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<document id="DEDF90A222B81B39D24A340A9B76AEDC" ID-CLB-Dataset="299409" ID-DOI="10.5281/zenodo.11067293" ID-GBIF-Dataset="9235db45-04c4-4570-8987-cb84765763a4" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie &amp; Liang, Hongbin" docDate="2023" docId="644DE316FF9EFFADD34FDDFBFD386134" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Tetragonoderus parviculus Kavanaugh and Cueva-Dabkoski 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="471" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="468" updateTime="1719954462017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="64127C4977DD99EDDE156E58CC61DB7F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E7847859A5CF872CD8704502D5EC9D68">
<mods:title id="A5FE200E2338CAFA134A44488332745D">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
</mods:titleInfo>
<mods:name id="0C7F0DAF3A28C7A81DDEE829975FD191" type="personal">
<mods:role id="7C1B71F9D52395687BB7999AA4C162FF">
<mods:roleTerm id="C85B4A4B4DFC23DD75639E87FD3D99E5">Author</mods:roleTerm>
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<mods:namePart id="CCA63D18DB25C014E1EA01D81013D182">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="9EA5F78BA655466B8D24E214DB092A2E">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="52E29B554758296DCCC847FE819E1305">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0ED7FAC662408F1528C782B9C364FF41">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="8914056A894575969A8F52DF44FF6B5D">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="6812A8FBB4F55B9932B1C96D811CB2A1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="538C3B34B354F92C1571845879915AC1">Liang, Hongbin</mods:namePart>
<mods:affiliation id="ADC66B4230D8195AA13A4B8EF491FC18">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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<mods:title id="03EE46A46F31056EDDB0EA1DA1361203">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="BF1626940D9F36E06D52B00F5762C7E0">
<mods:date id="6F4845BF499C70EC54409282DFD55AB5">2023</mods:date>
<mods:detail id="626B72C7F037EE45A32A6D5B8CBAB02E" type="pubDate">
<mods:number id="CECD346EE9E4136864714626A0B1C00D">2023-02-15</mods:number>
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<mods:number id="2BAE4AC8CF80EC8C5C1BF0E1A74789FB">67</mods:number>
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<treatment id="644DE316FF9EFFADD34FDDFBFD386134" ID-DOI="http://doi.org/10.5281/zenodo.12628250" ID-GBIF-Taxon="234412274" ID-Zenodo-Dep="12628250" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134" httpUri="http://treatment.plazi.org/id/644DE316FF9EFFADD34FDDFBFD386134" lastPageId="21" lastPageNumber="471" pageId="17" pageNumber="468">
<subSubSection id="A4FE018BFF9EFFA9D34FDDFBFC436594" box="[123,905,1436,1460]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="EC5B5200FF9EFFA9D34FDDFBFC436594" blockId="17.[123,905,1436,1490]" box="[123,905,1436,1460]" pageId="17" pageNumber="468">
<heading id="B713E56CFF9EFFA9D34FDDFBFC436594" box="[123,905,1436,1460]" fontSize="10" level="2" pageId="17" pageNumber="468" reason="4">
<emphasis id="DE908E12FF9EFFA9D34FDDFBFC436594" bold="true" box="[123,905,1436,1460]" pageId="17" pageNumber="468">
<taxonomicName id="2BE42983FF9EFFA9D34FDDFBFCEC6594" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[123,806,1436,1460]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9EFFA9D34FDDFBFE686593" bold="true" box="[123,418,1436,1460]" italics="true" pageId="17" pageNumber="468">Tetragonoderus parviculus</emphasis>
Kavanaugh and Cueva-Dabkoski
</taxonomicName>
,
<taxonomicNameLabel id="C5A33369FF9EFFA9D006DDFBFC436594" box="[818,905,1436,1459]" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9EFFA9D34FDDDBFD7F65F5" box="[123,693,1468,1490]" pageId="17" pageNumber="468" type="description">
<paragraph id="EC5B5200FF9EFFA9D34FDDDBFD7F65F5" blockId="17.[123,905,1436,1490]" box="[123,693,1468,1490]" pageId="17" pageNumber="468">
<figureCitation id="74DF4E85FF9EFFA9D34FDDDBFF2865F5" box="[123,226,1468,1490]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="17" pageNumber="468">Figures 11</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D3D9DDDBFED265F5" box="[237,280,1468,1490]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="17" pageNumber="468">13D</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D217DDDBFE8465F5" box="[291,334,1468,1490]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="17" pageNumber="468">14D</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D26DDDDBFE4E65F5" box="[345,388,1468,1490]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="17" pageNumber="468">15D</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D2BBDDDBFE7065F5" box="[399,442,1468,1490]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="17" pageNumber="468">16D</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D2F1DDDBFE3A65F5" box="[453,496,1468,1490]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="17" pageNumber="468">17D</figureCitation>
,
<figureCitation id="74DF4E85FF9EFFA9D2CFDDDBFDD865F5" box="[507,530,1468,1490]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="17" pageNumber="468">18</figureCitation>
C-D, 21, and 23.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9EFFAAD39FDD81FE2D6193" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="EC5B5200FF9EFFAAD39FDD81FD176153" blockId="17.[123,1174,1509,1725]" lastBlockId="18.[123,1174,219,1725]" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<emphasis id="DE908E12FF9EFFA9D39FDD81FE9365DA" bold="true" box="[171,345,1510,1533]" pageId="17" pageNumber="468">Type material.</emphasis>
<materialsCitation id="5C8C585DFF9EFFA9D25CDD82FCFD665A" ID-GBIF-Occurrence="4902827307" collectingDate="2006-06-03" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; R. L. Brett &amp; D. Z. Dong" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039396" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="335FECA2FF9EFFA9D25CDD82FE3265DA" box="[360,504,1509,1533]" pageId="17" pageNumber="468" type="holotype">HOLOTYPE</typeStatus>
, a
<specimenCount id="FAE29989FF9EFFA9D113DD82FD9765DA" box="[551,605,1509,1533]" pageId="17" pageNumber="468" type="male">male</specimenCount>
, deposited in
<collectionCode id="8AF5CAC5FF9EFFA9D03FDD82FCF065DA" box="[779,826,1509,1533]" pageId="17" pageNumber="468">IOZ</collectionCode>
, labeled: “
<specimenCode id="BC42FA7BFF9EFFA9D08FDD82FB4E65DA" box="[955,1156,1509,1533]" pageId="17" pageNumber="468">CASENT1039396</specimenCode>
”/ “
<collectingCountry id="94F31290FF9EFFA9D3B0DE62FF17663A" box="[132,221,1541,1565]" name="China" pageId="17" pageNumber="468">CHINA</collectingCountry>
,
<collectingRegion id="2E209CE2FF9EFFA9D3DFDE61FEF5663A" box="[235,319,1542,1565]" country="China" name="Yunnan" pageId="17" pageNumber="468">Yunnan</collectingRegion>
,
<collectingCounty id="053A2A8CFF9EFFA9D27ADE62FDD5663A" box="[334,543,1541,1565]" pageId="17" pageNumber="468">Tengchong County</collectingCounty>
,
<collectingMunicipality id="0C3FC87AFF9EFFA9D119DE62FD28663A" box="[557,738,1541,1565]" pageId="17" pageNumber="468">Wuhe Township</collectingMunicipality>
,
<location id="E93B04DBFF9EFFA9D1C4DE62FC61663A" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9EFFA9D1C4DE62FC61663A" box="[752,939,1541,1565]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan Jiang" pageId="17" pageNumber="468" stateProvince="Yunnan">Longchuan Jiang</location>
just below bridge at Menglian village, 24.89176°/98.67551°,”/ “
<quantity id="2B1CFFE5FF9EFFA9D166DE42FD62661A" box="[594,680,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">
<elevation id="67C9B533FF9EFFA9D166DE42FD62661A" box="[594,680,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">1230 m</elevation>
</quantity>
,
<date id="985A74C0FF9EFFA9D181DE42FCF1661A" box="[693,827,1573,1597]" pageId="17" pageNumber="468" value="2006-06-03">
<collectingDate id="881E8D28FF9EFFA9D181DE42FCF1661A" box="[693,827,1573,1597]" pageId="17" pageNumber="468" value="2006-06-03">3 June 2006</collectingDate>
</date>
, Stop # DHK-2006-054A,
<collectorName id="411137D6FF9EFFA9D756DE41FF3D667A" pageId="17" pageNumber="468">D.H. Kavanaugh</collectorName>
,
<collectorName id="411137D6FF9EFFA9D231DE21FEBF667A" box="[261,373,1606,1629]" pageId="17" pageNumber="468">R.L. Brett</collectorName>
&amp;
<collectorName id="411137D6FF9EFFA9D2ABDE21FDDF6679" box="[415,533,1606,1630]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
collectors”/ “HOLOTYPE
<taxonomicName id="2BE42983FF9EFFA9D07DDE22FBA7667A" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[841,1133,1605,1629]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9EFFA9D07DDE22FBA7667A" box="[841,1133,1605,1629]" italics="true" pageId="17" pageNumber="468">Tetragonoderus parviculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF9EFFA9D743DE21FF62665A" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label].
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D00BDE02FCB866BA" ID-GBIF-Occurrence="4902827318" collectingDate="2006-06-03" collectionCode="CAS, IOZ" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
A total of four paratypes:
<specimenCount id="FAE29989FF9EFFA9D754DE02FF7666BA" count="3" pageId="17" pageNumber="468" type="male">three males</specimenCount>
, in
<collectionCode id="8AF5CAC5FF9EFFA9D3D5DEE2FEDF66BA" box="[225,277,1669,1693]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
and
<collectionCode id="8AF5CAC5FF9EFFA9D27DDEE2FEB266BA" box="[329,376,1669,1693]" pageId="17" pageNumber="468">IOZ</collectionCode>
, same label data as holotype, except first label [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D046DEE2FE78669A" ID-GBIF-Occurrence="4902827332" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039394" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
unique identifier label] different, “
<specimenCode id="BC42FA7BFF9EFFA9D3EDDEC2FE6F669A" box="[217,421,1701,1725]" pageId="17" pageNumber="468">CASENT1039394</specimenCode>
”,
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D2F6DEC2FD1F669A" ID-GBIF-Occurrence="4902827308" box="[450,725,1701,1725]" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039395" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
“
<specimenCode id="BC42FA7BFF9EFFA9D2F8DEC2FD5F669A" box="[460,661,1701,1725]" pageId="17" pageNumber="468">CASENT1039395</specimenCode>
” and
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D1D0DEC2FB95669A" ID-GBIF-Occurrence="4902827329" box="[740,1119,1701,1725]" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039397" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
“
<specimenCode id="BC42FA7BFF9EFFA9D1DADEC2FC70669A" box="[750,954,1701,1725]" pageId="17" pageNumber="468">CASENT1039397</specimenCode>
”, respectively;
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFAAD75BDEC1FD136153" ID-GBIF-Occurrence="4902827335" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2016-06-25" collectingMethod="baited pitfall trap" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. - B. Wang" country="China" county="Tengchong County" elevation="1290" lastPageId="18" lastPageNumber="469" latitude="24.613" location="YUNNAN Prov." longLatPrecision="12" longitude="97.75633" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="1" specimenCount-female="1" specimenCount-male="3" stateProvince="YUNNAN Prov." typeStatus="holotype">
<specimenCount id="FAE29989FF9EFFA9D75BDEC1FB5C669A" box="[1135,1174,1702,1725]" count="1" pageId="17" pageNumber="468" type="female">one</specimenCount>
female, in
<collectionCode id="8AF5CAC5FF9DFFAAD3CFD8BCFE8360D4" box="[251,329,219,243]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="18" pageNumber="469">NMPC</collectionCode>
, labeled: “
<collectingCountry id="94F31290FF9DFFAAD2F3D8BCFDE860D4" box="[455,546,219,243]" name="China" pageId="18" pageNumber="469">CHINA</collectingCountry>
:
<collectingRegion id="2E209CE2FF9DFFAAD106D8BCFD2160D5" box="[562,747,219,243]" country="China" name="Yunnan" pageId="18" pageNumber="469">YUNNAN Prov.</collectingRegion>
<locationDeviation id="FDE94B22FF9DFFAAD1CDD8BCFC3560D4" box="[761,1023,219,243]" pageId="18" pageNumber="469">
<quantity id="2B1CFFE5FF9DFFAAD1CDD8BCFC8060D4" box="[761,842,219,243]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="18" pageNumber="469" unit="km" value="5.9">5.9 km</quantity>
W Tongbiguan
</locationDeviation>
,
<geoCoordinate id="89D034C7FF9DFFAAD725D8BCFB5B60D4" box="[1041,1169,219,243]" degrees="24" direction="north" minutes="36.78" orientation="latitude" pageId="18" pageNumber="469" precision="9" value="24.613">24°36.78’N</geoCoordinate>
,
<geoCoordinate id="89D034C7FF9DFFAAD34FD89CFF326134" box="[123,248,251,275]" degrees="97" direction="east" minutes="45.38" orientation="longitude" pageId="18" pageNumber="469" precision="9" value="97.75633">97°45.38’E</geoCoordinate>
,
<collectorName id="411137D6FF9DFFAAD236D89BFE966134" box="[258,348,251,275]" pageId="18" pageNumber="469">J. Hájek</collectorName>
,
<collectorName id="411137D6FF9DFFAAD251D89BFE1D6134" box="[357,471,251,275]" pageId="18" pageNumber="469">J. Růžička</collectorName>
, &amp;
<collectorName id="411137D6FF9DFFAAD2C8D89CFDB36133" box="[508,633,251,276]" pageId="18" pageNumber="469">C.-B. Wang</collectorName>
leg.”/ “(CH12)
<date id="985A74C0FF9DFFAAD017D89CFC086134" box="[803,962,251,275]" pageId="18" pageNumber="469" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">25.-27.vi.2016</date>
;
<quantity id="2B1CFFE5FF9DFFAAD0F8D89CFBD56134" box="[972,1055,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="469" unit="m" value="1290.0">
<elevation id="67C9B533FF9DFFAAD0F8D89CFBD56134" box="[972,1055,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="469" unit="m" value="1290.0">1290 m</elevation>
</quantity>
,
<collectingMethod id="35A52A17FF9DFFAAD71DD89CFF046114" pageId="18" pageNumber="469">baited pitfall trap</collectingMethod>
#06 (fish meat, ripening cheese), secondary broadleaved forest, valley near stream”. Each paratype also bears the following label: “PARATYPE
<taxonomicName id="2BE42983FF9DFFAAD1E1D95CFC326174" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[725,1016,315,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9DFFAAD1E1D95CFC326174" box="[725,1016,315,339]" italics="true" pageId="18" pageNumber="469">Tetragonoderus parviculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF9DFFAAD736D95BFB926174" box="[1026,1112,316,339]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FD91AFE2D6193" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
<emphasis id="DE908E12FF9DFFAAD39FD91AFE8C61B3" bold="true" box="[171,326,381,404]" pageId="18" pageNumber="469">
<typeStatus id="335FECA2FF9DFFAAD39FD91AFF2961B3" box="[171,227,381,404]" pageId="18" pageNumber="469">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="94F31290FF9DFFAAD265D91BFE5961B3" box="[337,403,380,404]" name="China" pageId="18" pageNumber="469">China</collectingCountry>
,
<collectingRegion id="2E209CE2FF9DFFAAD295D91AFE3F61B3" box="[417,501,381,404]" country="China" name="Yunnan" pageId="18" pageNumber="469">Yunnan</collectingRegion>
,
<collectingCounty id="053A2A8CFF9DFFAAD137D91BFD1E61B3" box="[515,724,380,404]" pageId="18" pageNumber="469">Tengchong County</collectingCounty>
,
<collectingMunicipality id="0C3FC87AFF9DFFAAD1D6D91BFC5D61B3" box="[738,919,380,404]" pageId="18" pageNumber="469">Wuhe Township</collectingMunicipality>
,
<location id="E93B04DBFF9DFFAAD092D91BFBAE61B3" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9DFFAAD092D91BFBAE61B3" box="[934,1124,380,404]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan River" pageId="18" pageNumber="469" stateProvince="Yunnan">Longchuan River</location>
just below bridge at
<location id="E93B04DBFF9DFFAAD21DD9FBFE296193" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9DFFAAD21DD9FBFE296193" box="[297,483,412,436]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Menglian village" pageId="18" pageNumber="469" stateProvince="Yunnan">Menglian village</location>
.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9DFFAAD39FD9DAFC1E6232" pageId="18" pageNumber="469" type="etymology">
<paragraph id="EC5B5200FF9DFFAAD39FD9DAFC1E6232" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
<emphasis id="DE908E12FF9DFFAAD39FD9DAFEE161F3" bold="true" box="[171,299,445,468]" pageId="18" pageNumber="469">Etymology.</emphasis>
The species epithet,
<taxonomicName id="2BE42983FF9DFFAAD126D9DBFD4961F3" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[530,643,444,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF9DFFAAD126D9DBFD4961F3" box="[530,643,444,468]" italics="true" pageId="18" pageNumber="469">parviculus</emphasis>
</taxonomicName>
, is a combination of the Latin adjective,
<emphasis id="DE908E12FF9DFFAAD772D9DAFB4561F3" box="[1094,1167,445,468]" italics="true" pageId="18" pageNumber="469">parvus</emphasis>
, meaning small, and the Latin qualifier, -ic
<emphasis id="DE908E12FF9DFFAAD10CD9BBFDAE61D3" box="[568,612,476,500]" italics="true" pageId="18" pageNumber="469">ulus</emphasis>
, meaning somewhat, here used as an adjective in the masculine form. It refers to the relatively small size of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9DFFAAD39FDA79FE9A63FF" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="EC5B5200FF9DFFAAD39FDA79FDA46350" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
<emphasis id="DE908E12FF9DFFAAD39FDA79FED56212" bold="true" box="[171,287,542,565]" pageId="18" pageNumber="469">Diagnosis.</emphasis>
Adults of
<taxonomicName id="2BE42983FF9DFFAAD2AADA79FDE46212" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[414,558,541,565]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF9DFFAAD2AADA79FDE46212" box="[414,558,541,565]" italics="true" pageId="18" pageNumber="469">T. parviculus</emphasis>
</taxonomicName>
can be distinguished from those of other
<emphasis id="DE908E12FF9DFFAAD726DA7AFB5C6212" box="[1042,1174,541,565]" italics="true" pageId="18" pageNumber="469">cyclosomine</emphasis>
species in the study area by the following combination of character states: Body size slightly small for genus, BL males =
<quantity id="2B1CFFE5FF9DFFAAD25ADA3AFDCF6252" box="[366,517,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="18" pageNumber="469" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7 to 4.2 mm</quantity>
, female =
<quantity id="2B1CFFE5FF9DFFAAD141DA3AFD006252" box="[629,714,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="469" unit="mm" value="4.4">4.4 mm</quantity>
; elytra dark without pale markings in basal half, with distinct transverse subapical band of pale spots extended from interval 4 to 8 (
<figureCitation id="74DF4E85FF9DFFAAD713DA19FB4262B1" box="[1063,1160,638,662]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="18" pageNumber="469">Fig. 11A</figureCitation>
), faintly present also on interval 2 and/or
<quantity id="2B1CFFE5FF9DFFAAD11EDAF9FD9E6291" box="[554,596,670,694]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="469" unit="in" value="3.0">3 in</quantity>
some individuals, femora dark black or piceous; dorsal surfaces shiny, with distinct aeneous-greenish metallic reflection, of similar sheen throughout; elytral microsculpture comprised of more or less regularly transverse meshes, except their orientation slightly distorted in or near discal setal pore punctures; pronotum widest anterior to middle; elytra slightly to distinctly and obliquely truncate apically; posterior discal setiferous puncture on interval 3 inserted more posteriad, at level of pale transverse subapical band; front tarsomeres 1 to 3 without lateral expansions (
<figureCitation id="74DF4E85FF9DFFAAD28CDB38FDEF6350" box="[440,549,863,887]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="18" pageNumber="469">Figs. 14D</figureCitation>
,
<figureCitation id="74DF4E85FF9DFFAAD105DB38FDA86350" box="[561,610,863,887]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="18" pageNumber="469">15D</figureCitation>
).
</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FDBE7FE9A63FF" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
In the study area, specimens of
<taxonomicName id="2BE42983FF9DFFAAD2C9DBE7FD4363B0" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[509,649,895,919]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF9DFFAAD2C9DBE7FD4363B0" box="[509,649,895,919]" italics="true" pageId="18" pageNumber="469">T. parviculus</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF9DFFAAD1A3DB18FD3263B0" box="[663,760,895,919]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="18" pageNumber="469">Fig. 12A</figureCitation>
) might be confused only with those of
<taxonomicName id="2BE42983FF9DFFAAD34FDBC6FF38639F" box="[123,242,929,952]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF9DFFAAD34FDBC6FF38639F" box="[123,242,929,952]" italics="true" pageId="18" pageNumber="469">T. arcuatus</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF9DFFAAD237DBF8FEAE6390" box="[259,356,927,951]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="18" pageNumber="469">Figs. 9A</figureCitation>
). Refer to the key and Diagnosis section for
<taxonomicName id="2BE42983FF9DFFAAD079DBC6FC0F639F" box="[845,965,929,952]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF9DFFAAD079DBC6FC0F639F" box="[845,965,929,952]" italics="true" pageId="18" pageNumber="469">T. arcuatus</emphasis>
</taxonomicName>
for features distinguishing these taxa.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9DFFADD39FDB86FD386134" lastPageId="21" lastPageNumber="472" pageId="18" pageNumber="469" type="description">
<paragraph id="EC5B5200FF9DFFAAD39FDB86FBD063DF" blockId="18.[123,1174,219,1725]" box="[171,1050,992,1017]" pageId="18" pageNumber="469">
<emphasis id="DE908E12FF9DFFAAD39FDB86FEFC63DF" bold="true" box="[171,310,993,1016]" pageId="18" pageNumber="469">Description.</emphasis>
Size. Small for genus, BL males =
<quantity id="2B1CFFE5FF9DFFAAD181DB87FC8463DF" box="[693,846,992,1016]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="18" pageNumber="469" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7 to 4.2 mm</quantity>
, female =
<quantity id="2B1CFFE5FF9DFFAAD0F4DB87FBDC63DF" box="[960,1046,992,1016]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="469" unit="mm" value="4.4">4.4 mm</quantity>
.
</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FDC67FEAC64DD" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
Color. (
<figureCitation id="74DF4E85FF9DFFAAD3C8DC67FE93643F" box="[252,345,1024,1048]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="18" pageNumber="469">Fig 11A</figureCitation>
). Head, including venter, piceous to black except mandibles paler subapically and antennae with antennomeres 1, 1 to 2, or 1 to 3 paler. Pronotum piceous to black. Elytra piceous to black but with transverse subapical band of pale spots at about five-sixths elytral length, extended from interval 2, 3, or 4 to interval 8, with spots on intervals 2 and 3 less distinct in most specimens, those on intervals 2 to 4 shorter, and spots on intervals 5 to 8 longer, extended more anteriad, and more distinct. Venter piceous laterally, rufo-piceous medially, elytral epipleurae paler rufopiceous. Legs, piceous to rufopiceous, with femora darker than tibiae or tarsi or tibiae paler than femora and tarsi.
</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FDD65FC5465DC" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">Reflection, luster, and microsculpture. Dorsum of head, pronotum and elytra with distinct aeneous-greenish metallic reflection. Head and pronotum slightly duller than elytra, with microsculpture moderately impressed and comprised of isodiametric meshes; elytra shiny, with microscuplture slightly less deeply impressed and comprised of moderately to markedly transverse meshes, meshes slightly distorted only in immediate vicinity of discal setae; abdominal venter dull laterally and shiny and slightly iridescent medially, with microsculpture moderately impressed and comprised of isodiametric to slightly transverse meshes laterally, more shallowly impressed and formed of transverse microlines and markedly transverse meshes medially.</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FDE63FCD5667B" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">Head. Eyes large, hemispheric; antennae slightly short, extended only to basal one-fifth of elytra; clypeus bisetose; labrum with three pairs of setae; mentum asetose, with a broadly rounded medial tooth; submentum anteriorly with a single pair of setae.</paragraph>
<paragraph id="EC5B5200FF9DFFACD39FDE03FC1A61F3" blockId="18.[123,1174,219,1725]" lastBlockId="20.[123,1174,219,1725]" lastPageId="20" lastPageNumber="471" pageId="18" pageNumber="469">
Prothorax. Pronotum (
<figureCitation id="74DF4E85FF9DFFAAD2AADE03FDCB665B" box="[414,513,1636,1660]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="18" pageNumber="469">Fig. 13D</figureCitation>
) slightly narrow (ratio PWM/EW = 0.61 to 0.65), distinctly narrowed basally (ratio PWB/PWA = 1.04 to 1.09), widest distinctly anterior to mid-length, with anterior angles very short and broadly rounded; posterior angles slightly obtuse and slightly projected laterally; apical margin straight or very slightly concave between anterior angles; lateral margins arcuate and with a short and shallow sinuation just anterior to posterior angles; basal margin faintly bisinuate, not or only slightly lobate medially; anterior margination distinct laterally, more or less effaced medially; lateral margination distinct and complete, joined with apical and basal margination at anterior and posterior angles, respectively; basal margination distinct laterally, less distinct medially; midlateral setae inserted on lateral margination (“lateral bead”) at about one-fourth pronotal length; basolateral setae inserted on margination at posterior angles. Prosternal intercoxal process short, rounded, with complete but faint margination apically.
</paragraph>
<caption id="B89B0288FF9CFFABD3AFDE5AFE3766BB" ID-DOI="http://doi.org/10.5281/zenodo.11067309" ID-Zenodo-Dep="11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="19" pageNumber="470" startId="19.[155,167,1597,1616]" targetBox="[123,1176,216,1583]" targetPageId="19" targetType="figure">
<paragraph id="EC5B5200FF9CFFABD3AFDE5AFE3766BB" blockId="19.[123,1174,1596,1692]" pageId="19" pageNumber="470">
<smallCapsWord id="EABDC4DCFF9CFFABD3AFDE5AFF126668" baselines="1611,1611" box="[155,216,1597,1616]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="19" pageNumber="470">FIGURE</smallCapsWord>
11.
<taxonomicName id="2BE42983FF9CFFABD3C9DE5BFD3D6668" authority="Kavanaugh &amp; Cueva-Dabkoski" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[253,759,1596,1615]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9CFFABD3C9DE5BFE296668" box="[253,483,1596,1615]" italics="true" pageId="19" pageNumber="470">Tetragonoderus parviculus</emphasis>
Kavanaugh &amp; Cueva-Dabkoski
</taxonomicName>
,
<taxonomicNameLabel id="C5A33369FF9CFFABD1CBDE5AFC886677" box="[767,834,1597,1616]" pageId="19" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<paragraph id="EC5B5200FF9BFFACD39FD9BBFC0F6292" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">Elytra. Broadly ovate, longer than wide (ratio EL/EW = 1.27 to 1.37), widest slightly posterior to mid-length, obliquely truncate apically, humeri obtusely rounded; elytral striae deeply impressed throughout, elytral intervals flat or slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 at junction of striae 1 and 2; two discal setiferous pores present on interval three, the anterior seta inserted adjacent to stria 3 near elytral midlength, the other inserted adjacent to stria 2 near apical one-fourth at level of subapical pale band; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 or 12 setae.</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDAD8FBAE6311" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
Legs. Tarsal claws finely denticulate medially. Males (
<figureCitation id="74DF4E85FF9BFFACD1C0DAD9FCAB62F1" box="[756,865,702,726]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="20" pageNumber="471">Figs. 14D</figureCitation>
,
<figureCitation id="74DF4E85FF9BFFACD059DAD9FC5762F1" box="[877,925,702,726]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="20" pageNumber="471">16D</figureCitation>
) with front and middle tarsi with tarsomeres 1 to 4 distinctly broader than in females; male front tarsomeres 1 to 3 with two rows of adhesive squamosetae ventrally (
<figureCitation id="74DF4E85FF9BFFACD15EDA99FD046331" box="[618,718,766,790]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="20" pageNumber="471">Fig. 15D</figureCitation>
), females without such setae; middle tarsi with tarsomeres 1 to 4 (
<figureCitation id="74DF4E85FF9BFFACD24EDB79FE116311" box="[378,475,798,822]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="20" pageNumber="471">Fig. 17D</figureCitation>
) with pads of adhesive setae ventrally, absent from females.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDB58FBC46390" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
Male genitalia. Median lobe (
<figureCitation id="74DF4E85FF9BFFACD2C4DB58FDA16370" box="[496,619,831,855]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="20" pageNumber="471">Fig. 18C,D</figureCitation>
) long and slender, shaft smoothly arcuate dorsally and ventrally, gradually tapered toward apex, with apical lamella slightly sinuate and tapered to a rounded point apically in lateral view, shaft symmetrically tapered and with apical lamella slightly expanded and apically rounded in dorsal view. Internal sac without evident spines.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDBA7FBAF645E" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
Geographical variation. The female specimen from Tongbiquan Township differs from the male specimens from Wuhe Township in having antennae with only antennomere 1 pale (antennomeres 1 to 2 or 3 pale in the other specimens), the tarsi darker than the tibiae (both tibiae and tarsi paler than the femur in the other specimens), and the elytral subapical pale band evident from interval 2 to interval 8 (pale band visible mainly from interval 4 to interval 8 and only very faintly evident on interval
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the other specimens). All of these specimens otherwise are similar.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDCE5FEE8653D" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
<emphasis id="DE908E12FF9BFFACD39FDCE5FE5064BE" bold="true" box="[171,410,1154,1177]" pageId="20" pageNumber="471">Habitat distribution.</emphasis>
Members of this species were found under small stones on the open sandy shore of the Longchuan River, a medium-sized stream, at an elevation of
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and together with adults of
<emphasis id="DE908E12FF9BFFACD205DCA5FD9C64FE" box="[305,598,1217,1241]" italics="true" pageId="20" pageNumber="471">
<taxonomicName id="2BE42983FF9BFFACD205DCA5FE5764FE" box="[305,413,1217,1241]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="elegans">T. elegans</taxonomicName>
and
<taxonomicName id="2BE42983FF9BFFACD2E7DCA5FD9C64FE" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[467,598,1218,1241]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="punctatus">T. punctatus</taxonomicName>
</emphasis>
. The specimen from the Tombiguan area was collected in a baited pitfall trap near a stream through secondary (disturbed) broadleaf forest at an elevation of
<quantity id="2B1CFFE5FF9BFFACD3FDDD65FED4653D" box="[201,286,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="20" pageNumber="471" unit="m" value="1290.0">1290 m</quantity>
.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDD44FC3965FC" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
<emphasis id="DE908E12FF9BFFACD39FDD44FD18651D" bold="true" box="[171,722,1315,1338]" pageId="20" pageNumber="471">Geographical distribution within the Gaoligong</emphasis>
<emphasis id="DE908E12FF9BFFACD1EFDD44FCD6651D" bold="true" box="[731,796,1315,1338]" pageId="20" pageNumber="471">
<collectingRegion id="2E209CE2FF9BFFACD1EFDD44FCDD651D" box="[731,791,1315,1338]" country="Myanmar" name="Shan" pageId="20" pageNumber="471">Shan</collectingRegion>
.
</emphasis>
<figureCitation id="74DF4E85FF9BFFACD011DD45FC42651D" box="[805,904,1314,1338]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="20" pageNumber="471">Fig. 11B</figureCitation>
.
<materialsCitation id="5C8C585DFF9BFFACD0A0DD44FD4E65BC" ID-GBIF-Occurrence="4902827320" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2006-06-03" collectionCode="NMPC, R, CAS, IOZ" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. B. Wang &amp; Longchuan Jiang &amp; D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1290" location="Tongbiguan Township" municipality="We" pageId="20" pageNumber="471" specimenCount="15" specimenCount-female="2" specimenCount-male="8" stateProvince="Shan">
<collectingMunicipality id="0C3FC87AFF9BFFACD0A0DD44FC7D651D" box="[916,951,1315,1338]" pageId="20" pageNumber="471">We</collectingMunicipality>
examined a total of
<specimenCount id="FAE29989FF9BFFACD34FDD25FED7657D" box="[123,285,1346,1370]" count="5" pageId="20" pageNumber="471" type="generic">five specimens</specimenCount>
(
<specimenCount id="FAE29989FF9BFFACD21BDD25FE6F657D" box="[303,421,1346,1370]" count="4" pageId="20" pageNumber="471" type="male">four males</specimenCount>
and
<specimenCount id="FAE29989FF9BFFACD2D4DD24FD91657D" box="[480,603,1346,1370]" count="1" pageId="20" pageNumber="471" type="female">one female</specimenCount>
) from the following localities:
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<collectingCounty id="053A2A8CFF9BFFACD08CDD24FB44657C" box="[952,1166,1347,1371]" pageId="20" pageNumber="471">Tengchong County</collectingCounty>
:
</emphasis>
<location id="E93B04DBFF9BFFACD34FDD05FEA3655D" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9BFFACD34FDD05FEA3655D" box="[123,361,1378,1402]" county="Tengchong County" municipality="We" name="Tongbiguan Township" pageId="20" pageNumber="471" stateProvince="Shan">Tongbiguan Township</location>
(
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W of Tongbiguan
</locationDeviation>
, 24.61276°/97.59121°,
<quantity id="2B1CFFE5FF9BFFACD0B4DD05FC19655D" box="[896,979,1378,1402]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="20" pageNumber="471" unit="m" value="1290.0">
<elevation id="67C9B533FF9BFFACD0B4DD05FC19655D" box="[896,979,1378,1402]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="20" pageNumber="471" unit="m" value="1290.0">1290 m</elevation>
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,
<date id="985A74C0FF9BFFACD0EADD05FB5B655D" box="[990,1169,1378,1402]" pageId="20" pageNumber="471" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">
<collectingDate id="881E8D28FF9BFFACD0EADD05FB5B655D" box="[990,1169,1378,1402]" pageId="20" pageNumber="471" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">25-27 June 2016</collectingDate>
</date>
,
<collectorName id="411137D6FF9BFFACD34FDDE4FF1365BC" box="[123,217,1411,1435]" pageId="20" pageNumber="471">J. Hájek</collectorName>
,
<collectorName id="411137D6FF9BFFACD3D2DDE4FE9165BC" box="[230,347,1411,1435]" pageId="20" pageNumber="471">J. Růžička</collectorName>
, &amp;
<collectorName id="411137D6FF9BFFACD2B3DDE4FDCA65BC" box="[391,512,1411,1435]" pageId="20" pageNumber="471">C.B. Wang</collectorName>
collectors [
</materialsCitation>
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<specimenCount id="FAE29989FF9BFFACD1B0DDE4FD2D65BC" box="[644,743,1411,1435]" count="1" pageId="20" pageNumber="471" type="female">1 female</specimenCount>
;
<collectionCode id="8AF5CAC5FF9BFFACD1C1DDE4FC8D65BC" box="[757,839,1411,1435]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="471">NMPC</collectionCode>
];
<location id="E93B04DBFF9BFFACD06EDDE4FBC165BC" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9BFFACD06EDDE4FBC165BC" box="[858,1035,1411,1435]" county="Tengchong County" municipality="We" name="Wuhe Township" pageId="20" pageNumber="471" stateProvince="Shan">Wuhe Township</location>
(
<collectorName id="411137D6FF9BFFACD728DDE4FF79659C" pageId="20" pageNumber="471">Longchuan Jiang</collectorName>
just below bridge at
<location id="E93B04DBFF9BFFACD291DDC4FDAA659C" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9BFFACD291DDC4FDAA659C" box="[421,608,1443,1467]" county="Tengchong County" municipality="We" name="Menglian village" pageId="20" pageNumber="471" stateProvince="Shan">Menglian village</location>
, 24.89176°/98.67551°,
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<elevation id="67C9B533FF9BFFACD050DDC4FC70659C" box="[868,954,1443,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="471" unit="m" value="1230.0">1230 m</elevation>
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,
<date id="985A74C0FF9BFFACD0FDDDC4FB99659C" box="[969,1107,1443,1467]" pageId="20" pageNumber="471" value="2006-06-03">
<collectingDate id="881E8D28FF9BFFACD0FDDDC4FB99659C" box="[969,1107,1443,1467]" pageId="20" pageNumber="471" value="2006-06-03">3 June 2006</collectingDate>
</date>
,
<collectorName id="411137D6FF9BFFACD756DDC3FF3D65FC" pageId="20" pageNumber="471">D.H. Kavanaugh</collectorName>
,
<collectionCode id="8AF5CAC5FF9BFFACD236DDA3FEDD65FC" box="[258,279,1476,1499]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="471">R</collectionCode>
.
<collectorName id="411137D6FF9BFFACD22FDDA3FEBB65FC" box="[283,369,1476,1499]" pageId="20" pageNumber="471">L. Brett</collectorName>
,
<collectorName id="411137D6FF9BFFACD248DDA3FE3E65FC" box="[380,500,1475,1499]" pageId="20" pageNumber="471">H.B. Liang</collectorName>
&amp;
<collectorName id="411137D6FF9BFFACD12CDDA3FD4165FB" box="[536,651,1476,1500]" pageId="20" pageNumber="471">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
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<specimenCount id="FAE29989FF9BFFACD03CDDA4FCAB65FC" box="[776,865,1475,1499]" count="4" pageId="20" pageNumber="471" type="male">4 males</specimenCount>
;
<collectionCode id="8AF5CAC5FF9BFFACD058DDA4FC6865FC" box="[876,930,1475,1499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="471" type="Museum">CAS</collectionCode>
,
<collectionCode id="8AF5CAC5FF9BFFACD09ADDA4FC2B65FC" box="[942,993,1475,1499]" pageId="20" pageNumber="471">IOZ</collectionCode>
]).
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</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDD84FF19663C" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">At present, this species is known only from the southwestern part of the study area (Core Area 6).</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDE43FE4D667B" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
<emphasis id="DE908E12FF9BFFACD39FDE43FDFA661C" bold="true" box="[171,560,1572,1595]" pageId="20" pageNumber="471">Overall geographical distribution.</emphasis>
(
<figureCitation id="74DF4E85FF9BFFACD10BDE43FD45661B" box="[575,655,1572,1596]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="20" pageNumber="471">Fig. 23</figureCitation>
). Known only from Tengchong County, western
<collectingRegion id="2E209CE2FF9BFFACD34FDE22FEFF667B" box="[123,309,1604,1628]" country="China" name="Yunnan" pageId="20" pageNumber="471">Yunnan Province</collectingRegion>
,
<collectingCountry id="94F31290FF9BFFACD274DE23FE49667B" box="[320,387,1604,1628]" name="China" pageId="20" pageNumber="471">China</collectingCountry>
.
</paragraph>
<paragraph id="EC5B5200FF9BFFADD39FDE02FD386134" blockId="20.[123,1174,219,1725]" lastBlockId="21.[123,1172,219,276]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
<emphasis id="DE908E12FF9BFFACD39FDE02FCB7665B" bold="true" box="[171,893,1636,1660]" pageId="20" pageNumber="471">
Geographical relationships with other
<taxonomicName id="2BE42983FF9BFFACD144DE03FCB7665B" box="[624,893,1636,1660]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="DE908E12FF9BFFACD144DE03FCD6665B" bold="true" box="[624,796,1636,1660]" italics="true" pageId="20" pageNumber="471">Tetragonoderus</emphasis>
species.
</taxonomicName>
</emphasis>
Members of this species have been found synotopic with those of
<taxonomicName id="2BE42983FF9BFFACD10EDEE2FD6C66BB" box="[570,678,1668,1692]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9BFFACD10EDEE2FD6C66BB" box="[570,678,1668,1692]" italics="true" pageId="20" pageNumber="471">T. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="2BE42983FF9BFFACD1E9DEE2FCAA66BB" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[733,864,1669,1692]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9BFFACD1E9DEE2FCAA66BB" box="[733,864,1669,1692]" italics="true" pageId="20" pageNumber="471">T. punctatus</emphasis>
</taxonomicName>
in the study area. The range of this species also overlaps that of
<taxonomicName id="2BE42983FF9BFFACD2C0DEC1FDA0669A" box="[500,618,1702,1725]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF9BFFACD2C0DEC1FDA0669A" box="[500,618,1702,1725]" italics="true" pageId="20" pageNumber="471">T. arcuatus</emphasis>
</taxonomicName>
in the study area, but these two species have not yet been found syntopic. Also, the range of
<taxonomicName id="2BE42983FF9AFFADD118D8BBFD0060D4" authorityName="Jian &amp; Tian" authorityYear="2009" box="[556,714,219,243]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF9AFFADD118D8BBFD0060D4" box="[556,714,219,243]" italics="true" pageId="21" pageNumber="472">T. microthorax</emphasis>
</taxonomicName>
overlaps that of
<taxonomicName id="2BE42983FF9AFFADD0B5D8BBFBC760D4" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[897,1037,219,243]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF9AFFADD0B5D8BBFBC760D4" box="[897,1037,219,243]" italics="true" pageId="21" pageNumber="472">T. parviculus</emphasis>
</taxonomicName>
, but the former species has not yet been recorded from the study area.
</paragraph>
</subSubSection>
</treatment>
</document>
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