237 lines
37 KiB
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237 lines
37 KiB
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<document id="178A82FED4015725F9FF002410F22E88" ID-CLB-Dataset="55594" ID-DOI="10.11646/zootaxa.5134.2.2" ID-GBIF-Dataset="47c35b15-3b64-4e49-8373-88d397136aeb" ID-ISSN="1175-5326" ID-Zenodo-Dep="6536905" ID-ZooBank="1D9EF14F-BFAA-45A4-B727-7CAFC55778FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652171664792" checkinUser="plazi" docAuthor="Cumming, Jeffrey M. & Brooks, Scott E." docDate="2022" docId="655241088A72FFF6FF0C24E8FA3FFE6A" docLanguage="en" docName="zootaxa.5134.2.2.pdf" docOrigin="Zootaxa 5134 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microphorella breviradia" docType="treatment" docVersion="6" lastPageNumber="199" masterDocId="996B39708A73FFF4FF9B201BFFA3FF88" masterDocTitle="Establishment of the Microphorella breviradia species group, with a key to the Nearctic species groups of Microphorella Becker (Diptera: Dolichopodidae sensu lato: Parathalassiinae)" masterLastPageNumber="214" masterPageNumber="197" pageNumber="198" updateTime="1699330120806" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="489EE2AF7F4B534B03F1AB806C890B37">Establishment of the Microphorella breviradia species group, with a key to the Nearctic species groups of Microphorella Becker (Diptera: Dolichopodidae sensu lato: Parathalassiinae)</mods:title>
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<mods:name id="AB898D30BC33393250C9919C315CBA4E" type="personal">
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<mods:namePart id="05CAB41BC8742004AF852E57FD0BC0DF">Cumming, Jeffrey M.</mods:namePart>
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<mods:nameIdentifier id="9AE23260E0724D6DCB61861C72CBCF0D" type="ORCID">0000-0002-3619-3700</mods:nameIdentifier>
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<mods:affiliation id="5EDE1273ED0D9D87070D581606196BED">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & jeff. cumming @ agr. gc. ca; https: // orcid. org / 0000 - 0002 - 3619 - 3700</mods:affiliation>
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<mods:nameIdentifier id="62196A544880FD19EEBE3530FDC3B356" type="email">jeff.cumming@agr.gc.ca</mods:nameIdentifier>
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<mods:namePart id="1EE45A65B744AB40037DFCC6B46C52B3">Brooks, Scott E.</mods:namePart>
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<mods:nameIdentifier id="2909DB4236B7647E8D4837588CD91C07" type="ORCID">0000-0001-7915-1941</mods:nameIdentifier>
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<mods:affiliation id="12ABBC26A7BD457182788C9326F125DD">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & scott. brooks @ agr. gc. ca; https: // orcid. org / 0000 - 0001 - 7915 - 1941</mods:affiliation>
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<mods:nameIdentifier id="933EDF71B2B75D12756CC5F7A7F1FCB0" type="email">scott.brooks@agr.gc.ca</mods:nameIdentifier>
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<mods:title id="BC61219E8E6B6E2337034360663EEEC8">Zootaxa</mods:title>
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<mods:date id="8D5C050A689D26ED6D6A73FA502796D9">2022</mods:date>
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<mods:number id="7B1498730BEA551CD22426B68ECCA31F">2022-05-10</mods:number>
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<mods:number id="2D2104444995BB60A342EB7C156E4DBB">5134</mods:number>
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<treatment id="655241088A72FFF6FF0C24E8FA3FFE6A" ID-DOI="http://doi.org/10.5281/zenodo.6537014" ID-GBIF-Taxon="195298249" ID-Zenodo-Dep="6537014" LSID="urn:lsid:plazi:treatment:655241088A72FFF6FF0C24E8FA3FFE6A" httpUri="http://treatment.plazi.org/id/655241088A72FFF6FF0C24E8FA3FFE6A" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198">
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<subSubSection id="A5E1A3958A72FFF5FF0C24E8FDDBFA85" box="[151,632,1267,1294]" pageId="1" pageNumber="198" type="nomenclature">
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<paragraph id="ED44F01E8A72FFF5FF0C24E8FDDBFA85" blockId="1.[151,632,1267,1294]" box="[151,632,1267,1294]" pageId="1" pageNumber="198">
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<heading id="B60C47728A72FFF5FF0C24E8FDDBFA85" bold="true" box="[151,632,1267,1294]" fontSize="11" level="1" pageId="1" pageNumber="198" reason="1">
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<emphasis id="DF8F2C0C8A72FFF5FF0C24E8FDDBFA85" bold="true" box="[151,632,1267,1294]" pageId="1" pageNumber="198">
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<taxonomicName id="2AFB8B9D8A72FFF5FF0C24E8FE6AFA86" ID-CoL="9J2P8" authority="Cumming & Brooks 2022" box="[151,457,1267,1294]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="breviradia">
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<emphasis id="DF8F2C0C8A72FFF5FF0C24E8FE6AFA86" bold="true" box="[151,457,1267,1294]" italics="true" pageId="1" pageNumber="198">Microphorella breviradia</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E1A3958A72FFF5FF0C2520FBB3F9FE" pageId="1" pageNumber="198" type="diagnosis">
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<paragraph id="ED44F01E8A72FFF5FF0C2520FBB3F9FE" blockId="1.[151,1437,1339,1977]" pageId="1" pageNumber="198">
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<emphasis id="DF8F2C0C8A72FFF5FF0C2520FEB1FADD" bold="true" box="[151,274,1339,1365]" pageId="1" pageNumber="198">Diagnosis.</emphasis>
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The
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<taxonomicName id="2AFB8B9D8A72FFF5FED22520FDC5FADD" box="[329,614,1339,1365]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="breviradia">
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<emphasis id="DF8F2C0C8A72FFF5FED22520FDC5FADD" box="[329,614,1339,1365]" italics="true" pageId="1" pageNumber="198">Microphorella breviradia</emphasis>
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</taxonomicName>
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species group can be distinguished from other parathalassiines by the following suite of characters: wing with short R
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<subScript id="717FF25B8A72FFF5FD1C2569FD2CFA08" attach="left" box="[647,655,1394,1408]" fontSize="6" pageId="1" pageNumber="198">1</subScript>
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vein that terminates at or before midpoint of wing (
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<figureCitation id="75C0EC9B8A72FFF5FB5D2544FAABFAF1" box="[1222,1288,1375,1401]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1657,1681]" captionTargetBox="[163,1436,309,1630]" captionTargetId="figure-97@6.[151,1436,309,1633]" captionTargetPageId="6" captionText="FIGURES 1–5. Microphorella breviradia sp. nov. and M. macdonaldi sp. nov. (males). 1. Habitus of M. breviradia sp. nov., left lateral view. 2. Head of M. breviradia sp. nov., anterior view (CNC487231). 3. Wing of M. breviradia sp. nov., anterior view (CNC487241). 4. Abdomen of M. breviradia sp. nov., right lateral view (CNC1155716). 5. Head of M. macdonaldi sp. nov., anterior view (holotype). Abbreviations: bm-m—basal medial crossvein; cua—anterior cubital (=anal) cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; h—humeral crossvein; M1, M2, M4—medial veins; (R)—right; R1, R2+3, R 4+5 —radial veins; Sc—subcostal vein; v epand proc—ventral epandrial process." figureDoi="http://doi.org/10.5281/zenodo.6536919" httpUri="https://zenodo.org/record/6536919/files/figure.png" pageId="1" pageNumber="198">Fig. 3</figureCitation>
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); mouthparts with sensory pit on palpus; male abdominal sternite 5 with short broad ventral projection; hypopygium (
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<figureCitation id="75C0EC9B8A72FFF5FABE2598FACEFA15" box="[1317,1389,1411,1437]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1657,1681]" captionTargetBox="[163,1436,309,1630]" captionTargetId="figure-97@6.[151,1436,309,1633]" captionTargetPageId="6" captionText="FIGURES 1–5. Microphorella breviradia sp. nov. and M. macdonaldi sp. nov. (males). 1. Habitus of M. breviradia sp. nov., left lateral view. 2. Head of M. breviradia sp. nov., anterior view (CNC487231). 3. Wing of M. breviradia sp. nov., anterior view (CNC487241). 4. Abdomen of M. breviradia sp. nov., right lateral view (CNC1155716). 5. Head of M. macdonaldi sp. nov., anterior view (holotype). Abbreviations: bm-m—basal medial crossvein; cua—anterior cubital (=anal) cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; h—humeral crossvein; M1, M2, M4—medial veins; (R)—right; R1, R2+3, R 4+5 —radial veins; Sc—subcostal vein; v epand proc—ventral epandrial process." figureDoi="http://doi.org/10.5281/zenodo.6536919" httpUri="https://zenodo.org/record/6536919/files/figure.png" pageId="1" pageNumber="198">Figs 4</figureCitation>
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,
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<figureCitation id="75C0EC9B8A72FFF5FAE22598FF7EFA49" captionStart="FIGURES 10–15" captionStartId="8.[151,264,1668,1692]" captionTargetBox="[156,1418,191,1635]" captionTargetId="figure-33@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 10–15. Hypopygium of Microphorella breviradia sp. nov. 10. Hypopygium, left lateral view (CNC DIPTERA # 192173). 11. Hypopygium, right lateral view (CNC DIPTERA # 192173). 12. Right epandrial lamella, right lateral view (CNC DIPTERA # 192167). 13. Right epandrial lamella, right lateral view (CNC487206). 14. Hypopygium and sternite 8, dorsal view (CNC DIPTERA # 192173). 15. Right cercus, dorsal view (CNC487206). Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536941" httpUri="https://zenodo.org/record/6536941/files/figure.png" pageId="1" pageNumber="198">10, 11, 14</figureCitation>
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,
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<figureCitation id="75C0EC9B8A72FFF5FF7225BCFEFBFA49" box="[233,344,1447,1473]" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1848,1872]" captionTargetBox="[154,1424,459,1815]" captionTargetId="figure-86@10.[151,1436,446,1825]" captionTargetPageId="10" captionText="FIGURES 19–21. Hypopygium of Microphorella vespera sp. nov. (CNC DIPTERA # 105212). 19. Hypopygium, left lateral view. 20. Hypopygium, right lateral view. 21. Hypopygium and sternite 8, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536947" httpUri="https://zenodo.org/record/6536947/files/figure.png" pageId="1" pageNumber="198">19, 20, 21</figureCitation>
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) with epandrium and hypandrium partially fused together, right epandrial lamella greatly enlarged and comprising most of hypopygium in right lateral view, hypandrium reduced to narrow subtriangular band positioned on left side of hypopygium; female abdomen (
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<figureCitation id="75C0EC9B8A72FFF5FD7125F4FCC9F981" box="[746,874,1519,1545]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1657,1681]" captionTargetBox="[151,1419,849,1634]" captionTargetId="figure-300@14.[151,1436,849,1634]" captionTargetPageId="14" captionText="FIGURES 31–36. Female abdomens of the Microphorella breviradia, M. chiragra and M. acroptera species groups. 31. M. breviradia sp. nov., left lateral view. 32. M. breviradia sp. nov., dorsal view. 33. Microphorella sp. 1 (M. chiragra species group), left lateral view. 34. Microphorella sp. 1 (M. chiragra species group), dorsal view. 35. Microphorella sp. 8 (M. acroptera species group), left lateral view. 36. Microphorella sp. 8 (M. acroptera species group), dorsal view. Abbreviation: tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.6536983" httpUri="https://zenodo.org/record/6536983/files/figure.png" pageId="1" pageNumber="198">Figs 31, 32</figureCitation>
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) with 7 exposed abdominal segments, telescoping terminalia retracted into segment 7, syntergite 9+10 with acanthophorous setae (
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<figureCitation id="75C0EC9B8A72FFF5FBB82608FB05F9A5" box="[1059,1190,1555,1581]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1513,1537]" captionTargetBox="[162,1424,190,1481]" captionTargetId="figure-19@9.[151,1436,181,1489]" captionTargetPageId="9" captionText="FIGURES 16–18. Female terminalia of Microphorella breviradia sp. nov. 16. Terminalia, dorsal view (CNC487214). 17. Terminalia, lateral view (CNC487214). 18. Spermatheca, lateral view (CNC487218). Abbreviations: cerc—cercus; st—sternite; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.6536945" httpUri="https://zenodo.org/record/6536945/files/figure.png" pageId="1" pageNumber="198">Figs 16, 17</figureCitation>
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), syntergite 9+10 and sternite 10 widely separated, cercus ovoid with long apical setae (
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<figureCitation id="75C0EC9B8A72FFF5FCEC262CFC6DF9D9" box="[887,974,1591,1617]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1513,1537]" captionTargetBox="[162,1424,190,1481]" captionTargetId="figure-19@9.[151,1436,181,1489]" captionTargetPageId="9" captionText="FIGURES 16–18. Female terminalia of Microphorella breviradia sp. nov. 16. Terminalia, dorsal view (CNC487214). 17. Terminalia, lateral view (CNC487214). 18. Spermatheca, lateral view (CNC487218). Abbreviations: cerc—cercus; st—sternite; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.6536945" httpUri="https://zenodo.org/record/6536945/files/figure.png" pageId="1" pageNumber="198">Figs 16</figureCitation>
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,
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<figureCitation id="75C0EC9B8A72FFF5FC40262CFC58F9D9" box="[987,1019,1591,1617]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1657,1681]" captionTargetBox="[151,1419,849,1634]" captionTargetId="figure-300@14.[151,1436,849,1634]" captionTargetPageId="14" captionText="FIGURES 31–36. Female abdomens of the Microphorella breviradia, M. chiragra and M. acroptera species groups. 31. M. breviradia sp. nov., left lateral view. 32. M. breviradia sp. nov., dorsal view. 33. Microphorella sp. 1 (M. chiragra species group), left lateral view. 34. Microphorella sp. 1 (M. chiragra species group), dorsal view. 35. Microphorella sp. 8 (M. acroptera species group), left lateral view. 36. Microphorella sp. 8 (M. acroptera species group), dorsal view. Abbreviation: tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.6536983" httpUri="https://zenodo.org/record/6536983/files/figure.png" pageId="1" pageNumber="198">32</figureCitation>
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). See
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<bibRefCitation id="896A8DEF8A72FFF5FBA5262CFAD6F9D9" author="Cumming, J. M. & Brooks, S. E." box="[1086,1397,1591,1617]" pageId="1" pageNumber="198" pagination="111 - 129" refId="ref11442" refString="Cumming, J. M. & Brooks, S. E. (2019) Phylogenetic analysis and preliminary classification of the Parathalassiinae (Empidoidea: Dolichopodidae sensu lato). Zootaxa, 4648 (1), 111 - 129. https: // doi. org / 10.11646 / zootaxa. 4648.1.5" type="journal article" year="2019">Cumming & Brooks (2019)</bibRefCitation>
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for additional parathalassiine features possessed by members of this species group.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E1A3958A72FFF5FF5C2664FF43F831" pageId="1" pageNumber="198" type="discussion">
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<paragraph id="ED44F01E8A72FFF5FF5C2664FF43F831" blockId="1.[151,1437,1339,1977]" pageId="1" pageNumber="198">
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<emphasis id="DF8F2C0C8A72FFF5FF5C2664FE98F911" bold="true" box="[199,315,1663,1689]" pageId="1" pageNumber="198">Remarks.</emphasis>
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The
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<taxonomicName id="2AFB8B9D8A72FFF5FEE92664FD2DF911" box="[370,654,1663,1689]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="breviradia">
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<emphasis id="DF8F2C0C8A72FFF5FEE92664FD2DF911" box="[370,654,1663,1689]" italics="true" pageId="1" pageNumber="198">Microphorella breviradia</emphasis>
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</taxonomicName>
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species group includes three new species from western North America (described below) that share a short R
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<subScript id="717FF25B8A72FFF5FDA326ADFDE3F94C" attach="left" box="[568,576,1718,1732]" fontSize="6" pageId="1" pageNumber="198">1</subScript>
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wing vein and a distinctively modified male hypopygium. The former feature is seen in some non-Nearctic parathalassiines (
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<emphasis id="DF8F2C0C8A72FFF5FD2926D3FD79F969" box="[690,730,1736,1761]" italics="true" pageId="1" pageNumber="198">e.g.</emphasis>
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, the
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<taxonomicName id="2AFB8B9D8A72FFF5FC8C26DCFBE1F969" authorityName="Shamshev & Grootaert" authorityYear="2004" box="[791,1090,1735,1761]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="malaysiana">
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<emphasis id="DF8F2C0C8A72FFF5FC8C26DCFBE1F969" box="[791,1090,1735,1761]" italics="true" pageId="1" pageNumber="198">Microphorella malaysiana</emphasis>
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</taxonomicName>
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species group,
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FB6326DCFA36F969" authorityName="Brooks & Cumming" authorityYear="2016" box="[1272,1429,1735,1761]" class="Insecta" family="Dolichopodidae" genus="Neothalassius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FB6326DCFA36F969" box="[1272,1429,1735,1761]" italics="true" pageId="1" pageNumber="198">Neothalassius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FF0C26F0FECBF88D" authorityName="Shamshev & Grootaert" authorityYear="2002" box="[151,360,1771,1797]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FF0C26F0FECBF88D" box="[151,360,1771,1797]" italics="true" pageId="1" pageNumber="198">Chimerothalassius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FEEE26F0FDA7F88D" authorityName="Shamshev & Grootaert" authorityYear="2005" box="[373,516,1771,1797]" class="Insecta" family="Dolichopodidae" genus="Eothalassius" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FEEE26F0FDA7F88D" box="[373,516,1771,1797]" italics="true" pageId="1" pageNumber="198">Eothalassius</emphasis>
|
||
</taxonomicName>
|
||
), but not in other known extant Nearctic species groups of
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FB0F26F0FA96F88D" authorityName="Becker" authorityYear="1909" box="[1172,1333,1771,1797]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FB0F26F0FA96F88D" box="[1172,1333,1771,1797]" italics="true" pageId="1" pageNumber="198">Microphorella</emphasis>
|
||
</taxonomicName>
|
||
. There is a fossil species of
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FEFA2714FDA1F8A1" authorityName="Becker" authorityYear="1909" box="[353,514,1807,1833]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FEFA2714FDA1F8A1" box="[353,514,1807,1833]" italics="true" pageId="1" pageNumber="198">Microphorella</emphasis>
|
||
</taxonomicName>
|
||
described from the middle Eocene Kishenehn Formation in northwestern
|
||
<collectingRegion id="2F3F3EFC8A72FFF5FAA2270BFA3EF8A2" box="[1337,1437,1808,1834]" country="United States of America" name="Montana" pageId="1" pageNumber="198">Montana</collectingRegion>
|
||
that may also belong to the
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FE56272FFDC4F8C5" box="[461,615,1843,1870]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="breviradia">
|
||
<emphasis id="DF8F2C0C8A72FFF5FE56272FFE47F8C5" box="[461,484,1844,1869]" italics="true" pageId="1" pageNumber="198">M</emphasis>
|
||
.
|
||
<emphasis id="DF8F2C0C8A72FFF5FE692728FDC4F8C5" box="[498,615,1843,1869]" italics="true" pageId="1" pageNumber="198">breviradia</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="896A8DEF8A72FFF5FC8D2728FBACF8C5" author="Greenwalt, D. E. & Amorim, D. S. & Hauser, M. & Kerr, P. H. & Fitzgerald, S. J. & Winterton, S. & Cumming, J. M. & Evenhuis, N. L. & Sinclair, B. J." box="[790,1039,1843,1870]" pageId="1" pageNumber="198" refId="ref11576" refString="Greenwalt, D. E., Amorim, D. S., Hauser, M., Kerr, P. H., Fitzgerald, S. J., Winterton, S. L, Cumming, J. M., Evenhuis, N. L. & Sinclair, B. J. (2022) Diptera of the middle Eocene Kishenehn formation. II. Palaeontologia Electronica. [accepted]" type="book" year="2022">
|
||
Greenwalt
|
||
<emphasis id="DF8F2C0C8A72FFF5FC08272FFC67F8C5" box="[915,964,1843,1869]" italics="true" pageId="1" pageNumber="198">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
). Members of the species group are generally found along fast flowing rivers and streams (
|
||
<figureCitation id="75C0EC9B8A72FFF5FD68274CFCFBF8FA" box="[755,856,1879,1906]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-19@7.[151,1436,181,1050]" captionTargetPageId="7" captionText="FIGURES 6–9. Collection localities and habitats of the Microphorella breviradia species group. 6–7. South Fork of American River (Eagle Rock), California, M. breviradia sp. nov. type locality (first author pictured in Fig. 7). 8–9. West Fork of Patrick Creek, California, M. vespera sp. nov. type locality." figureDoi="http://doi.org/10.5281/zenodo.6536929" httpUri="https://zenodo.org/record/6536929/files/figure.png" pageId="1" pageNumber="198">Figs 6–9</figureCitation>
|
||
).
|
||
<bibRefCitation id="896A8DEF8A72FFF5FCF1274CFB3EF8F9" author="Cumming, J. M. & Brooks, S. E." box="[874,1181,1879,1905]" pageId="1" pageNumber="198" pagination="111 - 129" refId="ref11442" refString="Cumming, J. M. & Brooks, S. E. (2019) Phylogenetic analysis and preliminary classification of the Parathalassiinae (Empidoidea: Dolichopodidae sensu lato). Zootaxa, 4648 (1), 111 - 129. https: // doi. org / 10.11646 / zootaxa. 4648.1.5" type="journal article" year="2019">Cumming & Brooks (2019)</bibRefCitation>
|
||
referred to this lineage as the “
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FF772760FE2EF81D" authorityName="Becker" authorityYear="1909" box="[236,397,1915,1941]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF8F2C0C8A72FFF5FF772760FE2EF81D" box="[236,397,1915,1941]" italics="true" pageId="1" pageNumber="198">Microphorella</emphasis>
|
||
</taxonomicName>
|
||
short R
|
||
<subScript id="717FF25B8A72FFF5FE7D2795FE4DF814" attach="left" box="[486,494,1934,1948]" fontSize="6" pageId="1" pageNumber="198">1</subScript>
|
||
species group” in their phylogenetic analysis of relationships within the
|
||
<taxonomicName id="2AFB8B9D8A72FFF5FABB2760FF7FF831" authorityName="Chvála" authorityYear="1981" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E1A3958A71FFF6FF0C20A0FC4FFF5D" box="[151,1004,186,213]" pageId="2" pageNumber="199" type="multiple">
|
||
<paragraph id="ED44F01E8A71FFF6FF0C20A0FC4FFF5D" blockId="2.[151,1004,186,213]" box="[151,1004,186,213]" pageId="2" pageNumber="199">
|
||
<heading id="B60C47728A71FFF6FF0C20A0FC4FFF5D" bold="true" box="[151,1004,186,213]" fontSize="11" level="1" pageId="2" pageNumber="199" reason="1">
|
||
<emphasis id="DF8F2C0C8A71FFF6FF0C20A0FC4FFF5D" bold="true" box="[151,1004,186,213]" pageId="2" pageNumber="199">
|
||
Key to males of species of the
|
||
<taxonomicName id="2AFB8B9D8A71FFF6FD9020A1FC9EFF5D" authority="Cumming & Brooks 2022" box="[523,829,186,213]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="breviradia">
|
||
<emphasis id="DF8F2C0C8A71FFF6FD9020A1FC9EFF5D" bold="true" box="[523,829,186,213]" italics="true" pageId="2" pageNumber="199">Microphorella breviradia</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E1A3958A71FFF6FF0C211BFA3FFE6A" pageId="2" pageNumber="199" type="key">
|
||
<keyStep id="560FE8BB8A71FFF6FF0C211BFA3FFEE5" pageId="2" pageNumber="199">
|
||
<paragraph id="ED44F01E8A71FFF6FF0C211BFA3FFEBA" blockId="2.[151,1437,255,483]" pageId="2" pageNumber="199">
|
||
<keyLead id="560A532B8A71FFF6FF0C211BFA3FFEBA" pageId="2" pageNumber="199">
|
||
1 Head with face broad, nearly twice width of anterior ocellus at narrowest point (
|
||
<figureCitation id="75C0EC9B8A71FFF6FC4C211BFBB2FE9D" box="[983,1041,255,278]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1657,1681]" captionTargetBox="[163,1436,309,1630]" captionTargetId="figure-97@6.[151,1436,309,1633]" captionTargetPageId="6" captionText="FIGURES 1–5. Microphorella breviradia sp. nov. and M. macdonaldi sp. nov. (males). 1. Habitus of M. breviradia sp. nov., left lateral view. 2. Head of M. breviradia sp. nov., anterior view (CNC487231). 3. Wing of M. breviradia sp. nov., anterior view (CNC487241). 4. Abdomen of M. breviradia sp. nov., right lateral view (CNC1155716). 5. Head of M. macdonaldi sp. nov., anterior view (holotype). Abbreviations: bm-m—basal medial crossvein; cua—anterior cubital (=anal) cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; h—humeral crossvein; M1, M2, M4—medial veins; (R)—right; R1, R2+3, R 4+5 —radial veins; Sc—subcostal vein; v epand proc—ventral epandrial process." figureDoi="http://doi.org/10.5281/zenodo.6536919" httpUri="https://zenodo.org/record/6536919/files/figure.png" pageId="2" pageNumber="199">Fig. 5</figureCitation>
|
||
); fore tarsomere 5 with tuft of long dorsal setae............................................................................
|
||
<emphasis id="DF8F2C0C8A71FFF6FB2F2106FA3FFEBA" bold="true" box="[1204,1436,284,306]" pageId="2" pageNumber="199">
|
||
<taxonomicName id="2AFB8B9D8A71FFF6FB2F2106FAE9FEBA" ID-CoL="9J2P7" box="[1204,1354,284,306]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="macdonaldi" status="sp. nov.">
|
||
<emphasis id="DF8F2C0C8A71FFF6FB2F2106FB6AFEBA" bold="true" box="[1204,1225,285,306]" italics="true" pageId="2" pageNumber="199">M</emphasis>
|
||
.
|
||
<emphasis id="DF8F2C0C8A71FFF6FB4D2107FAE9FEBA" bold="true" box="[1238,1354,284,306]" italics="true" pageId="2" pageNumber="199">macdonaldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BC91778A71FFF6FACB2106FA3FFEBA" box="[1360,1436,285,306]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="ED44F01E8A71FFF6FF0C2120FA3FFEE5" blockId="2.[151,1437,255,483]" pageId="2" pageNumber="199">
|
||
<keyLead id="560A532B8A71FFF6FF0C2120FA3FFEE5" pageId="2" pageNumber="199">
|
||
- Head with face narrow, about width of anterior ocellus at narrowest point (
|
||
<figureCitation id="75C0EC9B8A71FFF6FC342121FC4AFED8" box="[943,1001,314,336]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1657,1681]" captionTargetBox="[163,1436,309,1630]" captionTargetId="figure-97@6.[151,1436,309,1633]" captionTargetPageId="6" captionText="FIGURES 1–5. Microphorella breviradia sp. nov. and M. macdonaldi sp. nov. (males). 1. Habitus of M. breviradia sp. nov., left lateral view. 2. Head of M. breviradia sp. nov., anterior view (CNC487231). 3. Wing of M. breviradia sp. nov., anterior view (CNC487241). 4. Abdomen of M. breviradia sp. nov., right lateral view (CNC1155716). 5. Head of M. macdonaldi sp. nov., anterior view (holotype). Abbreviations: bm-m—basal medial crossvein; cua—anterior cubital (=anal) cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; h—humeral crossvein; M1, M2, M4—medial veins; (R)—right; R1, R2+3, R 4+5 —radial veins; Sc—subcostal vein; v epand proc—ventral epandrial process." figureDoi="http://doi.org/10.5281/zenodo.6536919" httpUri="https://zenodo.org/record/6536919/files/figure.png" pageId="2" pageNumber="199">Fig. 2</figureCitation>
|
||
); fore tarsomere 5 with shorter dorsal setae ................................................................................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="560FE8BB8A71FFF6FF0C216EFA3FFE6A" pageId="2" pageNumber="199">
|
||
<paragraph id="ED44F01E8A71FFF6FF0C216EFA3FFE2F" blockId="2.[151,1437,255,483]" pageId="2" pageNumber="199">
|
||
<keyLead id="560A532B8A71FFF6FF0C216EFA3FFE2F" pageId="2" pageNumber="199">
|
||
2 Hypopygium with basiventral projection(s) on right dorsal surstylar lobe (
|
||
<figureCitation id="75C0EC9B8A71FFF6FC28216EFBEBFE03" box="[947,1096,373,395]" captionStart="FIGURES 10–15" captionStartId="8.[151,264,1668,1692]" captionTargetBox="[156,1418,191,1635]" captionTargetId="figure-33@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 10–15. Hypopygium of Microphorella breviradia sp. nov. 10. Hypopygium, left lateral view (CNC DIPTERA # 192173). 11. Hypopygium, right lateral view (CNC DIPTERA # 192173). 12. Right epandrial lamella, right lateral view (CNC DIPTERA # 192167). 13. Right epandrial lamella, right lateral view (CNC487206). 14. Hypopygium and sternite 8, dorsal view (CNC DIPTERA # 192173). 15. Right cercus, dorsal view (CNC487206). Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536941" httpUri="https://zenodo.org/record/6536941/files/figure.png" pageId="2" pageNumber="199">Figs 11, 12, 13</figureCitation>
|
||
); phallus long and tubular, bearing very slender process medially (
|
||
<figureCitation id="75C0EC9B8A71FFF6FD962189FDF0FE20" box="[525,595,402,424]" captionStart="FIGURES 10–15" captionStartId="8.[151,264,1668,1692]" captionTargetBox="[156,1418,191,1635]" captionTargetId="figure-33@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 10–15. Hypopygium of Microphorella breviradia sp. nov. 10. Hypopygium, left lateral view (CNC DIPTERA # 192173). 11. Hypopygium, right lateral view (CNC DIPTERA # 192173). 12. Right epandrial lamella, right lateral view (CNC DIPTERA # 192167). 13. Right epandrial lamella, right lateral view (CNC487206). 14. Hypopygium and sternite 8, dorsal view (CNC DIPTERA # 192173). 15. Right cercus, dorsal view (CNC487206). Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536941" httpUri="https://zenodo.org/record/6536941/files/figure.png" pageId="2" pageNumber="199">Fig. 10</figureCitation>
|
||
).................................................
|
||
<emphasis id="DF8F2C0C8A71FFF6FB582188FA3FFE2F" bold="true" box="[1219,1436,402,424]" pageId="2" pageNumber="199">
|
||
<taxonomicName id="2AFB8B9D8A71FFF6FB582188FAEAFE20" ID-CoL="9J2P8" box="[1219,1353,402,424]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="breviradia" status="sp. nov.">
|
||
<emphasis id="DF8F2C0C8A71FFF6FB582188FB7BFE20" bold="true" box="[1219,1240,403,424]" italics="true" pageId="2" pageNumber="199">M</emphasis>
|
||
.
|
||
<emphasis id="DF8F2C0C8A71FFF6FB7F2189FAEAFE20" bold="true" box="[1252,1353,402,424]" italics="true" pageId="2" pageNumber="199">breviradia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BC91778A71FFF6FACB2189FA3FFE2F" box="[1360,1436,402,423]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="ED44F01E8A71FFF6FF0C21ABFA3FFE6A" blockId="2.[151,1437,255,483]" pageId="2" pageNumber="199">
|
||
<keyLead id="560A532B8A71FFF6FF0C21ABFA3FFE6A" pageId="2" pageNumber="199">
|
||
- Hypopygium without basiventral projections on right dorsal surstylar lobe (
|
||
<figureCitation id="75C0EC9B8A71FFF6FC2D21ABFC5EFE4E" box="[950,1021,432,454]" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1848,1872]" captionTargetBox="[154,1424,459,1815]" captionTargetId="figure-86@10.[151,1436,446,1825]" captionTargetPageId="10" captionText="FIGURES 19–21. Hypopygium of Microphorella vespera sp. nov. (CNC DIPTERA # 105212). 19. Hypopygium, left lateral view. 20. Hypopygium, right lateral view. 21. Hypopygium and sternite 8, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536947" httpUri="https://zenodo.org/record/6536947/files/figure.png" pageId="2" pageNumber="199">Fig. 20</figureCitation>
|
||
); phallus stout and bifid, bearing thick process ventrally (
|
||
<figureCitation id="75C0EC9B8A71FFF6FEE921D6FE1BFE6B" box="[370,440,461,483]" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1848,1872]" captionTargetBox="[154,1424,459,1815]" captionTargetId="figure-86@10.[151,1436,446,1825]" captionTargetPageId="10" captionText="FIGURES 19–21. Hypopygium of Microphorella vespera sp. nov. (CNC DIPTERA # 105212). 19. Hypopygium, left lateral view. 20. Hypopygium, right lateral view. 21. Hypopygium and sternite 8, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.6536947" httpUri="https://zenodo.org/record/6536947/files/figure.png" pageId="2" pageNumber="199">Fig. 19</figureCitation>
|
||
).................................................................
|
||
<emphasis id="DF8F2C0C8A71FFF6FB4521D6FA3FFE6A" bold="true" box="[1246,1436,461,482]" pageId="2" pageNumber="199">
|
||
<taxonomicName id="2AFB8B9D8A71FFF6FB4521D6FAE9FE6A" ID-CoL="9J2P9" box="[1246,1354,461,482]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="vespera" status="sp. nov.">
|
||
<emphasis id="DF8F2C0C8A71FFF6FB4521D6FB50FE6A" bold="true" box="[1246,1267,461,482]" italics="true" pageId="2" pageNumber="199">M</emphasis>
|
||
.
|
||
<emphasis id="DF8F2C0C8A71FFF6FB6421D6FAE9FE6A" bold="true" box="[1279,1354,461,482]" italics="true" pageId="2" pageNumber="199">vespera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BC91778A71FFF6FACB21D6FA3FFE6A" box="[1360,1436,461,482]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |