegion>
) and western 
<collectingCountry id="97BD76184D210472F05FB0296208FEF4" box="[1315,1399,311,337]" name="Turkey" pageId="36" pageNumber="37">Turkey</collectingCountry>
(
<figureCitation id="77912A0D4D210472F5BAB044665BFED1" box="[198,292,346,372]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[303,1293,291,1782]" captionTargetId="figure-70@18.[247,1340,265,1810]" captionTargetPageId="18" captionText="Fig. 9. Known geographical distribution of the species ofCholovocera Victor, 1838.A. Ch. formiceticola (Rosenhauer, 1856) (red dots) and Ch. gallica (Schaufuss, 1876) (yellow dots). B. Ch. punctata (Märkel, 1845) (red dots) and Ch. occulta sp. nov. (yellow dots). C. Ch. balcanica (Karaman, 1936) (red dots), Ch. attae (Kraatz, 1858) (yellow dots), Ch. formicaria (dark blue dots) and Ch. afghana Johnson, 1977 (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10424533" httpUri="https://zenodo.org/record/10424533/files/figure.png" pageId="36" pageNumber="37">Fig. 9C</figureCitation>
). However, the few available records of this species may not represent its total distributional range.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D210472F5C1B0A164DFFC23" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="EF1536884D210472F5C1B0A1664CFE7C" blockId="36.[189,307,447,473]" box="[189,307,447,473]" pageId="36" pageNumber="37">
<heading id="B45D81E44D210472F5C1B0A1664CFE7C" bold="true" box="[189,307,447,473]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="DDDEEA9A4D210472F5C1B0A1664CFE7C" bold="true" box="[189,307,447,473]" pageId="36" pageNumber="37">
<collectingRegion id="2D6EF86A4D210472F5C1B0A16787FE7C" box="[189,248,447,473]" country="Afghanistan" name="Khost" pageId="36" pageNumber="37">Host</collectingRegion>
ants
</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D210472F5C1B0F366B6FD7E" blockId="36.[188,1399,493,731]" pageId="36" pageNumber="37">
<bibRefCitation id="8B3B4B794D210472F5C1B0F36625FDAD" author="Smith F." box="[189,346,493,520]" pageId="36" pageNumber="37" pagination="354" refId="ref42470" refString="Smith F. 1874. Letter, Entomological Society Feb. 16. Societies and Academies. Nature 9: 354." type="journal article" year="1874">Smith (1874)</bibRefCitation>
cited studies by 
<bibRefCitation id="8B3B4B794D210472F75AB0F3647DFDAD" author="Moggridge J. T." box="[550,770,493,520]" pageId="36" pageNumber="37" refId="ref40851" refString="Moggridge J. T. 1873. Harvesting Ants and Trap-door Spiders. Lovell Reeve &amp; Co., London." type="book" year="1873">Moggridge (1873)</bibRefCitation>
, who collected 
<taxonomicName id="28AA4D0B4D210472F6C3B0F36352FDA2" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[959,1069,493,519]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F6C3B0F36352FDA2" box="[959,1069,493,519]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
in nests of 
<taxonomicName id="28AA4D0B4D210472F1C6B0F06616FD8E" authority="Santschi, 1925" authorityName="Santschi" authorityYear="1925" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="DDDEEA9A4D210472F1C6B0F06208FDA2" box="[1210,1399,493,519]" italics="true" pageId="36" pageNumber="37">Messor ibericus</emphasis>
Santschi, 1925
</taxonomicName>
(as 
<taxonomicName id="28AA4D0B4D210472F4EBB30F6554FD8F" authorityName="Jerdon" authorityYear="1851" box="[407,555,529,555]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D210472F4EBB30F6554FD8F" box="[407,555,529,555]" italics="true" pageId="36" pageNumber="37">Atta structor</emphasis>
</taxonomicName>
) in Menton (southern 
<collectingCountry id="97BD76184D210472F64DB30F64F9FD8E" box="[817,902,529,555]" name="France" pageId="36" pageNumber="37">France</collectingCountry>
). 
<taxonomicName id="28AA4D0B4D210472F6EBB30F632BFD8E" authorityName="Santschi" authorityYear="1925" box="[919,1108,529,555]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="DDDEEA9A4D210472F6EBB30F632BFD8E" box="[919,1108,529,555]" italics="true" pageId="36" pageNumber="37">Messor ibericus</emphasis>
</taxonomicName>
is the only species of the 
<taxonomicName id="28AA4D0B4D210472F5C0B32B6608FDEB" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[188,375,565,590]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D210472F5C0B32B6608FDEB" box="[188,375,565,590]" italics="true" pageId="36" pageNumber="37">Messor structor</emphasis>
</taxonomicName>
complex present in that region (
<bibRefCitation id="8B3B4B794D210472F78DB32A64B0FDEB" author="Steiner F. M. &amp; Csosz S. &amp; Marko B. &amp; Gamisch A. &amp; Rinnhofer L. &amp; Folterbauer C. &amp; Hammerle S. &amp; Stauffer C. &amp; Arthofer W. &amp; Schlick-Steiner B. C." box="[753,975,564,590]" pageId="36" pageNumber="37" pagination="387 - 404" refId="ref42533" refString="Steiner F. M., Csosz S., Marko B., Gamisch A., Rinnhofer L., Folterbauer C., Hammerle S., Stauffer C., Arthofer W. &amp; Schlick-Steiner B. C. 2018. Turning one into five: Integrative taxonomy uncovers complex evolution of cryptic species in the harvester ant Messor &quot; structor &quot;. Molecular Phylogenetics and Evolution 127: 387 - 404. https: // doi. org / 10.1016 / j. ympev. 2018.04.005" type="journal article" year="2018">
Steiner 
<emphasis id="DDDEEA9A4D210472F631B32B64F7FDEB" box="[845,904,564,590]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
2018
</bibRefCitation>
). However, according to our present knowledge, the beetles could not have been 
<taxonomicName id="28AA4D0B4D210472F7B5B349644AFDD4" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[713,821,599,625]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F7B5B349644AFDD4" box="[713,821,599,625]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
, but 
<taxonomicName id="28AA4D0B4D210472F60EB34963E3FDD4" box="[882,1180,599,626]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="variety" species="punctata" variety="gallica">
<emphasis id="DDDEEA9A4D210472F60EB3496375FDD4" box="[882,1034,599,625]" italics="true" pageId="36" pageNumber="37">Ch. punctata</emphasis>
, 
<emphasis id="DDDEEA9A4D210472F165B34963E3FDD4" box="[1049,1180,599,625]" italics="true" pageId="36" pageNumber="37">Ch. gallica</emphasis>
</taxonomicName>
or 
<taxonomicName id="28AA4D0B4D210472F1BAB3496232FDD4" authority="Delgado &amp; Palma, 2023" authorityName="Delgado &amp; Palma" authorityYear="2023" box="[1222,1357,599,625]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="DDDEEA9A4D210472F1BAB3496232FDD4" box="[1222,1357,599,625]" italics="true" pageId="36" pageNumber="37">Ch. occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6ED57E14D210472F029B346678FFD30" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
Subsequent authors perpetuated the incorrect identifications of both beetles and 
<vernacularName id="61A946A64D210472F1C8B365639CFD30" authorityName="Latreille" authorityYear="1802" box="[1204,1251,635,661]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">ants</vernacularName>
(e.g., 
<bibRefCitation id="8B3B4B794D210472F04DB3656784FD1D" author="Lucas H." pageId="36" pageNumber="37" pagination="239" refId="ref40549" refString="Lucas H. 1874. Note. Seance 25 Novembre 1874. Annales de la Societe entomologique de France (Bulletin des Seances et Bulletin bibliographique) 5 (4): 239." type="journal article" year="1874">Lucas 1874</bibRefCitation>
; 
<bibRefCitation id="8B3B4B794D210472F471B38366D3FD1D" author="Rucker H. W." box="[269,428,669,696]" pageId="36" pageNumber="37" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker 1980</bibRefCitation>
; 
<bibRefCitation id="8B3B4B794D210472F4C0B383658BFD1D" author="Angelini F. &amp; Rucker H. W." box="[444,756,669,696]" pageId="36" pageNumber="37" pagination="213 - 240" refId="ref37938" refString="Angelini F. &amp; Rucker H. W. 1999. Contributo alla conoscenza dei Merophysiidae e Lathridiidae dell'Italia meridionale e della Sicilia (Coleoptera). Memorie della Societa Entomologica Italiana 77: 213 - 240." type="journal article" year="1999">Angelini &amp; Rücker 1999</bibRefCitation>
). However, 
<bibRefCitation id="8B3B4B794D210472F6F0B3836303FD1D" author="Rucker W. H." box="[908,1148,669,696]" pageId="36" pageNumber="37" refId="ref41751" refString="Rucker W. H. 2018. Latridiidae und Merophysiidae der West-Palaarktis. Selbstverlag Wolfgang H. Rucker, Neuwied." type="book" year="2018">Rücker (2018: 576)</bibRefCitation>
correctly associated 
<taxonomicName id="28AA4D0B4D210472F5C1B3DF6656FD7F" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[189,297,705,731]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F5C1B3DF6656FD7F" box="[189,297,705,731]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
with 
<taxonomicName id="28AA4D0B4D210472F417B3DF66BEFD7F" authorityName="Forel" authorityYear="1890" box="[363,449,705,730]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D210472F417B3DF66BEFD7F" box="[363,449,705,730]" italics="true" pageId="36" pageNumber="37">Messor</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D210472F5C1B21C64DFFC23" blockId="36.[188,1399,770,902]" pageId="36" pageNumber="37">
At present, it is not possible to know the species of 
<vernacularName id="61A946A64D210472F66CB21D644BFCB8" authorityName="Latreille" authorityYear="1802" box="[784,820,771,797]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">ant</vernacularName>
associated with 
<taxonomicName id="28AA4D0B4D210472F68FB21C63B3FCB9" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1011,1228,770,796]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F68FB21C63B3FCB9" box="[1011,1228,770,796]" italics="true" pageId="36" pageNumber="37">Cholovocera attae</emphasis>
</taxonomicName>
in 
<collectingCountry id="97BD76184D210472F18CB21C6238FCB9" box="[1264,1351,770,796]" name="Greece" pageId="36" pageNumber="37">Greece</collectingCountry>
. We have examined one beetle preserved together with an 
<vernacularName id="61A946A64D210472F645B2386422FCE5" authorityName="Latreille" authorityYear="1802" box="[825,861,806,832]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">ant</vernacularName>
worker of the genus 
<taxonomicName id="28AA4D0B4D210472F127B23863CEFC9A" authorityName="Forel" authorityYear="1890" box="[1115,1201,806,831]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D210472F127B23863CEFC9A" box="[1115,1201,806,831]" italics="true" pageId="36" pageNumber="37">Messor</emphasis>
</taxonomicName>
but, considering the large number of species of this genus living in 
<collectingCountry id="97BD76184D210472F66CB257641CFCC6" box="[784,867,841,867]" name="Greece" pageId="36" pageNumber="37">Greece</collectingCountry>
and the lack of good descriptions, we cannot identify it to species (X. Espadaler pers. comm. 
<date id="9B1410484D210472F78EB27264ECFC23" box="[754,915,876,902]" pageId="36" pageNumber="37" value="2021-03-13">13 Mar. 2021</date>
).
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D210473F5C1B2B364DEFDC6" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="discussion">
<paragraph id="EF1536884D210472F5C1B2B36532FC62" blockId="36.[189,589,941,967]" box="[189,589,941,967]" pageId="36" pageNumber="37">
<heading id="B45D81E44D210472F5C1B2B36532FC62" bold="true" box="[189,589,941,967]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="DDDEEA9A4D210472F5C1B2B36532FC62" bold="true" box="[189,589,941,967]" pageId="36" pageNumber="37">Taxonomic history and remarks</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D210472F5C1B2C2631EFAF3" blockId="36.[189,1399,987,1366]" pageId="36" pageNumber="37">
<bibRefCitation id="8B3B4B794D210472F5C1B2C266CDFC53" author="Kraatz G." box="[189,434,987,1014]" pageId="36" pageNumber="37" pagination="123 - 148" refId="ref39933" refString="Kraatz G. 1858. Beitrag zur Kaferfauna Griechenlands. Staphylinidae (Schlufs), Trichopterygia, Histeridae, Phalacridae, Nitidularia, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrrhii. Berliner Entomologische Zeitschrift 2: 123 - 148." type="journal article" year="1858">Kraatz’s (1858: 140)</bibRefCitation>
description of 
<taxonomicName id="28AA4D0B4D210472F71BB2C2643DFC50" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[615,834,988,1014]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F71BB2C2643DFC50" box="[615,834,988,1014]" italics="true" pageId="36" pageNumber="37">Cholovocera attae</emphasis>
</taxonomicName>
is brief, but he accurately distinguished it from both 
<taxonomicName id="28AA4D0B4D210472F586B2E166EDFBBC" authorityName="Markel" authorityYear="1845" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[250,402,1023,1049]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D210472F586B2E166EDFBBC" box="[250,402,1023,1049]" italics="true" pageId="36" pageNumber="37">Ch. punctata</emphasis>
</taxonomicName>
and 
<taxonomicName id="28AA4D0B4D210472F4B2B2E165E3FBBC" authorityName="Rosenhauer" authorityYear="1856" box="[462,668,1023,1049]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D210472F4B2B2E165E3FBBC" box="[462,668,1023,1049]" italics="true" pageId="36" pageNumber="37">Ch. formiceticola</emphasis>
</taxonomicName>
by its body shape and size. In their catalogue, 
<bibRefCitation id="8B3B4B794D210472F1B0B2E166D8FB98" author="Gemminger M. &amp; Harold B. de" pageId="36" pageNumber="37" refId="ref39469" refString="Gemminger M. &amp; Harold B. de 1868. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. III. Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydidae, Rhysodidae, Cucujidae, Cryptophagidae, Derodontidae, Latrididae, Othnidae, Mycetophagidae, Thorictidae, Dermestidae, Byrrhidae, Georyssidae, Parnidae, Heteroceridae, Lucanidae. Sumptu E. H. Gummi, Monachii, Paris, London." type="book" year="1868">Gemminger &amp; Harold (1868: 905)</bibRefCitation>
gave the distribution of 
<taxonomicName id="28AA4D0B4D210472F7A4B53C6438FB99" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[728,839,1058,1084]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F7A4B53C6438FB99" box="[728,839,1058,1084]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
as 
<collectingCountry id="97BD76184D210472F609B53C64B4FB99" box="[885,971,1058,1084]" name="Greece" pageId="36" pageNumber="37">Greece</collectingCountry>
, and 
<bibRefCitation id="8B3B4B794D210472F16CB53C6253FB99" author="Schaufuss L. W." box="[1040,1324,1058,1084]" pageId="36" pageNumber="37" pagination="393 - 400" refId="ref42052" refString="Schaufuss L. W. 1876 a. Ueber Merophysia Luc., Coluocera Mtsch. und Reitteria Led. Nunquam otiosus 2: 393 - 400." type="journal article" year="1876">Schaufuss (1876a: 396)</bibRefCitation>
made comments about its morphology. 
<bibRefCitation id="8B3B4B794D210472F725B55B643BFBFA" author="Reitter E." box="[601,836,1093,1120]" pageId="36" pageNumber="37" pagination="297 - 340" refId="ref41208" refString="Reitter E. 1875. Revision der europaischen Lathridiidae. Entomologische Zeitung 36: 297 - 340." type="journal article" year="1875">Reitter (1875: 301)</bibRefCitation>
examined the 
<typeStatus id="3011882A4D210472F682B558634EFBC5" box="[1022,1073,1094,1120]" pageId="36" pageNumber="37">type</typeStatus>
of 
<taxonomicName id="28AA4D0B4D210472F11EB55B63ACFBFA" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1122,1235,1093,1119]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F11EB55B63ACFBFA" box="[1122,1235,1093,1119]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
and found it similar to 
<taxonomicName id="28AA4D0B4D210472F444B5766579FB27" authorityName="Rosenhauer" authorityYear="1856" box="[312,518,1128,1154]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D210472F444B5766579FB27" box="[312,518,1128,1154]" italics="true" pageId="36" pageNumber="37">Ch. formiceticola</emphasis>
</taxonomicName>
, but he listed both 
<taxonomicName id="28AA4D0B4D210472F793B5766423FB27" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[751,860,1128,1154]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F793B5766423FB27" box="[751,860,1128,1154]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
and 
<taxonomicName id="28AA4D0B4D210472F6E4B5766319FB27" authorityName="Rosenhauer" authorityYear="1856" box="[920,1126,1128,1154]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D210472F6E4B5766319FB27" box="[920,1126,1128,1154]" italics="true" pageId="36" pageNumber="37">Ch. formiceticola</emphasis>
</taxonomicName>
as junior synonyms of 
<taxonomicName id="28AA4D0B4D210472F5C2B5926612FB03" authorityName="Motschulsky" authorityYear="1838" box="[190,365,1164,1190]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D210472F5C2B5926612FB03" box="[190,365,1164,1190]" italics="true" pageId="36" pageNumber="37">Ch. formicaria</emphasis>
</taxonomicName>
, an action accepted by 
<bibRefCitation id="8B3B4B794D210472F7FAB59264D3FB03" author="Heyden L. von &amp; Reitter E. &amp; Weise J." box="[646,940,1164,1190]" pageId="36" pageNumber="37" refId="ref39594" refString="Heyden L. von, Reitter E. &amp; Weise J. 1883. Catalogus coleopterorum Europae et Caucasi. 3 th ed. Libraria Nicolai, Berlin." type="book" year="1883">
Heyden 
<emphasis id="DDDEEA9A4D210472F797B5926457FB03" box="[747,808,1164,1190]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(1883: 80)
</bibRefCitation>
in their catalogue. 
<bibRefCitation id="8B3B4B794D210472F1EDB592620EFB03" author="Walker J. J." box="[1169,1393,1164,1190]" pageId="36" pageNumber="37" pagination="175 - 187" refId="ref42952" refString="Walker J. J. 1888. A year's insect-hunting at Gibraltar. Entomologist's Monthly Magazine 24: 175 - 187." type="journal article" year="1888">
<vernacularName id="61A946A64D210472F1EDB592639AFB03" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1169,1253,1164,1190]" canonicalName="Polyphylla fullo" class="Insecta" family="Scarabaeidae" genus="Polyphylla" kingdom="Animalia" language="deu" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="fullo">Walker</vernacularName>
(1888: 181
</bibRefCitation>
, 
<bibRefCitation id="8B3B4B794D210472F5C2B5B16646FB6C" author="Walker J. J." box="[190,313,1199,1225]" pageId="36" pageNumber="37" pagination="248 - 249" refId="ref43006" refString="Walker J. J. 1892. A year's insect-hunting at Gibraltar III. The Mediterranean Naturalist 2: 248 - 249." type="journal article" year="1892">1892: 248</bibRefCitation>
) reported “ 
<taxonomicName id="28AA4D0B4D210472F4BDB5B1654FFB6C" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[449,560,1199,1225]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F4BDB5B1654FFB6C" box="[449,560,1199,1225]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
” from 
<collectingCountry id="97BD76184D210472F7F5B5B1658AFB6C" box="[649,757,1199,1225]" name="Gibraltar" pageId="36" pageNumber="37">Gibraltar</collectingCountry>
, but it was most likely 
<taxonomicName id="28AA4D0B4D210472F168B5B1639DFB6C" authorityName="Rosenhauer" authorityYear="1856" box="[1044,1250,1199,1225]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D210472F168B5B1639DFB6C" box="[1044,1250,1199,1225]" italics="true" pageId="36" pageNumber="37">Ch. formiceticola</emphasis>
</taxonomicName>
, and 
<bibRefCitation id="8B3B4B794D210472F05BB5B16634FB49" author="Fauvel A." pageId="36" pageNumber="37" pagination="330 - 344" refId="ref39359" refString="Fauvel A. 1890. Rectifications au catalogus coleopterorum Europae et Caucasi. Revue d'Entomologie 9: 330 - 344." type="journal article" year="1890">Fauvel (1890: 338)</bibRefCitation>
resurrected 
<taxonomicName id="28AA4D0B4D210472F49DB5CC6530FB49" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[481,591,1234,1260]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F49DB5CC6530FB49" box="[481,591,1234,1260]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
as a valid taxon. A report by 
<bibRefCitation id="8B3B4B794D210472F6CAB5CD63C0FB49" author="Wasmann E." box="[950,1215,1234,1261]" pageId="36" pageNumber="37" pagination="297 - 304" refId="ref43029" refString="Wasmann E. 1890. Verzeichniss der von Dr. August Forel (Zurich) in Sud-Tunesien und Ost-Algerien gesammelten Ameisen-Gaste. Deutsche Entomologische Zeitschrift 2: 297 - 304." type="journal article" year="1890">Wasmann (1890: 298)</bibRefCitation>
in 
<collectingCountry id="97BD76184D210472F194B5CC6240FB49" box="[1256,1343,1234,1260]" name="Tunisia" pageId="36" pageNumber="37">Tunisia</collectingCountry>
was most likely of 
<taxonomicName id="28AA4D0B4D210472F416B5EB657DFAAA" authorityName="Markel" authorityYear="1845" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[362,514,1269,1295]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D210472F416B5EB657DFAAA" box="[362,514,1269,1295]" italics="true" pageId="36" pageNumber="37">Ch. punctata</emphasis>
</taxonomicName>
or 
<taxonomicName id="28AA4D0B4D210472F756B5EB65D3FAAA" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[554,684,1269,1295]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D210472F756B5EB65D3FAAA" box="[554,684,1269,1295]" italics="true" pageId="36" pageNumber="37">Ch. gallica</emphasis>
</taxonomicName>
. 
<bibRefCitation id="8B3B4B794D210472F7C0B5EB64B8FAAA" author="Bodemeyer E. von" box="[700,967,1269,1295]" pageId="36" pageNumber="37" refId="ref38519" refString="Bodemeyer E. von 1900. Quer durch Klein-Asien in den Bulgar-Dagh. Eine naturwissenschaftliche Studien-reise. Druck und Verlags-Aktien Gesellschaft vormals D ˆ lter, Emmendingen." type="book" year="1900">Bodemeyer (1900: 24)</bibRefCitation>
recorded “ 
<taxonomicName id="28AA4D0B4D210472F136B5EB63C8FAAA" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1098,1207,1269,1295]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F136B5EB63C8FAAA" box="[1098,1207,1269,1295]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
” in “Karakeuy” (Istanbul), but we have examined some of Bodemeyer’s material and have identified it as 
<taxonomicName id="28AA4D0B4D210472F1B0B4066210FA97" box="[1228,1391,1304,1330]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D210472F1B0B4066210FA97" box="[1228,1391,1304,1330]" italics="true" pageId="36" pageNumber="37">Ch. balcanica</emphasis>
</taxonomicName>
. 
<bibRefCitation id="8B3B4B794D210472F5C1B42266FFFAF3" author="Reitter E." box="[189,384,1339,1366]" pageId="36" pageNumber="37" pagination="1 - 9" refId="ref41305" refString="Reitter E. 1902. Neue und seltene Coleopteren, gesammelt im Jahre 1901, in der Herzegowina, in Dalmatien und Bosnien. Wiener Entomologische Zeitung 21: 1 - 9." type="journal article" year="1902">Reitter (1902: 5)</bibRefCitation>
compared the morphology of 
<taxonomicName id="28AA4D0B4D210472F798B422642FFAF0" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[740,848,1340,1366]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F798B422642FFAF0" box="[740,848,1340,1366]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
with that of 
<taxonomicName id="28AA4D0B4D210472F69AB4226326FAF3" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[998,1113,1340,1366]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D210472F69AB4226326FAF3" box="[998,1113,1340,1366]" italics="true" pageId="36" pageNumber="37">C. gallica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D210472F5C1B4636321F928" blockId="36.[187,1399,1405,1678]" pageId="36" pageNumber="37">
<bibRefCitation id="8B3B4B794D210472F5C1B46366E1FA32" author="Rucker H. W." box="[189,414,1405,1431]" pageId="36" pageNumber="37" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker (1980: 144)</bibRefCitation>
included 
<taxonomicName id="28AA4D0B4D210472F776B463650BFA32" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[522,628,1405,1431]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F776B463650BFA32" box="[522,628,1405,1431]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
in his key for the identification of 
<taxonomicName id="28AA4D0B4D210472F686B4636391FA32" box="[1018,1262,1405,1431]" pageId="36" pageNumber="37">
<emphasis id="DDDEEA9A4D210472F686B46363EDFA32" box="[1018,1170,1405,1431]" italics="true" pageId="36" pageNumber="37">Cholovocera</emphasis>
species
</taxonomicName>
, illustrating the median lobe of the aedeagus (
<bibRefCitation id="8B3B4B794D210472F741B4BE646EFA1E" author="Rucker H. W." box="[573,785,1440,1467]" pageId="36" pageNumber="37" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker 1980: 145</bibRefCitation>
, fig. 22); also, he gave the geographic distribution of this species as: southern Europe, 
<collectingCountry id="97BD76184D210472F746B4DA65F2FA7B" box="[570,653,1476,1502]" name="Greece" pageId="36" pageNumber="37">Greece</collectingCountry>
and northern Africa, probably taken from previous publications, and associated it with 
<taxonomicName id="28AA4D0B4D210472F4B2B4F965E4F9A4" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[462,667,1511,1537]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D210472F4B2B4F965E4F9A4" box="[462,667,1511,1537]" italics="true" pageId="36" pageNumber="37">Messor barbarus</emphasis>
</taxonomicName>
and 
<taxonomicName id="28AA4D0B4D210472F7A6B4F9641DF9A5" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[730,866,1511,1536]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D210472F7A6B4F9641DF9A5" box="[730,866,1511,1536]" italics="true" pageId="36" pageNumber="37">M. structor</emphasis>
</taxonomicName>
. Later, 
<bibRefCitation id="8B3B4B794D210472F6BEB4F963EEF9A4" author="Rucker W. H." box="[962,1169,1511,1537]" pageId="36" pageNumber="37" pagination="1 - 73" refId="ref41572" refString="Rucker W. H. 1983. Kul ˆ nb ˆ zi csapu bogarak VI. - Diversicornia VI. Bunkoscsapu bogarak VII. - Clavicornia VII. In: Zoltan K. (ed.) Magyarorszag Allatvilaga (Fauna Hungariae) 8 (3): 1 - 73. Akademiai Kiado, Budapest." type="journal article" year="1983">Rücker (1983: 4)</bibRefCitation>
reported 
<taxonomicName id="28AA4D0B4D210472F074B4F96208F9A5" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1288,1399,1511,1537]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F074B4F96208F9A5" box="[1288,1399,1511,1537]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
from southern 
<collectingCountry id="97BD76184D210472F40DB71466A6F981" box="[369,473,1546,1572]" name="Hungary" pageId="36" pageNumber="37">Hungary</collectingCountry>
, including a habitus figure with metatibiae equal to those of 
<taxonomicName id="28AA4D0B4D210472F1BBB7146210F981" box="[1223,1391,1546,1572]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D210472F1BBB7146210F981" box="[1223,1391,1546,1572]" italics="true" pageId="36" pageNumber="37">Ch. balcanica</emphasis>
</taxonomicName>
, and an illustration of the distal part of the aedeagus (
<bibRefCitation id="8B3B4B794D210472F65AB73364A0F9ED" author="Rucker W. H." box="[806,991,1581,1608]" pageId="36" pageNumber="37" pagination="1 - 73" refId="ref41572" refString="Rucker W. H. 1983. Kul ˆ nb ˆ zi csapu bogarak VI. - Diversicornia VI. Bunkoscsapu bogarak VII. - Clavicornia VII. In: Zoltan K. (ed.) Magyarorszag Allatvilaga (Fauna Hungariae) 8 (3): 1 - 73. Akademiai Kiado, Budapest." type="journal article" year="1983">Rücker 1983: 5</bibRefCitation>
, fig. f), which also matches that of 
<taxonomicName id="28AA4D0B4D210472F5C1B74E661CF9CF" box="[189,355,1616,1642]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D210472F5C1B74E661CF9CF" box="[189,355,1616,1642]" italics="true" pageId="36" pageNumber="37">Ch. balcanica</emphasis>
</taxonomicName>
. Actually, 
<bibRefCitation id="8B3B4B794D210472F49FB74E65DCF9CF" author="Rucker W. H." box="[483,675,1616,1642]" pageId="36" pageNumber="37" pagination="1 - 73" refId="ref41572" refString="Rucker W. H. 1983. Kul ˆ nb ˆ zi csapu bogarak VI. - Diversicornia VI. Bunkoscsapu bogarak VII. - Clavicornia VII. In: Zoltan K. (ed.) Magyarorszag Allatvilaga (Fauna Hungariae) 8 (3): 1 - 73. Akademiai Kiado, Budapest." type="journal article" year="1983">Rücker’s (1983)</bibRefCitation>
figure f, labelled as 
<taxonomicName id="28AA4D0B4D210472F6EAB74E637CF9CF" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[918,1027,1616,1642]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F6EAB74E637CF9CF" box="[918,1027,1616,1642]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
, represents the same species as his figure e, labelled as 
<taxonomicName id="28AA4D0B4D210472F4AEB76D6508F928" box="[466,631,1651,1677]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D210472F4AEB76D6508F928" box="[466,631,1651,1677]" italics="true" pageId="36" pageNumber="37">Ch. balcanica</emphasis>
</taxonomicName>
, but both shown in different orientations.
</paragraph>
<paragraph id="EF1536884D210472F5C1B7AB641BF8F9" blockId="36.[189,1399,1717,1884]" pageId="36" pageNumber="37">
D̂bler (1987: 14) recorded two “ 
<typeStatus id="3011882A4D210472F736B7AB65CAF96A" box="[586,693,1717,1743]" pageId="36" pageNumber="37" type="syntype">syntypes</typeStatus>
” of 
<taxonomicName id="28AA4D0B4D210472F797B7AB6427F96A" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[747,856,1717,1743]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F797B7AB6427F96A" box="[747,856,1717,1743]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
in the SDEI collection, although one of them had a label reading “ 
<typeStatus id="3011882A4D210472F4CAB7C6654AF957" box="[438,565,1752,1778]" pageId="36" pageNumber="37" type="holotype">Holotypus</typeStatus>
” (see 
<bibRefCitation id="8B3B4B794D210472F7FDB7C66455F957" author="Rucker W. H." box="[641,810,1752,1778]" pageId="36" pageNumber="37" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker 2011a</bibRefCitation>
: fig. 12). 
<bibRefCitation id="8B3B4B794D210472F6E7B7C663CAF957" author="Audisio P. &amp; Canepari C. &amp; De Biase A. &amp; Poggi R. &amp; Ratti E. &amp; Zampetti M. F." box="[923,1205,1752,1779]" pageId="36" pageNumber="37" pagination="1 - 19" refId="ref38021" refString="Audisio P., Canepari C., De Biase A., Poggi R., Ratti E. &amp; Zampetti M. F. 1995. Coleoptera Polyphaga XI (Clavicornia II). In: Minelli A., Ruffo S. &amp; La Posta S. (eds) Checklist delle specie della fauna italiana 56: 1 - 19. Calderini, Bologna." type="journal article" year="1995">
Audisio 
<emphasis id="DDDEEA9A4D210472F17DB7C76341F957" box="[1025,1086,1752,1778]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(1995: 9)
</bibRefCitation>
and 
<bibRefCitation id="8B3B4B794D210472F18CB7C666DCF8B0" author="Angelini F. &amp; Rucker H. W." pageId="36" pageNumber="37" pagination="213 - 240" refId="ref37938" refString="Angelini F. &amp; Rucker H. W. 1999. Contributo alla conoscenza dei Merophysiidae e Lathridiidae dell'Italia meridionale e della Sicilia (Coleoptera). Memorie della Societa Entomologica Italiana 77: 213 - 240." type="journal article" year="1999">Angelini &amp; Rücker (1999: 218)</bibRefCitation>
cited 
<collectingCountry id="97BD76184D210472F497B7E5655EF8B0" box="[491,545,1787,1813]" name="Italy" pageId="36" pageNumber="37">Italy</collectingCountry>
as the location of 
<taxonomicName id="28AA4D0B4D210472F788B7E56420F8B0" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[756,863,1787,1813]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F788B7E56420F8B0" box="[756,863,1787,1813]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
, again an error by confusing this species with 
<taxonomicName id="28AA4D0B4D210472F5C1B600662AF89D" authorityName="Markel" authorityYear="1845" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[189,341,1822,1848]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D210472F5C1B600662AF89D" box="[189,341,1822,1848]" italics="true" pageId="36" pageNumber="37">Ch. punctata</emphasis>
</taxonomicName>
. We have examined over 
<specimenCount id="F9ACFD014D210472F7F5B6016456F89D" box="[649,809,1822,1849]" count="30" pageId="36" pageNumber="37" type="generic">30 specimens</specimenCount>
collected by F. Angelini in 
<collectingRegion id="2D6EF86A4D210472F10FB60063C6F89D" box="[1139,1209,1822,1848]" country="Italy" name="Sicilia" pageId="36" pageNumber="37">Sicily</collectingRegion>
, which we have identified as 
<taxonomicName id="28AA4D0B4D210472F42FB65C6694F8FE" authorityName="Markel" authorityYear="1845" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[339,491,1858,1884]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D210472F42FB65C6694F8FE" box="[339,491,1858,1884]" italics="true" pageId="36" pageNumber="37">Ch. punctata</emphasis>
</taxonomicName>
(see Material examined above).
</paragraph>
<paragraph id="EF1536884D210473F5C1B69D6258FEED" blockId="36.[188,1401,1923,2020]" lastBlockId="37.[189,1398,265,328]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<bibRefCitation id="8B3B4B794D210472F5C1B69D66C0F838" author="Sar J. &amp; Dudas G. &amp; Merkl O." box="[189,447,1923,1949]" pageId="36" pageNumber="37" pagination="329 - 332" refId="ref41952" refString="Sar J., Dudas G. &amp; Merkl O. 2004. A hangyasz alb ˆ de (Pleganophorus bispinosus Hampe, 1855) Magyarorszagon (Coleoptera: Endomychidae). Somogyi Muzeumok Kozlemenyei 16: 329 - 332." type="journal article" year="2004">
Sár 
<emphasis id="DDDEEA9A4D210472F591B69A6655F838" box="[237,298,1923,1949]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(2004: 331)
</bibRefCitation>
expressed doubt about the record of 
<taxonomicName id="28AA4D0B4D210472F603B69D6495F838" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[895,1002,1923,1949]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D210472F603B69D6495F838" box="[895,1002,1923,1949]" italics="true" pageId="36" pageNumber="37">Ch. attae</emphasis>
</taxonomicName>
in 
<bibRefCitation id="8B3B4B794D210472F16FB69D63BFF838" author="Rucker W. H." box="[1043,1216,1923,1949]" pageId="36" pageNumber="37" pagination="1 - 73" refId="ref41572" refString="Rucker W. H. 1983. Kul ˆ nb ˆ zi csapu bogarak VI. - Diversicornia VI. Bunkoscsapu bogarak VII. - Clavicornia VII. In: Zoltan K. (ed.) Magyarorszag Allatvilaga (Fauna Hungariae) 8 (3): 1 - 73. Akademiai Kiado, Budapest." type="journal article" year="1983">Rücker (1983)</bibRefCitation>
from 
<collectingCountry id="97BD76184D210472F076B69A620DF83B" box="[1290,1394,1924,1950]" name="Hungary" pageId="36" pageNumber="37">Hungary</collectingCountry>
, commenting that some records and specimens found in museum collections labelled as from “ 
<collectingCountry id="97BD76184D210472F187B6B86216F865" box="[1275,1385,1958,1984]" name="Hungary" pageId="36" pageNumber="37">Hungary</collectingCountry>
”, may have originated from territories which once were part of the Austro-Hungarian Empire, such as 
<collectingCountry id="97BD76184D200473F5C1B0176674FE86" box="[189,267,265,291]" name="Serbia" pageId="37" pageNumber="38">Serbia</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F464B01766FDFE86" box="[280,386,265,291]" name="Romania" pageId="37" pageNumber="38">Romania</collectingCountry>
and nearby countries. Therefore, we believe that, unless specimens are re-examined, reports from those countries should be considered doubtful, probably referring to 
<taxonomicName id="28AA4D0B4D200473F107B033625FFEE2" box="[1147,1312,301,327]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D200473F107B033625FFEE2" box="[1147,1312,301,327]" italics="true" pageId="37" pageNumber="38">Ch. balcanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D200473F5C1B06E64DEFDC6" blockId="37.[187,1399,368,612]" pageId="37" pageNumber="38">
Several subsequent catalogues and checklists gave large geographic distributions for 
<taxonomicName id="28AA4D0B4D200473F1F3B06E6386FE2F" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1167,1273,368,394]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D200473F1F3B06E6386FE2F" box="[1167,1273,368,394]" italics="true" pageId="37" pageNumber="38">Ch. attae</emphasis>
</taxonomicName>
, including 
<collectingCountry id="97BD76184D200473F5C7B08B6671FE0A" box="[187,270,405,431]" name="France" pageId="37" pageNumber="38">France</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F460B08A660DFE0B" box="[284,370,404,430]" name="Greece" pageId="37" pageNumber="38">Greece</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F4FCB08A66C8FE0B" box="[384,439,404,430]" name="Italy" pageId="37" pageNumber="38">Italy</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F4B9B08B6552FE0A" box="[453,557,405,431]" name="Hungary" pageId="37" pageNumber="38">Hungary</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F746B08A65F7FE0B" box="[570,648,404,430]" name="Serbia" pageId="37" pageNumber="38">Serbia</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F7EAB08B6457FE0A" box="[662,808,405,431]" name="Serbia" pageId="37" pageNumber="38">Montenegro</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F649B08A64EFFE0B" box="[821,912,404,430]" name="Algeria" pageId="37" pageNumber="38">Algeria</collectingCountry>
, 
<collectingCountry id="97BD76184D200473F6E2B08B6377FE0A" box="[926,1032,405,431]" name="Morocco" pageId="37" pageNumber="38">Morocco</collectingCountry>
and 
<collectingCountry id="97BD76184D200473F139B08A63E0FE0B" box="[1093,1183,404,430]" name="Tunisia" pageId="37" pageNumber="38">Tunisia</collectingCountry>
, which we regard as erroneous, except for 
<collectingCountry id="97BD76184D200473F49EB0A6654AFE77" box="[482,565,440,466]" name="Greece" pageId="37" pageNumber="38">Greece</collectingCountry>
(L̂bl &amp; 
<bibRefCitation id="8B3B4B794D200473F7D8B0A664F4FE76" author="Smetana A." box="[676,907,440,467]" pageId="37" pageNumber="38" refId="ref40417" refString="L ˆ bl I. &amp; Smetana A. 2007. Catalogue of Palaearctic Coleoptera, vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup. https: // doi. org / 10.1163 / 9789004260894" type="book" year="2007">Smetana 2007: 557</bibRefCitation>
; 
<bibRefCitation id="8B3B4B794D200473F6E4B0A6631FFE77" author="Rucker W. H." box="[920,1120,440,466]" pageId="37" pageNumber="38" pagination="1 - 15" refId="ref41629" refString="Rucker W. H. 2009. Checklist Latridiidae &amp; Merophysiinae of the World. Latridiidae und Merophysiinae 9: 1 - 15." type="journal article" year="2009">Rücker 2009: 14</bibRefCitation>
, 
<bibRefCitation id="8B3B4B794D200473F111B0A663C6FE77" author="Rucker W. H." box="[1133,1209,440,466]" pageId="37" pageNumber="38" refId="ref41721" refString="Rucker W. H. 2011 b. Fauna Europaea: Coleoptera, Endomychidae: Merophysiinae. Fauna Europaea ver. 2017.06 http: // www. faunaeur. org" type="url" year="2011">2011b</bibRefCitation>
; 
<bibRefCitation id="8B3B4B794D200473F1BAB0A6664AFE52" author="Shockley F. W. &amp; Tomaszewska K. W. &amp; McHugh J. V." pageId="37" pageNumber="38" pagination="1 - 113" refId="ref42306" refString="Shockley F. W., Tomaszewska K. W. &amp; McHugh J. V. 2009 b. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113. https: // doi. org / 10.11646 / zootaxa. 1999.1.1" type="journal article" year="2009">
Shockley 
<emphasis id="DDDEEA9A4D200473F040B0A76208FE77" box="[1340,1399,440,466]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2009b: 65
</bibRefCitation>
). However, 
<bibRefCitation id="8B3B4B794D200473F4BCB0C265E4FE52" author="Rucker W. H." box="[448,667,476,503]" pageId="37" pageNumber="38" refId="ref41751" refString="Rucker W. H. 2018. Latridiidae und Merophysiidae der West-Palaarktis. Selbstverlag Wolfgang H. Rucker, Neuwied." type="book" year="2018">Rücker (2018: 576</bibRefCitation>
, 
<bibRefCitation id="8B3B4B794D200473F7D9B0C36472FE52" author="Rucker W. H." box="[677,781,477,503]" pageId="37" pageNumber="38" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">2020: 34</bibRefCitation>
) restricted that distribution to only 
<collectingRegion id="2D6EF86A4D200473F1DEB0C3639DFE52" box="[1186,1250,477,503]" country="Greece" name="Kriti" pageId="37" pageNumber="38">Crete</collectingRegion>
and Rhodes, which is partially correct. Also, 
<bibRefCitation id="8B3B4B794D200473F74AB31F6468FDB9" author="Rucker W. H." box="[566,791,513,540]" pageId="37" pageNumber="38" refId="ref41751" refString="Rucker W. H. 2018. Latridiidae und Merophysiidae der West-Palaarktis. Selbstverlag Wolfgang H. Rucker, Neuwied." type="book" year="2018">Rücker (2018: 576</bibRefCitation>
, fig. 1187) gave a detailed description of 
<taxonomicName id="28AA4D0B4D200473F07FB31F6210FDBE" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[1283,1391,513,539]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D200473F07FB31F6210FDBE" box="[1283,1391,513,539]" italics="true" pageId="37" pageNumber="38">Ch. attae</emphasis>
</taxonomicName>
, including a figure of the aedeagus in lateral view. Finally, 
<bibRefCitation id="8B3B4B794D200473F61FB33B6340FD9A" author="Stalling T." box="[867,1087,549,575]" pageId="37" pageNumber="38" pagination="11 - 13" refId="ref42492" refString="Stalling T. 2019. Aktuelle funde einiger Merophysiinae und eine neue Wirtsameise von Reitteria escherichi Wasmann, 1896 von der insel Kos, Griechenland (Coleoptera: Merophysiidae). Folia Latridiidae et Merophysiidae 2: 11 - 13." type="journal article" year="2019">Stalling (2019: 13)</bibRefCitation>
reported the island of Kos (
<collectingCountry id="97BD76184D200473F5B9B357662AFDC6" box="[197,341,585,611]" name="Greece" pageId="37" pageNumber="38">Dodecanese</collectingCountry>
Islands) as a new locality for 
<taxonomicName id="28AA4D0B4D200473F7C9B357645EFDC6" baseAuthorityName="Kraatz" baseAuthorityYear="1858" box="[693,801,585,611]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D200473F7C9B357645EFDC6" box="[693,801,585,611]" italics="true" pageId="37" pageNumber="38">Ch. attae</emphasis>
</taxonomicName>
in 
<collectingCountry id="97BD76184D200473F63AB35764E2FDC6" box="[838,925,585,611]" name="Greece" pageId="37" pageNumber="38">Greece</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>