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<mods:title id="D0F4F52ECF9693041CFD1AAF507A068C">A comprehensive taxonomic list of brittle stars (Echinodermata: Ophiuroidea) from submarine caves of the Ryukyu Islands, southwestern Japan, with a description of a rare species, Dougaloplus echinatus (Amphiuridae)</mods:title>
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<treatment id="6551470CFFA1FF83FF04DF28E1ACFACB" ID-DOI="http://doi.org/10.5281/zenodo.5924588" ID-GBIF-Taxon="154494667" ID-Zenodo-Dep="5924588" LSID="urn:lsid:plazi:treatment:6551470CFFA1FF83FF04DF28E1ACFACB" httpUri="http://treatment.plazi.org/id/6551470CFFA1FF83FF04DF28E1ACFACB" lastPageNumber="89" pageId="16" pageNumber="89">
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<heading id="B60F4176FFA1FF83FF04DF28E7E1FCCF" bold="true" box="[151,692,798,824]" fontSize="11" level="1" pageId="16" pageNumber="89" reason="1">
<taxonomicName id="2AF88D99FFA1FF83FF04DF28E7E1FCCF" authority="(H. L. Clark, 1911)" baseAuthorityName="H. L. Clark" baseAuthorityYear="1911" box="[151,692,798,824]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiopsammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="89" phylum="Echinodermata" rank="species" species="anchista">
<emphasis id="DF8C2A08FFA1FF83FF04DF28E7E1FCCF" bold="true" box="[151,692,798,824]" pageId="16" pageNumber="89">
<emphasis id="DF8C2A08FFA1FF83FF04DF28E494FCCF" bold="true" box="[151,449,798,824]" italics="true" pageId="16" pageNumber="89">Ophiopsammus anchista</emphasis>
(
<bibRefCitation id="89698BEBFFA1FF83FE42DF28E7F8FCCF" author="Clark, H. L." box="[465,685,798,824]" pageId="16" pageNumber="89" pagination="1 - 302" refId="ref11913" refString="Clark, H. L. (1911) North Pacific Ophiurans in the collection of the United States National Museum. Smithsonian Institution United States National Museum Bulletin, 75, 1 - 302. https: // doi. org / 10.5479 / si. 03629236.75.1" type="journal article" year="1911">H. L. Clark, 1911</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="ED47F61AFFA1FF83FF04DF74E71AFCAD" blockId="16.[151,692,798,894]" box="[151,591,833,858]" pageId="16" pageNumber="89">[Japanese name: Menashi-kumohitode]</paragraph>
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<paragraph id="ED47F61AFFA1FF83FF04DF53E5AAFC89" blockId="16.[151,692,798,894]" box="[151,255,869,894]" pageId="16" pageNumber="89">
(
<figureCitation id="75C3EA9FFFA1FF83FF0CDF53E5A2FC89" box="[159,247,869,894]" captionStart="FIGURE 9" captionStartId="19.[151,250,1877,1899]" captionTargetBox="[214,1364,178,1865]" captionTargetId="figure@19.[214,1388,178,1865]" captionTargetPageId="19" captionText="FIGURE 9. Aboral views of living specimens. A, Ophiolepis superba H. L. Clark, 1915 (RUMF-ZE-02063); B, Ophiozonella cavernalis Okanishi &amp; Fujita, 2018a (RUMF-ZE-02064); C, Ophiostriatus sexradiatus irimura, 1993 (RUMF-ZE-02067); D, Ophiocoma pusilla (Brock, 1888) (RUMF-ZE-02083); E, Cryptopelta cf. callista H. L. Clark, 1938 (RUMF-ZE-02068); F, Ophiarachnella gorgonia Müller &amp; Troschel, 1842 (RUMF-ZE-02073); G, Ophiopsammus anchista (H. L. Clark, 1911) (RUMF-ZE-02079); H, Ophiarachna mauritiensis de Loriol, 1893a (RUMF-ZE-02070)." httpUri="https://zenodo.org/record/2605994/files/figure.png" pageId="16" pageNumber="89">Fig. 9G</figureCitation>
)
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<taxonomicName id="2AF88D99FFA1FF83FF04DF98E761FC34" authority="H. L. Clark 1911: 23" authorityName="H. L. Clark" authorityPageNumber="23" authorityYear="1911" box="[151,564,941,963]" class="Ophiuroidea" family="Ophiodermatidae" genus="Pectinura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="89" phylum="Echinodermata" rank="species" species="anchista">
<emphasis id="DF8C2A08FFA1FF83FF04DF98E40DFC34" box="[151,344,941,963]" italics="true" pageId="16" pageNumber="89">Pectinura anchista</emphasis>
<treatmentCitation id="6C59D00BFFA1FF83FECDDF98E761FC34" author="Clark, H. L." box="[350,564,941,963]" page="23" pageId="16" pageNumber="89" year="1911">
<bibRefCitation id="89698BEBFFA1FF83FECDDF98E75AFC34" author="Clark, H. L." box="[350,527,941,963]" pageId="16" pageNumber="89" pagination="1 - 302" refId="ref11913" refString="Clark, H. L. (1911) North Pacific Ophiurans in the collection of the United States National Museum. Smithsonian Institution United States National Museum Bulletin, 75, 1 - 302. https: // doi. org / 10.5479 / si. 03629236.75.1" type="journal article" year="1911">H. L. Clark 1911</bibRefCitation>
: 23
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, fig. 1; 1915: 303.—
<treatmentCitation id="6C59D00BFFA1FF83FC94DF9BE6BEFC34" author="Matsumoto, H." box="[775,1003,941,963]" page="322" pageId="16" pageNumber="89" year="1917">
<bibRefCitation id="89698BEBFFA1FF83FC94DF9BE6EFFC34" author="Matsumoto, H." box="[775,954,941,963]" pageId="16" pageNumber="89" pagination="1 - 408" refId="ref15066" refString="Matsumoto, H. (1917) A monograph of Japanese Ophiuroidea, arranged according to a new classification. Journal of the College of Science Imperial University Tokyo, 38, 1 - 408." type="journal article" year="1917">Matsumoto 1917</bibRefCitation>
: 322
</treatmentCitation>
.—
<treatmentCitation id="6C59D00BFFA1FF83FB9BDF9BE186FC34" author="Murakami, S." box="[1032,1235,941,963]" page="33" pageId="16" pageNumber="89" year="1942">
<bibRefCitation id="89698BEBFFA1FF83FB9BDF9BE1E4FC34" author="Murakami, S." box="[1032,1201,941,963]" pageId="16" pageNumber="89" pagination="1 - 36" refId="ref15434" refString="Murakami, S. (1942) Ophiurans of Izu, Japan. Journal of the Department of Agriculture, Kyusyu Imperial University, 7, 1 - 36." type="journal article" year="1942">Murakami 1942</bibRefCitation>
: 33
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.—
<treatmentCitation id="6C59D00BFFA1FF83FB62DF9BE0C2FC34" author="Irimura, S." box="[1265,1431,941,963]" page="5" pageId="16" pageNumber="89" year="1979">
<bibRefCitation id="89698BEBFFA1FF83FB62DF9BE028FC34" author="Irimura, S." box="[1265,1405,941,963]" pageId="16" pageNumber="89" pagination="1 - 6" refId="ref13227" refString="Irimura, S. (1979) Ophiuroidea of Sado Island, the Sea of Japan. Annual Report of the Sado Marine Biological Station, Niigata University, 9, 1 - 6." type="journal article" year="1979">Irimura 1979</bibRefCitation>
: 5
</treatmentCitation>
; 1981: 45; 1982: 64, text-fig. 38.
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</paragraph>
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<treatmentCitationGroup id="CDE8D134FFA1FF83FF04DFDCE4E4FBE8" pageId="16" pageNumber="89">
<emphasis id="DF8C2A08FFA1FF83FF04DFDCE4CDFBF7" box="[151,408,1002,1024]" italics="true" pageId="16" pageNumber="89">
<taxonomicName id="2AF88D99FFA1FF83FF04DFDCE4C1FBF7" baseAuthorityName="H. L. Clark" baseAuthorityYear="1911" box="[151,404,1002,1024]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiopsammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="89" phylum="Echinodermata" rank="species" species="anchista">Ophiopsammus anchista</taxonomicName>
.
</emphasis>
<treatmentCitation id="6C59D00BFFA1FF83FE0EDFDCE7DAFBF7" author="Vail, L. L. &amp; Rowe, F. W. E." box="[413,655,1002,1024]" page="286" pageId="16" pageNumber="89" year="1989">
<bibRefCitation id="89698BEBFFA1FF83FE0EDFDCE708FBF7" author="Vail, L. L. &amp; Rowe, F. W. E." box="[413,605,1002,1024]" pageId="16" pageNumber="89" pagination="267 - 288" refId="ref17288" refString="Vail, L. L. &amp; Rowe, F. W. E. (1989) Status of the genera Ophiopeza and Ophiopsammus (Echinodermata: Ophiuroidea) in Australian waters with the description of a new species. Proceedings of the Linnean Society of New South Wales, 110, 267 - 288." type="journal article" year="1989">Vail &amp; Rowe 1989</bibRefCitation>
: 286
</treatmentCitation>
, fig. 9A.—
<treatmentCitation id="6C59D00BFFA1FF83FD6EDFDCE162FBF7" author="Irimura, S. &amp; Tachikawa, H." box="[765,1079,1002,1024]" page="19" pageId="16" pageNumber="89" year="2002">
<bibRefCitation id="89698BEBFFA1FF83FD6EDFDCE142FBF7" author="Irimura, S. &amp; Tachikawa, H." box="[765,1047,1002,1024]" pageId="16" pageNumber="89" pagination="1 - 27" refId="ref13426" refString="Irimura, S. &amp; Tachikawa, H. (2002) Ophiuroids from the Ogasawara Islands (Echinodermata: Ophiuroidea). Ogasawara Research, 28, 1 - 27." type="journal article" year="2002">Irimura &amp; Tachikawa 2002</bibRefCitation>
: 19
</treatmentCitation>
.—Okanishi 2016: 669.—Okanishi
<emphasis id="DF8C2A08FFA1FF83FF54D83CE5A6FBE8" box="[199,243,1033,1055]" italics="true" pageId="16" pageNumber="89">et al</emphasis>
. 2016: 25, fig.
<specimenCount id="FBFE3D93FFA1FF83FE1DD83FE4E4FBE8" box="[398,433,1033,1055]" pageId="16" pageNumber="89" type="generic">9L.</specimenCount>
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</paragraph>
</subSubSection>
<subSubSection id="A5E2A591FFA1FF83FF04D87AE4D5FB27" pageId="16" pageNumber="89" type="materials_examined">
<paragraph id="ED47F61AFFA1FF83FF04D87AE4D5FB27" blockId="16.[151,1436,1100,1341]" pageId="16" pageNumber="89">
<emphasis id="DF8C2A08FFA1FF83FF04D87AE4C4FB92" bold="true" box="[151,401,1100,1125]" pageId="16" pageNumber="89">Material examined.</emphasis>
Okinawa Island (RUMF-ZE-02064[1], RUMF-ZE-02077[1], RUMF-ZE-02078[1]): “entrance” of the “Hedo Dome” cave, under rocks or coral rubble, approximately
<quantity id="2A005BFFFFA1FF83FBADD859E12CFB70" box="[1086,1145,1135,1160]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="89" unit="m" value="15.0">15 m</quantity>
depth. Ie
<collectingCountry id="95EFB68AFFA1FF83FB7ED859E064FB7F" box="[1261,1329,1135,1160]" name="Iceland" pageId="16" pageNumber="89">Island</collectingCountry>
(RUMF- ZE-02079[1], RUMF-ZE-02076[1]): “entrance” of the “Unnamed cave”, under coral rubble, approximately
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depth,
<date id="9946D0DAFFA1FF83FF70D88EE429FB27" box="[227,380,1208,1232]" pageId="16" pageNumber="89" value="2017-06-24">24 June 2017</date>
.
</paragraph>
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<subSubSection id="A5E2A591FFA1FF83FF54D8EAE659FB03" box="[199,780,1244,1269]" pageId="16" pageNumber="89" type="distribution">
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<emphasis id="DF8C2A08FFA1FF83FF54D8EAE435FB02" bold="true" box="[199,352,1244,1269]" pageId="16" pageNumber="89">Distribution.</emphasis>
Around
<collectingCountry id="95EFB68AFFA1FF83FE57D8EBE752FB03" box="[452,519,1245,1268]" name="Japan" pageId="16" pageNumber="89">Japan</collectingCountry>
. Depth range
<quantity id="2A005BFFFFA1FF83FD31D8EAE65DFB03" box="[674,776,1244,1269]" metricMagnitude="2" metricUnit="m" metricValue="2.815" metricValueMax="5.58" metricValueMin="0.05" pageId="16" pageNumber="89" unit="m" value="281.5" valueMax="558.0" valueMin="5.0">
5
<specimenCount id="FBFE3D93FFA1FF83FD2ED8EAE65DFB03" box="[701,776,1244,1269]" pageId="16" pageNumber="89" type="generic">558 m</specimenCount>
</quantity>
.
</paragraph>
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<subSubSection id="A5E2A591FFA1FF83FF54D8C9E1ACFACB" pageId="16" pageNumber="89" type="discussion">
<paragraph id="ED47F61AFFA1FF83FF54D8C9E1ACFACB" blockId="16.[151,1436,1100,1341]" pageId="16" pageNumber="89">
<emphasis id="DF8C2A08FFA1FF83FF54D8C9E46EFAEF" bold="true" box="[199,315,1279,1304]" pageId="16" pageNumber="89">Remarks.</emphasis>
The examined specimens fall within
<taxonomicName id="2AF88D99FFA1FF83FD7AD936E6EAFAEF" authorityName="H. L. Clark" authorityYear="1911" box="[745,959,1280,1304]" class="Ophiuroidea" family="Ophiodermatidae" genus="Pectinura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="89" phylum="Echinodermata" rank="species" species="anchista">
<emphasis id="DF8C2A08FFA1FF83FD7AD936E6EAFAEF" box="[745,959,1280,1304]" italics="true" pageId="16" pageNumber="89">Pectinura anchista</emphasis>
</taxonomicName>
by virtue of having wider, approximately two to three times than long, dorsal arm plates on proximal portion of the arm, and cylindrical arms.
</paragraph>
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</treatment>
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