594 lines
72 KiB
XML
594 lines
72 KiB
XML
<document id="E9A35623A95C6C5F460A3C4A775DAB62" ID-CLB-Dataset="8271" ID-DOI="10.11646/zootaxa.4859.4.4" ID-GBIF-Dataset="1bbcdb6e-555e-4e19-9160-46753ee0174a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4413354" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609698821548" checkinUser="plazi" docAuthor="Calcetas, Orlando A., Adorada, Jessamyn R. & Navasero, Mario V." docDate="2020" docId="656A3E54FFEAD112FF61FC149A851D37" docLanguage="en" docName="zootaxa.4859.4.4.pdf" docOrigin="Zootaxa 4859 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exopholis philippinica Brenske. Highest 1894" docType="treatment" docVersion="7" lastPageNumber="535" masterDocId="9953462CFFE8D11AFFF6FFC69F1F1E0E" masterDocTitle="Review of the genus Exopholis Motschulsky, 1859 (Coleoptera: Scarabaeidae Melolonthinae: Leucopholini) in the Philippines" masterLastPageNumber="536" masterPageNumber="527" pageNumber="529" updateTime="1698851825736" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="301A4EE5F34C14804788CE3EB275467E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="395833D46774B73B7E9F0395C79D0C6F">
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<mods:title id="FAC40FC4829366C82B5F650DB0E75B50">Review of the genus Exopholis Motschulsky, 1859 (Coleoptera: Scarabaeidae Melolonthinae: Leucopholini) in the Philippines</mods:title>
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<mods:name id="A1B4C31DD9FB17FA7ED6AB89487B02AD" type="personal">
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<mods:roleTerm id="1FD13754FE6680A732FD4ADA3A7E451C">Author</mods:roleTerm>
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<mods:namePart id="1147AC927988AAC3BB533DB68264C123">Calcetas, Orlando A.</mods:namePart>
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<mods:affiliation id="46FB323E289511C99EFEF533E4C7CC6D">Regional Crop Protection Center - Region IV-CALABARZON, Timugan, Los Baños, Laguna 4030, Philippines.</mods:affiliation>
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<mods:name id="F5A23DD9D1F0DDEC8F1A4E31C77921A7" type="personal">
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<mods:roleTerm id="42E23F0CC3D4B15BF11B92F069A2E9CA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="97013963E5D1A62EF21928EE52575322">Adorada, Jessamyn R.</mods:namePart>
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<mods:affiliation id="3338483D89F4C9DAB5FFE7270E7A9745">Crop Protection Cluster-College of Agriculture, University of the Philippines Los Baños, 4031 College, Laguna, Philippines. jradorada @ up. edu. ph; https: // orcid. org / 0000 - 0001 - 7204 - 0966</mods:affiliation>
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<mods:nameIdentifier id="5E708167A211C0AAC6ABD9484E9EBB0A" type="email">jradorada@up.edu.ph</mods:nameIdentifier>
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<mods:roleTerm id="5BB0FBD429B56A30B9F83699677B0B06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C5844DDB5D8052F2DE123010F5666575">Navasero, Mario V.</mods:namePart>
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<mods:affiliation id="F8026FE576DE3F468EC5ABB77F021A08">Crop Protection Cluster-College of Agriculture, University of the Philippines Los Baños, 4031 College, Laguna, Philippines. marnavasero @ gmail. com</mods:affiliation>
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<mods:typeOfResource id="5672C91EDA73FA6C2B2226DC9B7B17DF">text</mods:typeOfResource>
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<mods:title id="13F9FFE7CB5469EAFEE3266D3D6C678B">Zootaxa</mods:title>
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<mods:part id="C9C824E89CDD3D2E339701F0760D4AB4">
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<mods:date id="99766E6E818D6A37392F935DFD170A7E">2020</mods:date>
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<mods:number id="00FC0A60A7C404EF679F41CBAE99391C">2020-10-08</mods:number>
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<mods:number id="35CE35768A42E6CCAD0E73A2CC209058">4859</mods:number>
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<treatment id="656A3E54FFEAD112FF61FC149A851D37" ID-DOI="http://doi.org/10.5281/zenodo.4413364" ID-GBIF-Taxon="177319791" ID-Zenodo-Dep="4413364" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37" httpUri="http://treatment.plazi.org/id/656A3E54FFEAD112FF61FC149A851D37" lastPageId="8" lastPageNumber="535" pageId="2" pageNumber="529">
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<subSubSection id="A5D9DCC9FFEAD118FF61FC149D451DE2" box="[151,602,978,1005]" pageId="2" pageNumber="529" type="nomenclature">
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<paragraph id="ED7C8F42FFEAD118FF61FC149D451DE2" blockId="2.[151,602,978,1040]" box="[151,602,978,1005]" pageId="2" pageNumber="529">
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<heading id="B634382EFFEAD118FF61FC149D451DE2" bold="true" box="[151,602,978,1005]" fontSize="11" level="1" pageId="2" pageNumber="529" reason="1">
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<taxonomicName id="2AC3F4C1FFEAD118FF61FC149D451DE2" authority="Brenske 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[151,602,978,1005]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="philippinica">
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<emphasis id="DFB75350FFEAD118FF61FC149D451DE2" bold="true" box="[151,602,978,1005]" pageId="2" pageNumber="529">
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<emphasis id="DFB75350FFEAD118FF61FC149EB51DE3" bold="true" box="[151,426,978,1005]" italics="true" pageId="2" pageNumber="529">Exopholis philippinica</emphasis>
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<bibRefCitation id="8952F2B3FFEAD118FE47FC149D451DE2" author="Brenske, E." box="[433,602,978,1004]" pageId="2" pageNumber="529" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske 1894</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5D9DCC9FFEAD118FF61FC309E3F1A1E" box="[151,288,1014,1040]" pageId="2" pageNumber="529" type="description">
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<paragraph id="ED7C8F42FFEAD118FF61FC309E3F1A1E" blockId="2.[151,602,978,1040]" box="[151,288,1014,1040]" pageId="2" pageNumber="529">
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(
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<figureCitation id="75F893C7FFEAD118FF69FC309E071A1E" box="[159,280,1014,1040]" captionStart-0="FIGURES 4–6" captionStart-1="FIGURES 7–9" captionStart-2="FIGURES 10–12" captionStart-3="FIGURE 13" captionStartId-0="4.[151,264,1849,1873]" captionStartId-1="5.[151,264,1867,1891]" captionStartId-2="6.[151,264,1981,2005]" captionStartId-3="7.[151,250,1721,1745]" captionTargetBox-0="[193,1329,216,1808]" captionTargetBox-1="[227,1378,227,1833]" captionTargetBox-2="[320,1230,185,1933]" captionTargetBox-3="[245,1279,181,1697]" captionTargetId-0="figure-20@4.[151,1436,181,1825]" captionTargetId-1="figure-18@5.[151,1436,181,1843]" captionTargetId-2="figure-20@6.[258,1330,181,1957]" captionTargetId-3="figure-18@7.[151,1436,178,1697]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 4–6. Exopholis philippinica Brenske, 1894, female, holotype from SMT. 4, habitus, dorsal aspect; 5, habitus, ventral aspect; 6, habitus, lateral aspect (photograph of the female type specimen from SMT, courtesy of Dr. Olaf Jaeger)." captionText-1="FIGURES 7–9. Lepidiota corpulenta Moser, 1910, female, syntype from ZMHB. 7, habitus, dorsal aspect; 8, habitus, ventral aspect; 9, habitus, lateral aspect (photograph of the female from ZMHB, courtesy of Dr. Johannes Frisch and Dr. Joachim Willers)." captionText-2="FIGURES 10–12. Exopholis philippinica Brenske, 1894, male from IRRI. 10, habitus, dorsal aspect; 11, male genitalia, dorsal aspect; 12, male genitalia, lateral aspect." captionText-3="FIGURE 13. Distribution of examined specimens of Exopholis philippinica Brenske, 1894 in the Philippines." figureDoi-0="http://doi.org/10.5281/zenodo.4413360" figureDoi-1="http://doi.org/10.5281/zenodo.4413362" figureDoi-2="http://doi.org/10.5281/zenodo.4413366" figureDoi-3="http://doi.org/10.5281/zenodo.4413368" httpUri-0="https://zenodo.org/record/4413360/files/figure.png" httpUri-1="https://zenodo.org/record/4413362/files/figure.png" httpUri-2="https://zenodo.org/record/4413366/files/figure.png" httpUri-3="https://zenodo.org/record/4413368/files/figure.png" pageId="2" pageNumber="529">Figs. 4–13</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="A5D9DCC9FFEAD118FF61FBFD9C6C1AA1" pageId="2" pageNumber="529" type="reference_group">
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<paragraph id="ED7C8F42FFEAD118FF61FBFD9D7F1A7F" blockId="2.[151,1437,1082,1199]" pageId="2" pageNumber="529">
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<treatmentCitationGroup id="CDD3A86CFFEAD118FF61FBFD9D7F1A7F" pageId="2" pageNumber="529">
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<taxonomicName id="2AC3F4C1FFEAD118FF61FBFD9D041A5D" authority="Brenske, 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[151,539,1083,1107]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="philippinica">
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||
<emphasis id="DFB75350FFEAD118FF61FBFD9E611A5D" box="[151,382,1083,1107]" italics="true" pageId="2" pageNumber="529">Exopholis philippinica</emphasis>
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<bibRefCitation id="8952F2B3FFEAD118FE73FBFD9D041A5D" author="Brenske, E." box="[389,539,1083,1107]" pageId="2" pageNumber="529" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske, 1894</bibRefCitation>
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</taxonomicName>
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||
, 38: 354-356 (original description,
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<typeStatus id="327831E0FFEAD118FC73FBFD9CAE1A5D" box="[901,945,1083,1107]" pageId="2" pageNumber="529">type</typeStatus>
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locality: “
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<collectingCountry id="95D4CFD2FFEAD118FBEDFBFD9B871A5D" box="[1051,1176,1083,1107]" name="Philippines" pageId="2" pageNumber="529">Philippinen</collectingCountry>
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”);
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||
<treatmentCitation id="6C62A953FFEAD118FB42FBFD9A831A5D" author="Dalla Torre, K. W." box="[1204,1436,1083,1107]" page="169" pageId="2" pageNumber="529" year="1912">
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<bibRefCitation id="8952F2B3FFEAD118FB42FBFD9A831A5D" author="Dalla Torre, K. W." box="[1204,1436,1083,1107]" pageId="2" pageNumber="529" pagination="135 - 290" refId="ref4138" refString="Dalla Torre, K. W. (1912) Pars 49. Scarabaeidae: Melolonthinae III. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290." type="book chapter" year="1912">Dalla Torre, 1912: 169</bibRefCitation>
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||
</treatmentCitation>
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||
(catalog);
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||
<treatmentCitation id="6C62A953FFEAD118FED9FB9F9EE61A7F" author="Schultze, W. A." box="[303,505,1113,1137]" page="178" pageId="2" pageNumber="529" year="1916">
|
||
<bibRefCitation id="8952F2B3FFEAD118FED9FB9F9EE61A7F" author="Schultze, W. A." box="[303,505,1113,1137]" pageId="2" pageNumber="529" pagination="1 - 178" refId="ref4321" refString="Schultze, W. A. (1916) Catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 1 - 178." type="journal article" year="1916">Schultze, 1916: 178</bibRefCitation>
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</treatmentCitation>
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||
(catalog).
|
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="ED7C8F42FFEAD118FF61FBBE9C6C1AA1" blockId="2.[151,1437,1082,1199]" pageId="2" pageNumber="529">
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<treatmentCitationGroup id="CDD3A86CFFEAD118FF61FBBE9C6C1AA1" pageId="2" pageNumber="529">
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<treatmentCitation id="6C62A953FFEAD118FF61FBBE9D2E1A9E" author="Moser, J." box="[151,561,1144,1168]" page="185" pageId="2" pageNumber="529" year="1910">
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||
<taxonomicName id="2AC3F4C1FFEAD118FF61FBBE9D2E1A9E" authority="Moser, 1910: 185" authorityName="Moser. To Dr. Olaf Jaeger" authorityPageNumber="185" authorityYear="1910" box="[151,561,1144,1168]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="corpulenta">
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||
<emphasis id="DFB75350FFEAD118FF61FBBE9E6F1A9E" box="[151,368,1144,1168]" italics="true" pageId="2" pageNumber="529">Lepidiota corpulenta</emphasis>
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<bibRefCitation id="8952F2B3FFEAD118FE8FFBBE9D2E1A9E" author="Moser, J." box="[377,561,1144,1168]" pageId="2" pageNumber="529" pagination="149 - 150" refId="ref4269" refString="Moser, J. (1910) Neue Coleoptera Lamellicornia von den Philippen. The Philippine Journal of Science, 5, 149 - 150." type="journal article" year="1910">Moser, 1910: 185</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(primary description,
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<typeStatus id="327831E0FFEAD118FCEBFBBE9C561A9E" box="[797,841,1144,1168]" pageId="2" pageNumber="529">type</typeStatus>
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locality: “Mindanao, Camp Keithley, P.I.”);
|
||
<treatmentCitation id="6C62A953FFEAD118FAD6FBBE9E321AA1" author="Dalla Torre, K. W." page="170" pageId="2" pageNumber="529" year="1912">
|
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<bibRefCitation id="8952F2B3FFEAD118FAD6FBBE9E321AA1" author="Dalla Torre, K. W." pageId="2" pageNumber="529" pagination="135 - 290" refId="ref4138" refString="Dalla Torre, K. W. (1912) Pars 49. Scarabaeidae: Melolonthinae III. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290." type="book chapter" year="1912">Dalla Torre, 1912: 170</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalog);
|
||
<treatmentCitation id="6C62A953FFEAD118FE6DFB519D7A1AA1" author="Schultze, W. A." box="[411,613,1175,1199]" page="178" pageId="2" pageNumber="529" year="1916">
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<bibRefCitation id="8952F2B3FFEAD118FE6DFB519D7A1AA1" author="Schultze, W. A." box="[411,613,1175,1199]" pageId="2" pageNumber="529" pagination="1 - 178" refId="ref4321" refString="Schultze, W. A. (1916) Catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 1 - 178." type="journal article" year="1916">Schultze, 1916: 178</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalog);
|
||
<emphasis id="DFB75350FFEAD118FD25FB519C6C1AA1" bold="true" box="[723,883,1175,1199]" pageId="2" pageNumber="529">new synonymy</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
|
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<subSubSection id="A5D9DCC9FFEAD118FF61FB1B9B8E1851" pageId="2" pageNumber="529" type="materials_examined">
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<paragraph id="ED7C8F42FFEAD118FF61FB1B9A471B6D" blockId="2.[151,1437,1245,2027]" pageId="2" pageNumber="529">
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<emphasis id="DFB75350FFEAD118FF61FB1B9EA81AF9" bold="true" box="[151,439,1245,1271]" pageId="2" pageNumber="529">Type material examined</emphasis>
|
||
.
|
||
<taxonomicName id="2AC3F4C1FFEAD118FE33FB1B9DDC1AF9" authorityName="Brenske. Highest" authorityYear="1894" box="[453,707,1245,1271]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="philippinica">
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||
<emphasis id="DFB75350FFEAD118FE33FB1B9DDC1AF9" box="[453,707,1245,1271]" italics="true" pageId="2" pageNumber="529">Exopholis philippinica</emphasis>
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||
</taxonomicName>
|
||
,
|
||
<materialsCitation id="5DAB851FFFEAD118FD27FB1B9D141B31" ID-GBIF-Occurrence="3034045305" collectionCode="SMT" collectorName="Coll. C. Felsche" country="Philippines" county="Unicum" location="Brsk" municipality="Typus" pageId="2" pageNumber="529" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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||
<typeStatus id="327831E0FFEAD118FD27FB1B9C291AF9" box="[721,822,1245,1271]" pageId="2" pageNumber="529" type="holotype">holotype</typeStatus>
|
||
, female (
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||
<collectionCode id="8BD21787FFEAD118FC6AFB1B9CC31AF9" box="[924,988,1245,1271]" pageId="2" pageNumber="529">SMT</collectionCode>
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||
): “
|
||
<collectingCountry id="95D4CFD2FFEAD118FC0EFB1B9B9E1AF9" box="[1016,1153,1245,1271]" name="Philippines" pageId="2" pageNumber="529">Philippinen</collectingCountry>
|
||
”, “
|
||
<collectorName id="4036EA94FFEAD118FB57FB1B9A461AF9" box="[1185,1369,1245,1271]" pageId="2" pageNumber="529">Coll. C. Felsche</collectorName>
|
||
, Kauf 20, 1918”, “
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||
<taxonomicName id="2AC3F4C1FFEAD118FED2FAC79E841B15" authorityName="Motschulsky" authorityYear="1859" box="[292,411,1281,1307]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Exopholis</taxonomicName>
|
||
, philippinica, Type,
|
||
<specimenCount id="FBC544CBFFEAD118FD7FFAC79D811B15" box="[649,670,1281,1307]" pageId="2" pageNumber="529" type="female">♀</specimenCount>
|
||
,
|
||
<location id="E81CD999FFEAD118FD59FAC79DF61B15" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FD59FAC79DF61B15" box="[687,745,1281,1307]" country="Philippines" county="Unicum" municipality="Typus" name="Brsk" pageId="2" pageNumber="529">Brsk</location>
|
||
”, “
|
||
<collectingMunicipality id="0D181538FFEAD118FCFAFAC79C4A1B15" box="[780,853,1281,1307]" pageId="2" pageNumber="529">Typus</collectingMunicipality>
|
||
,
|
||
<taxonomicName id="2AC3F4C1FFEAD118FC92FAC79B431B15" authority=", Brenske" authorityName=", Brenske" authorityYear="1896" box="[868,1116,1281,1307]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="philippinica">philippinica, Brenske</taxonomicName>
|
||
”, “
|
||
<collectingCounty id="041DF7CEFFEAD118FB89FAC79BF91B15" box="[1151,1254,1281,1307]" pageId="2" pageNumber="529">Unicum</collectingCounty>
|
||
!”, “
|
||
<collectingCountry id="95D4CFD2FFEAD118FAFAFAC79A8C1B15" box="[1292,1427,1281,1307]" name="Philippines" pageId="2" pageNumber="529">Philippinen</collectingCountry>
|
||
” (
|
||
<figureCitation id="75F893C7FFEAD118FF69FAE39E131B4E" box="[159,268,1317,1344]" captionStart="FIGURES 4–6" captionStartId="4.[151,264,1849,1873]" captionTargetBox="[193,1329,216,1808]" captionTargetId="figure-20@4.[151,1436,181,1825]" captionTargetPageId="4" captionText="FIGURES 4–6. Exopholis philippinica Brenske, 1894, female, holotype from SMT. 4, habitus, dorsal aspect; 5, habitus, ventral aspect; 6, habitus, lateral aspect (photograph of the female type specimen from SMT, courtesy of Dr. Olaf Jaeger)." figureDoi="http://doi.org/10.5281/zenodo.4413360" httpUri="https://zenodo.org/record/4413360/files/figure.png" pageId="2" pageNumber="529">Figs. 4–6</figureCitation>
|
||
).
|
||
<taxonomicName id="2AC3F4C1FFEAD118FED6FAE39D141B31" authorityName="Moser. To Dr. Olaf Jaeger" authorityYear="1910" box="[288,523,1317,1343]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="corpulenta">
|
||
<emphasis id="DFB75350FFEAD118FED6FAE39D141B31" box="[288,523,1317,1343]" italics="true" pageId="2" pageNumber="529">Lepidiota corpulenta</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="5DAB851FFFEAD118FDEFFAE39A4B1B6D" ID-GBIF-Occurrence="3034045302" collectionCode="ZMHB" collectorName="Mrs M. Clemens" country="Philippines" location="Bu. of Sci." pageId="2" pageNumber="529" specimenCount="1" typeStatus="syntype">
|
||
<typeStatus id="327831E0FFEAD118FDEFFAE39D6C1B31" box="[537,627,1317,1343]" pageId="2" pageNumber="529" type="syntype">syntype</typeStatus>
|
||
, female (
|
||
<collectionCode id="8BD21787FFEAD118FD2FFAE39C2E1B31" box="[729,817,1317,1343]" pageId="2" pageNumber="529">ZMHB</collectionCode>
|
||
): “
|
||
<collectingCountry id="95D4CFD2FFEAD118FCBBFAE39CCB1B31" box="[845,980,1317,1343]" name="Philippines" pageId="2" pageNumber="529">Philippinen</collectingCountry>
|
||
, Mindanao”, “Collected by,
|
||
<collectorName id="4036EA94FFEAD118FAE7FAE39FC81B6D" pageId="2" pageNumber="529">Mrs M. Clemens</collectorName>
|
||
”, “Acc. No. 6883, Lot,
|
||
<location id="E81CD999FFEAD118FE29FA8F9D451B6D" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FE29FA8F9D451B6D" box="[479,602,1353,1379]" country="Philippines" name="Bu. of Sci." pageId="2" pageNumber="529">Bu. of Sci.</location>
|
||
, P.I.” (
|
||
<figureCitation id="75F893C7FFEAD118FD53FA8F9C0D1B6D" box="[677,786,1353,1379]" captionStart="FIGURES 7–9" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[227,1378,227,1833]" captionTargetId="figure-18@5.[151,1436,181,1843]" captionTargetPageId="5" captionText="FIGURES 7–9. Lepidiota corpulenta Moser, 1910, female, syntype from ZMHB. 7, habitus, dorsal aspect; 8, habitus, ventral aspect; 9, habitus, lateral aspect (photograph of the female from ZMHB, courtesy of Dr. Johannes Frisch and Dr. Joachim Willers)." figureDoi="http://doi.org/10.5281/zenodo.4413362" httpUri="https://zenodo.org/record/4413362/files/figure.png" pageId="2" pageNumber="529">Figs. 7–9</figureCitation>
|
||
). Both specimens were only examined from photos
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEAD118FF31FAAB9B8E1851" blockId="2.[151,1437,1245,2027]" pageId="2" pageNumber="529">
|
||
<emphasis id="DFB75350FFEAD118FF31FAAB9D2D1B89" bold="true" box="[199,562,1389,1415]" pageId="2" pageNumber="529">Additional material examined.</emphasis>
|
||
<materialsCitation id="5DAB851FFFEAD118FDCEFAAB9E091BA5" ID-GBIF-Occurrence="3034045309" collectingDate="1972-03" collectionCode="IRRI-AC" collectorName="San Leonardo & A. D. Pawar" country="Philippines" location="Ecija" municipality="Nueva" pageId="2" pageNumber="529" specimenCount="1" specimenCount-female="1">
|
||
<collectingCountry id="95D4CFD2FFEAD118FDCEFAAB9DA41B89" box="[568,699,1389,1415]" name="Philippines" pageId="2" pageNumber="529">Philippines</collectingCountry>
|
||
:
|
||
<emphasis id="DFB75350FFEAD118FD32FAAB9CD01B89" bold="true" box="[708,975,1389,1415]" pageId="2" pageNumber="529">
|
||
LUZON:
|
||
<collectingMunicipality id="0D181538FFEAD118FCC1FAAB9C9A1B89" box="[823,901,1389,1415]" pageId="2" pageNumber="529">Nueva</collectingMunicipality>
|
||
,
|
||
<location id="E81CD999FFEAD118FC67FAAB9CD01B89" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FC67FAAB9CD01B89" box="[913,975,1389,1415]" country="Philippines" municipality="Nueva" name="Ecija" pageId="2" pageNumber="529">Ecija</location>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="FBC544CBFFEAD118FC2BFAAB9B1F1B89" box="[989,1024,1389,1415]" pageId="2" pageNumber="529" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FBFBFAAB9BB41B89" box="[1037,1195,1389,1415]" pageId="2" pageNumber="529">San Leonardo</collectorName>
|
||
,
|
||
<date id="997DA982FFEAD118FB43FAAB9A0F1B89" box="[1205,1296,1389,1415]" pageId="2" pageNumber="529" value="1972-03">
|
||
<collectingDate id="8939506AFFEAD118FB43FAAB9A0F1B89" box="[1205,1296,1389,1415]" pageId="2" pageNumber="529" value="1972-03">iii-1972</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FAECFAAB9A831B89" box="[1306,1436,1389,1415]" pageId="2" pageNumber="529">A.D. Pawar</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FF69FA579E111BA5" box="[159,270,1425,1451]" pageId="2" pageNumber="529">IRRI-AC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FED2FA579A281BA5" ID-GBIF-Occurrence="3034045306" box="[292,1335,1425,1451]" collectingDate="1973-07-02" collectionCode="IRRI-AC" country="Philippines" location="Camarines Sur" pageId="2" pageNumber="529" specimenCount="1" specimenCount-female="1" stateProvince="Camarines Sur">
|
||
<emphasis id="DFB75350FFEAD118FED2FA579EC21BA5" bold="true" box="[292,477,1425,1451]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FED2FA579EC21BA5" box="[292,477,1425,1451]" country="Philippines" name="Camarines Sur" pageId="2" pageNumber="529">Camarines Sur</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="FBC544CBFFEAD118FE19FA579D0D1BA5" box="[495,530,1425,1451]" pageId="2" pageNumber="529" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FDD5FA579D441BA5" box="[547,603,1425,1451]" pageId="2" pageNumber="529">Iriga</collectorName>
|
||
,
|
||
<date id="997DA982FFEAD118FD9CFA579DEE1BA5" box="[618,753,1425,1451]" pageId="2" pageNumber="529" value="1973-07-02">
|
||
<collectingDate id="8939506AFFEAD118FD9CFA579DEE1BA5" box="[618,753,1425,1451]" pageId="2" pageNumber="529" value="1973-07-02">02-vii-1973</collectingDate>
|
||
</date>
|
||
,
|
||
<taxonomicName id="2AC3F4C1FFEAD118FD09FA579C8E1BA5" box="[767,913,1425,1451]" class="Liliopsida" family="Poaceae" genus="Oryza" kingdom="Plantae" order="Poales" pageId="2" pageNumber="529" phylum="Tracheophyta" rank="species" species="sativa">
|
||
<emphasis id="DFB75350FFEAD118FD09FA579C8E1BA5" box="[767,913,1425,1451]" italics="true" pageId="2" pageNumber="529">Oryza sativa</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<collectorName id="4036EA94FFEAD118FC33FA579B461BA5" box="[965,1113,1425,1451]" pageId="2" pageNumber="529">Upland, A.D</collectorName>
|
||
.
|
||
<collectorName id="4036EA94FFEAD118FB9EFA579BB11BA5" box="[1128,1198,1425,1451]" pageId="2" pageNumber="529">Pawar</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FB36FA579A301BA5" box="[1216,1327,1425,1451]" pageId="2" pageNumber="529">IRRI-AC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<emphasis id="DFB75350FFEAD118FAB3FA579FEC1BC1" bold="true" pageId="2" pageNumber="529">MIND- ANAO:</emphasis>
|
||
<materialsCitation id="5DAB851FFFEAD118FF0CFA739BBC1BC1" ID-GBIF-Occurrence="3034045307" box="[250,1187,1461,1487]" collectingDate="1986-06-28" collectionCode="IRRI-AC" collectorName="Claveria & A. Barrion" country="Philippines" location="Kalingagan" pageId="2" pageNumber="529" specimenCount="6" specimenCount-male="6" stateProvince="Misamis Oriental">
|
||
<emphasis id="DFB75350FFEAD118FF0CFA739ED41BC1" bold="true" box="[250,459,1461,1487]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FF0CFA739ED41BC1" box="[250,459,1461,1487]" country="Philippines" name="Misamis Oriental" pageId="2" pageNumber="529">Misamis Oriental</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="FBC544CBFFEAD118FE2CFA739D0A1BC1" box="[474,533,1461,1487]" pageId="2" pageNumber="529" type="male">6♂♂</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FDD6FA739D9D1BC1" box="[544,642,1461,1487]" pageId="2" pageNumber="529">Claveria</collectorName>
|
||
,
|
||
<location id="E81CD999FFEAD118FD7BFA739C0E1BC1" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FD7BFA739C0E1BC1" box="[653,785,1461,1487]" country="Philippines" name="Kalingagan" pageId="2" pageNumber="529" stateProvince="Misamis Oriental">Kalingagan</location>
|
||
,
|
||
<date id="997DA982FFEAD118FCEAFA739C841BC1" box="[796,923,1461,1487]" pageId="2" pageNumber="529" value="1986-06-28">
|
||
<collectingDate id="8939506AFFEAD118FCEAFA739C841BC1" box="[796,923,1461,1487]" pageId="2" pageNumber="529" value="1986-06-28">28-vi-1986</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FC53FA739B011BC1" box="[933,1054,1461,1487]" pageId="2" pageNumber="529">A. Barrion</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FBDAFA739B841BC1" box="[1068,1179,1461,1487]" pageId="2" pageNumber="529">IRRI-AC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FB58FA739DDC1BFD" ID-GBIF-Occurrence="3034045303" collectingDate="1950-05-01" collectionCode="NMM" collectorName="Camp Maramog & P. Anonuevo" country="Philippines" location="Bukidnon" pageId="2" pageNumber="529" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Bukidnon">
|
||
<emphasis id="DFB75350FFEAD118FB58FA739A3B1BC1" bold="true" box="[1198,1316,1461,1487]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FB58FA739A3B1BC1" box="[1198,1316,1461,1487]" country="Philippines" name="Bukidnon" pageId="2" pageNumber="529">Bukidnon</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="FBC544CBFFEAD118FAC5FA739A741BC1" box="[1331,1387,1461,1487]" pageId="2" pageNumber="529" type="male">2♂♂</specimenCount>
|
||
<specimenCount id="FBC544CBFFEAD118FA84FA739A8A1BC1" box="[1394,1429,1461,1487]" pageId="2" pageNumber="529" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FF61FA1F9E4E1BFD" box="[151,337,1497,1523]" pageId="2" pageNumber="529">Camp Maramog</collectorName>
|
||
,
|
||
<date id="997DA982FFEAD118FEAAFA1F9EDA1BFA" box="[348,453,1497,1524]" pageId="2" pageNumber="529" value="1950-05-01">
|
||
<collectingDate id="8939506AFFEAD118FEAAFA1F9EDA1BFA" box="[348,453,1497,1524]" pageId="2" pageNumber="529" value="1950-05-01">1-v-1950</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FE27FA1F9D7F1BFD" box="[465,608,1497,1523]" pageId="2" pageNumber="529">P. Anonuevo</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FD98FA1F9DA51BFD" box="[622,698,1497,1523]" pageId="2" pageNumber="529">NMM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FD38FA1F9A101BFD" ID-GBIF-Occurrence="3034045308" box="[718,1295,1497,1523]" collectionCode="UNSM" country="Philippines" pageId="2" pageNumber="529" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
|
||
<specimenCount id="FBC544CBFFEAD118FD38FA1F9C101BFD" box="[718,783,1497,1523]" pageId="2" pageNumber="529" type="male">3♂♂</specimenCount>
|
||
<specimenCount id="FBC544CBFFEAD118FCF9FA1F9C341BFD" box="[783,811,1497,1523]" pageId="2" pageNumber="529" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FCC1FA1F9CB11BFD" box="[823,942,1497,1523]" pageId="2" pageNumber="529">Tangcolan</collectorName>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FC4FFA1F9B4D1BFD" box="[953,1106,1497,1523]" pageId="2" pageNumber="529">Bukinon, C.F</collectorName>
|
||
.
|
||
<collectorName id="4036EA94FFEAD118FBABFA1F9BBF1BFD" box="[1117,1184,1497,1523]" pageId="2" pageNumber="529">Baker</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FB59FA1F9A191BFD" box="[1199,1286,1497,1523]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2" pageNumber="529">UNSM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FAECFA1F9DC61819" ID-GBIF-Occurrence="3034045311" collectingDate="1970-06-21" collectionCode="UPMNH" collectorName="Casisang & L. C. Raros" country="Philippines" pageId="2" pageNumber="529" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="FBC544CBFFEAD118FAECFA1F9A211BFD" box="[1306,1342,1497,1523]" pageId="2" pageNumber="529" type="male">1♂</specimenCount>
|
||
, Malay- balay,
|
||
<collectorName id="4036EA94FFEAD118FF16FA3B9E571819" box="[224,328,1533,1559]" pageId="2" pageNumber="529">Casisang</collectorName>
|
||
,
|
||
<date id="997DA982FFEAD118FEA5FA3B9ECD1819" box="[339,466,1533,1559]" pageId="2" pageNumber="529" value="1970-06-21">
|
||
<collectingDate id="8939506AFFEAD118FEA5FA3B9ECD1819" box="[339,466,1533,1559]" pageId="2" pageNumber="529" value="1970-06-21">21-vi-1970</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FE2BFA3B9D481819" box="[477,599,1533,1559]" pageId="2" pageNumber="529">L.C. Raros</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FD93FA3B9DCE1819" box="[613,721,1533,1559]" pageId="2" pageNumber="529">UPMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FD12FA3B9BF61819" ID-GBIF-Occurrence="3034045310" box="[740,1257,1533,1559]" collectingDate="2012-05-10" collectionCode="UPMNH" collectorName="Navasero" country="Philippines" pageId="2" pageNumber="529" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
|
||
<specimenCount id="FBC544CBFFEAD118FD12FA3B9C181819" box="[740,775,1533,1559]" pageId="2" pageNumber="529" type="male">1♂</specimenCount>
|
||
<specimenCount id="FBC544CBFFEAD118FCF8FA3B9C581819" box="[782,839,1533,1559]" pageId="2" pageNumber="529" type="female">3♀♀</specimenCount>
|
||
,
|
||
<date id="997DA982FFEAD118FCA2FA3B9CD41819" box="[852,971,1533,1559]" pageId="2" pageNumber="529" value="2012-05-10">
|
||
<collectingDate id="8939506AFFEAD118FCA2FA3B9CD41819" box="[852,971,1533,1559]" pageId="2" pageNumber="529" value="2012-05-10">10-v-2012</collectingDate>
|
||
</date>
|
||
, M.
|
||
<collectorName id="4036EA94FFEAD118FC0BFA3B9B781819" box="[1021,1127,1533,1559]" pageId="2" pageNumber="529">Navasero</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FB83FA3B9BFE1819" box="[1141,1249,1533,1559]" pageId="2" pageNumber="529">UPMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FB02FA3B9D501835" ID-GBIF-Occurrence="3034045312" collectionCode="UPMNH" collectorName="Banga & Uichanco" country="Philippines" location="Cotabato" pageId="2" pageNumber="529" specimenCount="1" specimenCount-female="1" stateProvince="South Cotabato">
|
||
<emphasis id="DFB75350FFEAD118FB02FA3B9A741819" bold="true" box="[1268,1387,1533,1559]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FB02FA3B9A781819" box="[1268,1383,1533,1559]" country="Philippines" name="South Cotabato" pageId="2" pageNumber="529">Cotabato</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="FBC544CBFFEAD118FA84FA3B9A8A1819" box="[1394,1429,1533,1559]" pageId="2" pageNumber="529" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FF61F9E79FFE1835" box="[151,225,1569,1595]" pageId="2" pageNumber="529">Banga</collectorName>
|
||
, 24-1950,
|
||
<collectorName id="4036EA94FFEAD118FEA8F9E79ED61835" box="[350,457,1569,1595]" pageId="2" pageNumber="529">Uichanco</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FE2DF9E79D581835" box="[475,583,1569,1595]" pageId="2" pageNumber="529">UPMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FDAAF9E79FEC1851" ID-GBIF-Occurrence="3034045304" collectionCode="NMM" country="Philippines" location="Mt. Makaayat" pageId="2" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Davao del Norte">
|
||
<emphasis id="DFB75350FFEAD118FDAAF9E79C361835" bold="true" box="[604,809,1569,1595]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FDAAF9E79C361835" box="[604,809,1569,1595]" country="Philippines" name="Davao del Norte" pageId="2" pageNumber="529">Davao Del Norte</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="FBC544CBFFEAD118FCCDF9E79C401835" box="[827,863,1569,1595]" pageId="2" pageNumber="529" type="male">1♂</specimenCount>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FC99F9E79CE61835" box="[879,1017,1569,1595]" pageId="2" pageNumber="529">Malamboon</collectorName>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FBF1F9E79B721835" box="[1031,1133,1569,1595]" pageId="2" pageNumber="529">Malabog</collectorName>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FB8DF9E79BF11835" box="[1147,1262,1569,1595]" pageId="2" pageNumber="529">Paquibato</collectorName>
|
||
,
|
||
<location id="E81CD999FFEAD118FB0AF9E79A831835" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FB0AF9E79A831835" box="[1276,1436,1569,1595]" country="Philippines" name="Mt. Makaayat" pageId="2" pageNumber="529" stateProvince="Davao del Norte">Mt. Makaayat</location>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FF69F9839FF41851" box="[159,235,1605,1631]" pageId="2" pageNumber="529">NMM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5DAB851FFFEAD118FF09F9839EFD1851" ID-GBIF-Occurrence="3034045313" box="[255,482,1605,1631]" country="Philippines" location="Davao Del Sur" pageId="2" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Davao del Sur">
|
||
<emphasis id="DFB75350FFEAD118FF09F9839EA91851" bold="true" box="[255,438,1605,1631]" pageId="2" pageNumber="529">
|
||
<collectingRegion id="2F0741A0FFEAD118FF09F9839EAE1851" box="[255,433,1605,1631]" country="Philippines" name="Davao del Sur" pageId="2" pageNumber="529">Davao Del Sur</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="FBC544CBFFEAD118FE4BF9839EFD1851" box="[445,482,1605,1631]" pageId="2" pageNumber="529" type="male">1♂</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="5DAB851FFFEAD118FE18F9839B911851" ID-GBIF-Occurrence="3034045301" box="[494,1166,1605,1632]" collectingDate="1957-07-04" collectionCode="UPMNH" collectorName="P. M. Narciso" country="Philippines" location="Tuganay" municipality="City" pageId="2" pageNumber="529" specimenCount="1" stateProvince="Davao">
|
||
<collectingRegion id="2F0741A0FFEAD118FE18F9839D281851" box="[494,567,1605,1631]" country="Philippines" name="Davao" pageId="2" pageNumber="529">Davao</collectingRegion>
|
||
<collectingMunicipality id="0D181538FFEAD118FDC8F9839D701851" box="[574,623,1605,1631]" pageId="2" pageNumber="529">City</collectingMunicipality>
|
||
,
|
||
<location id="E81CD999FFEAD118FD8CF9839DC31851" LSID="urn:lsid:plazi:treatment:656A3E54FFEAD112FF61FC149A851D37:E81CD999FFEAD118FD8CF9839DC31851" box="[634,732,1605,1631]" country="Philippines" municipality="City" name="Tuganay" pageId="2" pageNumber="529" stateProvince="Davao">Tuganay</location>
|
||
,
|
||
<date id="997DA982FFEAD118FD11F9839C71186E" box="[743,878,1605,1632]" pageId="2" pageNumber="529" value="1957-07-04">
|
||
<collectingDate id="8939506AFFEAD118FD11F9839C71186E" box="[743,878,1605,1632]" pageId="2" pageNumber="529" value="1957-07-04">04-vii-1957</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4036EA94FFEAD118FC8CF9839B141851" box="[890,1035,1605,1631]" pageId="2" pageNumber="529">P.M. Narciso</collectorName>
|
||
(
|
||
<collectionCode id="8BD21787FFEAD118FBECF9839B991851" box="[1050,1158,1605,1631]" pageId="2" pageNumber="529">UPMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5D9DCC9FFEAD119FF31F9AF9FF4199B" lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529" type="description">
|
||
<paragraph id="ED7C8F42FFEAD118FF31F9AF9A59191D" blockId="2.[151,1437,1245,2027]" pageId="2" pageNumber="529">
|
||
<emphasis id="DFB75350FFEAD118FF31F9AF9E6A188D" bold="true" box="[199,373,1641,1667]" pageId="2" pageNumber="529">Redescription.</emphasis>
|
||
Males: length 19.0–22.0 mm, width 10.0–12.0 mm. Body ovoid, lustrous, monochromatic to bichromatic; dark reddish brown, blackish brown to light orange; head, pronotum and scutellum usually much darker than elytra in bichromatic specimens. Dorsum covered occasionally with minute, lanceolate, yellowish-white scales; except for a row of few large, overlapping, elongate-lanceolate, parallel laterally blunt scales adjacent to fronto-lateral margin apically. Venter same dominant coloration as head and pronotum of dorsum (
|
||
<figureCitation id="75F893C7FFEAD118FB13F93F9A27191D" box="[1253,1336,1785,1811]" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1981,2005]" captionTargetBox="[320,1230,185,1933]" captionTargetId="figure-20@6.[258,1330,181,1957]" captionTargetPageId="6" captionText="FIGURES 10–12. Exopholis philippinica Brenske, 1894, male from IRRI. 10, habitus, dorsal aspect; 11, male genitalia, dorsal aspect; 12, male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4413366" httpUri="https://zenodo.org/record/4413366/files/figure.png" pageId="2" pageNumber="529">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEAD119FF31F8DB9DCA1F6B" blockId="2.[151,1437,1245,2027]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529">
|
||
<emphasis id="DFB75350FFEAD118FF31F8DB9E141939" box="[199,267,1821,1847]" italics="true" pageId="2" pageNumber="529">Head.</emphasis>
|
||
Labrum with subrugose to rugose punctures; posterior half evenly sloping downward towards posterior margin medially; each side of posterior margin with long, thick, stiff, fringed, bristle-like yellowish to orange setae; surface with few, very long yellowish to orange stiff hairs. Clypeus with medial anterior margin nearly straight to weakly convex anteriorly, without medial cleft; with round punctures of variable size near posterior margin anteriorly; longitudinally flattened laterally; margin narrowly, slightly deflexed backward laterally; surface with dense, shallow round to irregular-shaped to rugose punctures dorsally; each puncture with minute, narrow, elongate, yel- lowish white scales; not extending beyond length of puncture. Vertex with moderate to sparsely spaced, minute, round punctures; subrugose; scales distinctly larger on each side adjacent lateral margin and ocular canthus.
|
||
<collectingRegion id="2F0741A0FFEBD119FA96FF7D9FDC1EF7" country="Myanmar" name="Chin" pageId="3" pageNumber="530">Mentum</collectingRegion>
|
||
sub-anterior margin with wide, triangulate carina; above carina with deep, rugose punctures and groove; below carina smooth, lustrous; without hairs and scales; adjacent posterior margin of mentum with few, short, fine whitish, stiff setae. Terminal maxillary palpomere cylindrical, spindle-shaped; slightly longer in males than females. Lamellae length 1.0–
|
||
<quantity id="2A3B22A7FFEBD119FEC8FE8D9E8A1F6B" box="[318,405,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="530" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, slightly longer than female.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FEA99E111C17" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FEA99E291F86" box="[199,310,367,392]" italics="true" pageId="3" pageNumber="530">Pronotum</emphasis>
|
||
anterior margin widely concave, not carinate medially; margin slightly to strongly carinate towards anterior angle; anterior angle rectangular, rounded apically; lateral margin convex, narrow carinate; posterior angle obtuse, round apically; apical margin smooth, not carinate, concave, widely rounded medially; each side slightly sloping downward; surface with evenly spaced round to subrugose punctures; scales minute, much larger in clypeus and elytra.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FDE59E051DAB" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FDE59EE31C32" box="[199,508,547,573]" italics="true" pageId="3" pageNumber="530">Ventral surface of pronotum</emphasis>
|
||
. Prosternum with deep mesal groove or depression; densely setose medially, sparsely setose on each side, without scales. Prosternal process triangulate, short, tapering towards anterior margin, with blunt apex, wide basally; convex laterally. Mesoventral process triangulate, rounded apically; rounded laterally; with or without acicular scales apically; surrounded with moderate-sized, stiff, yellowish-white hairs. Metaventral process length
|
||
<quantity id="2A3B22A7FFEBD119FEC9FD759ED91CC3" box="[319,454,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="3" pageNumber="530" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
|
||
(n=10), short; thickly carinate medially, narrowing towards basal margin; without hairs and scales medially. Metaventrite sparsely to evenly covered with long stiff, yellowish-white hairs except medially; lustrous medially; with or without few, dense, long lanceolate to acicular whitish to yellowish scales near posterior angle and posterior margin; mesepisternum densely covered with long, lanceolate, tapered apex scales; metepisternum and metepimera densely covered with short to moderate-sized whitish to yellowish white hairs; usually without hairs and scales medially; mesepisternum densely covered with same scales, with numerous long, stiff yellowish white hairs.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FC699BC31DE3" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FC699FEE1DC6" box="[199,241,943,968]" italics="true" pageId="3" pageNumber="530">Leg</emphasis>
|
||
. Foretibiae tridentate; apical and medial dents distinctly large; basal dent distinctly small. Longer and shorter metatibial spurs very narrow, tapered apically. Metatibiae with 17–27 spicules posteriorly.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FC319E9D1A57" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FC319E2A1A1F" box="[199,309,1015,1041]" italics="true" pageId="3" pageNumber="530">Scutellum</emphasis>
|
||
posterior margin sub-circular; anterior margin widely, evenly concave, smooth, rounded laterally; lateral margin minutely carinate, not carinate medially; surface with slightly larger scales than clypeus, vertex, pronotum and elytra.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FBA59C6E1B77" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FBA59E051A73" box="[199,282,1123,1149]" italics="true" pageId="3" pageNumber="530">Elytron</emphasis>
|
||
anterior margin slightly sloping downward towards scutellum, smooth, not carinate; anterior angle obtuse, round apically; lateral margin evenly convex; upper lateral margin narrowly carinate and explanate; lower lateral margin thinly carinate medially, disappearing towards posterior angle; posterior angle widely rounded, smooth, not carinate; apical margin not carinate, smooth; sutural angle slightly obtuse, round apically; sutural margin with distinctly wide costa, lustrous, mostly non-scaled; with four costae; costa 1 distinct, wide, complete; costa 2 narrow, distinct, incomplete towards anterior margin, basally attached to costa 3; costa 3 faint, incomplete towards anterior margin; costa 4 indistinct, faint, incomplete towards anterior margin; apical knob of elytra without patch of scales; surface with subrugose punctures, with same scales as pronotum.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31FA459FE01807" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31FA459E2E1B93" box="[199,305,1411,1437]" italics="true" pageId="3" pageNumber="530">Abdomen</emphasis>
|
||
. Adjacent to anterior margin of abdominal sternites III–VI with narrow, lustrous, non-scaled area; wider towards medially, narrower towards lateral margin; abdominal sternites with short, ovoid, lanceolate yellowish-white scales, most dense towards lateral margin; mostly with short, elongate, lanceolate scales, sparsely spaced medially.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31F9D59C7A187B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31F9D59E2D1823" box="[199,306,1555,1581]" italics="true" pageId="3" pageNumber="530">Pygidium</emphasis>
|
||
. Anterior margin evenly convex, thinly carinate; lateral margin thickly carinate, narrowing towards apical margin; apical margin rounded, narrowly explanate, not carinate; surface evenly covered with short lanceolate scales, tapered or round apically; with subrugose punctures.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31F9B99EC3190B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31F9B99E231897" box="[199,316,1663,1689]" italics="true" pageId="3" pageNumber="530">Aedeagus.</emphasis>
|
||
Paramere and phallobase combined length
|
||
<quantity id="2A3B22A7FFEBD119FCDAF9B99CA81897" box="[812,951,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="7.7" metricValueMin="7.4" pageId="3" pageNumber="530" unit="mm" value="7.55" valueMax="7.7" valueMin="7.4">7.4–7.7 mm</quantity>
|
||
. Phallobase dorsal anterior margin slightly convex mesally (
|
||
<figureCitation id="75F893C7FFEBD119FEA2F9659EBA18B3" box="[340,421,1699,1725]" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1981,2005]" captionTargetBox="[320,1230,185,1933]" captionTargetId="figure-20@6.[258,1330,181,1957]" captionTargetPageId="6" captionText="FIGURES 10–12. Exopholis philippinica Brenske, 1894, male from IRRI. 10, habitus, dorsal aspect; 11, male genitalia, dorsal aspect; 12, male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4413366" httpUri="https://zenodo.org/record/4413366/files/figure.png" pageId="3" pageNumber="530">Fig. 11</figureCitation>
|
||
). Paramere dorsal posterior margin triangulate; with inverted lanceolate-shaped depression laterally. Apical process short, triangulate laterally; apex round; pointed at 45
|
||
<emphasis id="DFB75350FFEBD119FC04F9009CE218EE" bold="true" box="[1010,1021,1734,1760]" pageId="3" pageNumber="530">°</emphasis>
|
||
angle; base of apical process narrow, elongate, neck-like (
|
||
<figureCitation id="75F893C7FFEBD119FE8AF92D9ED0190B" box="[380,463,1771,1797]" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1981,2005]" captionTargetBox="[320,1230,185,1933]" captionTargetId="figure-20@6.[258,1330,181,1957]" captionTargetPageId="6" captionText="FIGURES 10–12. Exopholis philippinica Brenske, 1894, male from IRRI. 10, habitus, dorsal aspect; 11, male genitalia, dorsal aspect; 12, male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4413366" httpUri="https://zenodo.org/record/4413366/files/figure.png" pageId="3" pageNumber="530">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEBD119FF31F8C99FF4199B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31F8C99E3A1927" bold="true" box="[199,293,1807,1833]" pageId="3" pageNumber="530">Female.</emphasis>
|
||
Length
|
||
<quantity id="2A3B22A7FFEBD119FE72F8C89D361927" box="[388,553,1806,1833]" metricMagnitude="-2" metricUnit="m" metricValue="2.315" metricValueMax="2.65" metricValueMin="1.98" pageId="3" pageNumber="530" unit="mm" value="23.15" valueMax="26.5" valueMin="19.8">19.8–26.5 mm</quantity>
|
||
, width
|
||
<quantity id="2A3B22A7FFEBD119FD8BF8C99DE71927" box="[637,760,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="530" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
|
||
. Antennal club length
|
||
<quantity id="2A3B22A7FFEBD119FC03F8C99B751927" box="[1013,1130,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="530" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1–1.1 mm</quantity>
|
||
, slightly shorter than male. Terminal maxillary palpomere cylindrical, spindle-shaped; slightly shorter in male. Foretibiae broad. Longer and shorter metatibial spurs little wider, slightly broad or rounded apically. Metatibiae with 23–30 spicules posteriorly (n=20).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5D9DCC9FFEBD119FF31F8589C2D19B4" box="[199,818,1950,1978]" pageId="3" pageNumber="530" type="distribution">
|
||
<paragraph id="ED7C8F42FFEBD119FF31F8589C2D19B4" blockId="3.[151,1437,151,2013]" box="[199,818,1950,1978]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31F8589E7E19B6" bold="true" box="[199,353,1950,1976]" pageId="3" pageNumber="530">Distribution.</emphasis>
|
||
<collectingCountry id="95D4CFD2FFEBD119FE91F8599EFA19B7" box="[359,485,1951,1977]" name="Philippines" pageId="3" pageNumber="530">Philippines</collectingCountry>
|
||
(Luzon, Visayas, Mindanao).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5D9DCC9FFEBD119FF31F8059ECC19D3" box="[199,467,1987,2013]" pageId="3" pageNumber="530" type="biology_ecology">
|
||
<paragraph id="ED7C8F42FFEBD119FF31F8059ECC19D3" blockId="3.[151,1437,151,2013]" box="[199,467,1987,2013]" pageId="3" pageNumber="530">
|
||
<emphasis id="DFB75350FFEBD119FF31F8059E4919D3" bold="true" box="[199,342,1987,2013]" pageId="3" pageNumber="530">Host plants.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B9BCDFCAFFECD11EFF61F8FF9A03197B" ID-DOI="http://doi.org/10.5281/zenodo.4413360" ID-Zenodo-Dep="4413360" httpUri="https://zenodo.org/record/4413360/files/figure.png" pageId="4" pageNumber="531" startId="4.[151,264,1849,1873]" targetBox="[193,1329,216,1808]" targetPageId="4">
|
||
<paragraph id="ED7C8F42FFECD11EFF61F8FF9A03197B" blockId="4.[151,1436,1849,1910]" pageId="4" pageNumber="531">
|
||
<emphasis id="DFB75350FFECD11EFF61F8FF9E25195F" bold="true" box="[151,314,1849,1873]" pageId="4" pageNumber="531">FIGURES 4–6.</emphasis>
|
||
<taxonomicName id="2AC3F4C1FFECD11EFEC9F8FF9DA2195F" authority="Brenske, 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[319,701,1849,1873]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="531" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFECD11EFEC9F8FF9D3B195F" box="[319,548,1849,1873]" italics="true" pageId="4" pageNumber="531">Exopholis philippinica</emphasis>
|
||
<bibRefCitation id="8952F2B3FFECD11EFDDCF8FF9DA2195F" author="Brenske, E." box="[554,701,1849,1873]" pageId="4" pageNumber="531" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
, female, holotype from SMT. 4, habitus, dorsal aspect; 5, habitus, ventral aspect; 6, habitus, lateral aspect (photograph of the female type specimen from SMT, courtesy of Dr. Olaf Jaeger).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9BCDFCAFFEDD11FFF61F88D9FDB19A5" ID-DOI="http://doi.org/10.5281/zenodo.4413362" ID-Zenodo-Dep="4413362" httpUri="https://zenodo.org/record/4413362/files/figure.png" pageId="5" pageNumber="532" startId="5.[151,264,1867,1891]" targetBox="[227,1378,227,1833]" targetPageId="5">
|
||
<paragraph id="ED7C8F42FFEDD11FFF61F88D9FDB19A5" blockId="5.[151,1436,1867,1963]" pageId="5" pageNumber="532">
|
||
<emphasis id="DFB75350FFEDD11FFF61F88D9E24196D" bold="true" box="[151,315,1867,1891]" pageId="5" pageNumber="532">FIGURES 7–9.</emphasis>
|
||
<taxonomicName id="2AC3F4C1FFEDD11FFEB4F88D9DBE196D" authority="Moser, 1910" authorityName="Moser. To Dr. Olaf Jaeger" authorityYear="1910" box="[322,673,1867,1891]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="532" phylum="Arthropoda" rank="species" species="corpulenta">
|
||
<emphasis id="DFB75350FFEDD11FFEB4F88D9D07196D" box="[322,536,1867,1891]" italics="true" pageId="5" pageNumber="532">Lepidiota corpulenta</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEDD11FFDE9F88A9DBE196D" author="Moser, J." box="[543,673,1867,1891]" pageId="5" pageNumber="532" pagination="149 - 150" refId="ref4269" refString="Moser, J. (1910) Neue Coleoptera Lamellicornia von den Philippen. The Philippine Journal of Science, 5, 149 - 150." type="journal article" year="1910">Moser, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
, female, syntype from ZMHB. 7, habitus, dorsal aspect; 8, habitus, ventral aspect; 9, habitus, lateral aspect (photograph of the female from ZMHB, courtesy of Dr. Johannes Frisch and Dr. Joachim Willers).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9BCDFCAFFEED11CFF61F87B9D2D19F7" ID-DOI="http://doi.org/10.5281/zenodo.4413366" ID-Zenodo-Dep="4413366" httpUri="https://zenodo.org/record/4413366/files/figure.png" pageId="6" pageNumber="533" startId="6.[151,264,1981,2005]" targetBox="[320,1230,185,1933]" targetPageId="6">
|
||
<paragraph id="ED7C8F42FFEED11CFF61F87B9D2D19F7" blockId="6.[151,1436,1981,2041]" pageId="6" pageNumber="533">
|
||
<emphasis id="DFB75350FFEED11CFF61F87B9E4B19DB" bold="true" box="[151,340,1981,2005]" pageId="6" pageNumber="533">FIGURES 10–12.</emphasis>
|
||
<taxonomicName id="2AC3F4C1FFEED11CFEACF87B9DC519DB" authority="Brenske, 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[346,730,1981,2005]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="533" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFEED11CFEACF87B9D2019DB" box="[346,575,1981,2005]" italics="true" pageId="6" pageNumber="533">Exopholis philippinica</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEED11CFDB0F87B9DC519DB" author="Brenske, E." box="[582,730,1981,2005]" pageId="6" pageNumber="533" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
, male from IRRI. 10, habitus, dorsal aspect; 11, male genitalia, dorsal aspect; 12, male genitalia, lateral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9BCDFCAFFEFD11DFF61F97F9BEB18DF" ID-DOI="http://doi.org/10.5281/zenodo.4413368" ID-Zenodo-Dep="4413368" box="[151,1268,1721,1745]" httpUri="https://zenodo.org/record/4413368/files/figure.png" pageId="7" pageNumber="534" startId="7.[151,250,1721,1745]" targetBox="[245,1279,181,1697]" targetPageId="7">
|
||
<paragraph id="ED7C8F42FFEFD11DFF61F97F9BEB18DF" blockId="7.[151,1268,1721,1745]" box="[151,1268,1721,1745]" pageId="7" pageNumber="534">
|
||
<emphasis id="DFB75350FFEFD11DFF61F97F9E3F18DF" bold="true" box="[151,288,1721,1745]" pageId="7" pageNumber="534">FIGURE 13.</emphasis>
|
||
Distribution of examined specimens of
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFD40F97F9B2A18DF" authority="Brenske, 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[694,1077,1721,1745]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFEFD11DFD40F97F9C8318DF" box="[694,924,1721,1745]" italics="true" pageId="7" pageNumber="534">Exopholis philippinica</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEFD11DFC55F97F9B2A18DF" author="Brenske, E." box="[931,1077,1721,1745]" pageId="7" pageNumber="534" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
in the Philippines.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A5D9DCC9FFEFD112FF31F8C69A851D37" lastPageId="8" lastPageNumber="535" pageId="7" pageNumber="534" type="discussion">
|
||
<paragraph id="ED7C8F42FFEFD11DFF31F8C69C0C19FC" blockId="7.[151,1437,1791,2034]" pageId="7" pageNumber="534">
|
||
<emphasis id="DFB75350FFEFD11DFF31F8C69E241914" bold="true" box="[199,315,1792,1818]" pageId="7" pageNumber="534">Remarks.</emphasis>
|
||
The species can be distinguished from other species of
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFC40F9399B391917" authorityName="Motschulsky" authorityYear="1859" box="[950,1062,1791,1817]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEFD11DFC40F9399B391917" box="[950,1062,1791,1817]" italics="true" pageId="7" pageNumber="534">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters. Elytra with four distinct costae, sutural costa distinctly wide while the rest are narrow and less distinct. The pygidium has the apical margin rounded. The clypeus is distinctly upturned laterally. The apical margin of the paramere of the male genitalia is triangulate and widely rounded apically while the basal paramere is inverted, lanceolate-shaped and with a lateral depression.
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFD6CF8499C8619A7" authorityName="Brenske. Highest" authorityYear="1894" box="[666,921,1935,1961]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFEFD11DFD6CF8499C8619A7" box="[666,921,1935,1961]" italics="true" pageId="7" pageNumber="534">Exopholis philippinica</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from other species with four costae,
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFED2F8729EA819C3" baseAuthorityName="Weidemann" baseAuthorityYear="1819" box="[292,439,1971,1997]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="hypoleuca">
|
||
<emphasis id="DFB75350FFEFD11DFED2F8729EA819C3" box="[292,439,1971,1997]" italics="true" pageId="7" pageNumber="534">E. hypoleuca</emphasis>
|
||
</taxonomicName>
|
||
with variable number of costae,
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFCC6F8729CD519C3" authorityName="Brenske" authorityYear="1894" box="[816,970,1971,1997]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="DFB75350FFEFD11DFCC6F8729CD519C3" box="[816,970,1971,1997]" italics="true" pageId="7" pageNumber="534">E. borneensis</emphasis>
|
||
</taxonomicName>
|
||
with four, which are different from each other,
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFF28F81E9E6619FF" authorityName="Waterhouse" authorityYear="1867" box="[222,377,2007,2033]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="lacordairei">
|
||
<emphasis id="DFB75350FFEFD11DFF28F81E9E6619FF" box="[222,377,2007,2033]" italics="true" pageId="7" pageNumber="534">E. lacordairei</emphasis>
|
||
</taxonomicName>
|
||
with five and
|
||
<taxonomicName id="2AC3F4C1FFEFD11DFDECF81E9D9419FF" authorityName="Lansberge" authorityYear="1879" box="[538,651,2008,2033]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="534" phylum="Arthropoda" rank="species" species="pinguis">
|
||
<emphasis id="DFB75350FFEFD11DFDECF81E9D9419FF" box="[538,651,2008,2033]" italics="true" pageId="7" pageNumber="534">E. pinguis</emphasis>
|
||
</taxonomicName>
|
||
with seven.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFE0D112FF31FF519FF11FA3" blockId="8.[151,1437,151,825]" pageId="8" pageNumber="535">
|
||
Sexual dimorphism is not strongly developed, the antennal club is slightly longer in males than in females. The terminal maxillary palpomere is slightly longer in males than in females. The longer and shorter metatibial spur has a tapered apex in males while it has a slightly broader apex in females. The number of posterior metatibial spicules is overlapping in range in both sexes. The species also exhibits polyphenic trait with the occurrence of variable body coloration; some specimens are reddish brown while others are blackish brown.
|
||
<bibRefCitation id="8952F2B3FFE0D112FBE8FEE19BD41F4F" author="Brenske, E." box="[1054,1227,295,321]" pageId="8" pageNumber="535" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske (1894)</bibRefCitation>
|
||
also observed this color variability in the populations of
|
||
<taxonomicName id="2AC3F4C1FFE0D112FDBCFE8D9DC11F6B" baseAuthorityName="Weidemann" baseAuthorityYear="1819" box="[586,734,331,357]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="hypoleuca">
|
||
<emphasis id="DFB75350FFE0D112FDBCFE8D9DC11F6B" box="[586,734,331,357]" italics="true" pageId="8" pageNumber="535">E. hypoleuca</emphasis>
|
||
</taxonomicName>
|
||
from Borneo,
|
||
<collectingRegion id="2F0741A0FFE0D112FC7EFE8D9CF71F6B" box="[904,1000,331,357]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="535">Sumatra</collectingRegion>
|
||
, Nias, and
|
||
<collectingCountry id="95D4CFD2FFE0D112FB9EFE8D9BA31F6B" box="[1128,1212,331,357]" name="Indonesia" pageId="8" pageNumber="535">Ambon</collectingCountry>
|
||
he found the elytra light brown to dark chestnut-brown while some individuals from
|
||
<collectingRegion id="2F0741A0FFE0D112FC8FFEA99CC91F87" box="[889,982,367,393]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="535">Sumatra</collectingRegion>
|
||
were entirely black and with brown pygidium.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFE0D112FF31FE719A4A1C6F" blockId="8.[151,1437,151,825]" pageId="8" pageNumber="535">
|
||
<bibRefCitation id="8952F2B3FFE0D112FF31FE719E7B1FDF" author="Moser, J." box="[199,356,439,465]" pageId="8" pageNumber="535" pagination="149 - 150" refId="ref4269" refString="Moser, J. (1910) Neue Coleoptera Lamellicornia von den Philippen. The Philippine Journal of Science, 5, 149 - 150." type="journal article" year="1910">Moser (1910)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="2AC3F4C1FFE0D112FE10FE719D9F1FDF" authorityName="Moser. To Dr. Olaf Jaeger" authorityYear="1910" box="[486,640,439,465]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="corpulenta">
|
||
<emphasis id="DFB75350FFE0D112FE10FE719D9F1FDF" box="[486,640,439,465]" italics="true" pageId="8" pageNumber="535">L. corpulenta</emphasis>
|
||
</taxonomicName>
|
||
from three female specimens collected on Mindanao and
|
||
<collectingRegion id="2F0741A0FFE0D112FAD8FE719A721FDF" box="[1326,1389,439,465]" country="Philippines" name="Cebu" pageId="8" pageNumber="535">Cebu</collectingRegion>
|
||
. He described
|
||
<taxonomicName id="2AC3F4C1FFE0D112FEFDFE1D9EBC1FFB" authorityName="Moser. To Dr. Olaf Jaeger" authorityYear="1910" box="[267,419,475,501]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="corpulenta">
|
||
<emphasis id="DFB75350FFE0D112FEFDFE1D9EBC1FFB" box="[267,419,475,501]" italics="true" pageId="8" pageNumber="535">L. corpulenta</emphasis>
|
||
</taxonomicName>
|
||
using the following distinguishing characters that fit very well with the description of
|
||
<taxonomicName id="2AC3F4C1FFE0D112FA72FE1D9E011C17" authorityName="Brenske. Highest" authorityYear="1894" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFE0D112FA72FE1D9E011C17" italics="true" pageId="8" pageNumber="535">E. philippinica</emphasis>
|
||
</taxonomicName>
|
||
. The antennae with nine antennomeres, chestnut body color, sides of clypeus rounded, the anterior angle of the pronotum at right angle, the posterior angle of the pronotum obtuse, the scales minute and not densely punctured, the scutellum semicircular, the sides of the abdomen densely scaled and tridentate anterior tibiae.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFE0D112FF31FDAD9A851D37" blockId="8.[151,1437,151,825]" pageId="8" pageNumber="535">
|
||
The authors had the possibility to study only a digital photo of the
|
||
<typeStatus id="327831E0FFE0D112FC51FDAD9CE11C8B" box="[935,1022,619,645]" pageId="8" pageNumber="535" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="2AC3F4C1FFE0D112FBD4FDAD9BA81C8B" authorityName="Moser. To Dr. Olaf Jaeger" authorityYear="1910" box="[1058,1207,619,645]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="corpulenta">
|
||
<emphasis id="DFB75350FFE0D112FBD4FDAD9BA81C8B" box="[1058,1207,619,645]" italics="true" pageId="8" pageNumber="535">L. corpulenta</emphasis>
|
||
</taxonomicName>
|
||
deposited in ZMHB (
|
||
<figureCitation id="75F893C7FFE0D112FF69FD499E0E1CA7" box="[159,273,655,681]" captionStart="FIGURES 7–9" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[227,1378,227,1833]" captionTargetId="figure-18@5.[151,1436,181,1843]" captionTargetPageId="5" captionText="FIGURES 7–9. Lepidiota corpulenta Moser, 1910, female, syntype from ZMHB. 7, habitus, dorsal aspect; 8, habitus, ventral aspect; 9, habitus, lateral aspect (photograph of the female from ZMHB, courtesy of Dr. Johannes Frisch and Dr. Joachim Willers)." figureDoi="http://doi.org/10.5281/zenodo.4413362" httpUri="https://zenodo.org/record/4413362/files/figure.png" pageId="8" pageNumber="535">Figs. 7–9</figureCitation>
|
||
).
|
||
<taxonomicName id="2AC3F4C1FFE0D112FEDEFD499D351CA7" authorityName="Brenske. Highest" authorityYear="1894" box="[296,554,655,681]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFE0D112FEDEFD499D351CA7" box="[296,554,655,681]" italics="true" pageId="8" pageNumber="535">Exopholis philippinica</emphasis>
|
||
</taxonomicName>
|
||
was described according to female
|
||
<typeStatus id="327831E0FFE0D112FC25FD499B2A1CA7" box="[979,1077,655,681]" pageId="8" pageNumber="535" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="8952F2B3FFE0D112FBB1FD499BEC1CA4" author="Brenske, E." box="[1095,1267,655,682]" pageId="8" pageNumber="535" pagination="178 - 205" refId="ref4065" refString="Brenske, E. (1896) Neue Melolonthiden aus Africa und Asien. Stettiner Entomologische Zeitschrift, 57, 178 - 205. https: // doi. org / 10.1002 / mmnd. 48018960708" type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
). Also for this species, we studied its digital photo only (
|
||
<figureCitation id="75F893C7FFE0D112FD84FD759DC01CC0" box="[626,735,691,718]" captionStart="FIGURES 4–6" captionStartId="4.[151,264,1849,1873]" captionTargetBox="[193,1329,216,1808]" captionTargetId="figure-20@4.[151,1436,181,1825]" captionTargetPageId="4" captionText="FIGURES 4–6. Exopholis philippinica Brenske, 1894, female, holotype from SMT. 4, habitus, dorsal aspect; 5, habitus, ventral aspect; 6, habitus, lateral aspect (photograph of the female type specimen from SMT, courtesy of Dr. Olaf Jaeger)." figureDoi="http://doi.org/10.5281/zenodo.4413360" httpUri="https://zenodo.org/record/4413360/files/figure.png" pageId="8" pageNumber="535">Figs. 4–6</figureCitation>
|
||
). Comparison of photos of types of both species enabled us to propose following new synonymy. Examination of 23 non-type specimens of
|
||
<taxonomicName id="2AC3F4C1FFE0D112FBFFFD119BAF1CFF" authorityName="Brenske. Highest" authorityYear="1894" box="[1033,1200,727,753]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFE0D112FBFFFD119BAF1CFF" box="[1033,1200,727,753]" italics="true" pageId="8" pageNumber="535">E. philippinica</emphasis>
|
||
</taxonomicName>
|
||
deposited in various Philippine institutional collections (see material examined) made us possible to clarify geographic distribution of the species in the
|
||
<collectingCountry id="95D4CFD2FFE0D112FEA8FCD99EFF1D37" box="[350,480,799,825]" name="Philippines" pageId="8" pageNumber="535">Philippines</collectingCountry>
|
||
. As well as it is known,
|
||
<taxonomicName id="2AC3F4C1FFE0D112FD19FCD99C8A1D37" authorityName="Brenske. Highest" authorityYear="1894" box="[751,917,799,825]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="535" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFE0D112FD19FCD99C8A1D37" box="[751,917,799,825]" italics="true" pageId="8" pageNumber="535">E. philippinica</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the
|
||
<collectingCountry id="95D4CFD2FFE0D112FB94FCD99A351D37" box="[1122,1322,799,825]" name="Philippines" pageId="8" pageNumber="535">Philippine islands</collectingCountry>
|
||
(
|
||
<figureCitation id="75F893C7FFE0D112FACFFCD99A931D37" box="[1337,1420,799,825]" captionStart="FIGURE 13" captionStartId="7.[151,250,1721,1745]" captionTargetBox="[245,1279,181,1697]" captionTargetId="figure-18@7.[151,1436,178,1697]" captionTargetPageId="7" captionText="FIGURE 13. Distribution of examined specimens of Exopholis philippinica Brenske, 1894 in the Philippines." figureDoi="http://doi.org/10.5281/zenodo.4413368" httpUri="https://zenodo.org/record/4413368/files/figure.png" pageId="8" pageNumber="535">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |